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<document id="CE523FCBF4B0A287FA52CAFAA51BBA29" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A005CFFD9FF182F5E11ED3772" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus variolatus LeConte. The 1852" docType="treatment" docVersion="8" lastPageNumber="481" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="481" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="318EE87B220FD11B8A74BFB14EE069B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBA72279ED5F6F16C36174D493B04D81">
<mods:title id="8A580B5214178CF03B0C2281892F110C">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="D1B2711BCF5EBA03966F07DD655A1FA6" type="personal">
<mods:role id="2996AAE1D2BA0250A4402BA37A1517C5">
<mods:roleTerm id="5D31064365435624E6E4C2DA3B2D5837">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDC062D1509FFCD8E98D9C5E25D6CBBD">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="992B61DFB266CC4B8FA1622985CF73B5">text</mods:typeOfResource>
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<mods:title id="5566ECD3884948DAA4C19ED03D3E1707">Zootaxa</mods:title>
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<mods:part id="6DC9FA0C6341008C847FC3743EC10720">
<mods:date id="6E6D6387A7EED2A40363303D9FE290FB">2021</mods:date>
<mods:detail id="396FC7A1FED4A635B5A38226ADB08231" type="pubDate">
<mods:number id="A6DF6B2A5ABEB1AB08E50C99287F4760">2021-05-21</mods:number>
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<mods:detail id="D3707E64C91D1E7F9E1B0626F4A596A7" type="volume">
<mods:number id="AE097C9A666D2310307C20201B3AFEB7">4974</mods:number>
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<mods:number id="ADBAA5DF7449B3D5CA471FCD6B04375A">3</mods:number>
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<mods:start id="4F25E5D73C743FCC028F41322BB8E682">459</mods:start>
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<mods:classification id="0F07F8F63574C0467778D1B3F9C1B74E">journal article</mods:classification>
<mods:identifier id="076679B9B551991BD7C7BD39A7D98BAB" type="CLB-Dataset">6214</mods:identifier>
<mods:identifier id="15F6283885146382E2294E1CE45DDCCD" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
<mods:identifier id="1193462E8F71D740BE71791DACBE7816" type="GBIF-Dataset">5b4eaeb3-5722-4021-84ad-125ccd80855f</mods:identifier>
<mods:identifier id="09E6EB2E2F9FA5E7986BDBA9A770BE13" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="20D80A57A64E2364932609AD8F421617" type="Zenodo-Dep">4777839</mods:identifier>
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</mods:mods>
<treatment id="4772526A005CFFD9FF182F5E11ED3772" ID-DOI="http://doi.org/10.5281/zenodo.4777948" ID-GBIF-Taxon="182642554" ID-Zenodo-Dep="4777948" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772" httpUri="http://treatment.plazi.org/id/4772526A005CFFD9FF182F5E11ED3772" lastPageNumber="481" pageId="22" pageNumber="481">
<subSubSection id="87C1B0F7005CFFD9FF182F5E170D3250" box="[151,528,582,609]" pageId="22" pageNumber="481" type="nomenclature">
<paragraph id="CF64E37C005CFFD9FF182F5E170D3250" blockId="22.[151,528,582,645]" box="[151,528,582,609]" pageId="22" pageNumber="481">
<heading id="942C5410005CFFD9FF182F5E170D3250" bold="true" box="[151,528,582,609]" fontSize="11" level="1" pageId="22" pageNumber="481" reason="1">
<taxonomicName id="08DB98FF005CFFD9FF182F5E170D3250" ID-CoL="3N6CY" authority="LeConte" authorityName="LeConte. The" authorityYear="1852" box="[151,528,582,609]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="variolatus">
<emphasis id="FDAF3F6E005CFFD9FF182F5E170D3250" bold="true" box="[151,528,582,609]" pageId="22" pageNumber="481">
<emphasis id="FDAF3F6E005CFFD9FF182F5E14803251" bold="true" box="[151,413,582,609]" italics="true" pageId="22" pageNumber="481">Hydrochus variolatus</emphasis>
LeConte
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005CFFD9FF182F73149632B5" box="[151,395,619,645]" pageId="22" pageNumber="481" type="description">
<paragraph id="CF64E37C005CFFD9FF182F73149632B5" blockId="22.[151,528,582,645]" box="[151,395,619,645]" pageId="22" pageNumber="481">
<figureCitation id="57E0FFF9005CFFD9FF182F7315F832B5" box="[151,229,619,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="22" pageNumber="481">Figs. 2</figureCitation>
,
<figureCitation id="57E0FFF9005CFFD9FF7E2F73142432B5" box="[241,313,619,645]" captionStart="FIGURES 17–18" captionStartId="30.[151,264,1922,1946]" captionTargetBox="[170,1430,188,1882]" captionTargetId="figure-22@30.[151,1435,181,1899]" captionTargetPageId="30" captionText="FIGURES 17–18. Habitus and male genitalia. 17. Hydrochus variolatus LeConte, non-type. 18. Hydrochus variolatus LeConte, holotype." figureDoi="http://doi.org/10.5281/zenodo.4777877" httpUri="https://zenodo.org/record/4777877/files/figure.png" pageId="22" pageNumber="481">17, 18</figureCitation>
,
<figureCitation id="57E0FFF9005CFFD9FECA2F73147E32B5" box="[325,355,619,645]" captionStart="FIGURES 25–27. 25" captionStartId="34.[151,264,1944,1968]" captionTargetBox="[170,1427,194,1905]" captionTargetId="figure-22@34.[151,1435,181,1920]" captionTargetPageId="34" captionText="FIGURES 25–27. 25. Hydrochus aeruginosus n. sp., holotype. 26. Hydrochus variolatus LeConte, type labels. 27. Hydrochus vagus LeConte, type labels" figureDoi="http://doi.org/10.5281/zenodo.4777900" httpUri="https://zenodo.org/record/4777900/files/figure.png" pageId="22" pageNumber="481">26</figureCitation>
,
<figureCitation id="57E0FFF9005CFFD9FEE02F73149632B5" box="[367,395,619,645]" captionStart="FIGURES 34–36" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[166,1421,181,1920]" captionTargetId="figure-15@37.[166,1421,181,1920]" captionTargetPageId="37" captionText="FIGURES 34–36. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777912" httpUri="https://zenodo.org/record/4777912/files/figure.png" pageId="22" pageNumber="481">34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005CFFD9FF182FB717A632D5" pageId="22" pageNumber="481" type="reference_group">
<paragraph id="CF64E37C005CFFD9FF182FB7175E32F7" blockId="22.[151,699,687,742]" box="[151,579,687,711]" pageId="22" pageNumber="481">
<treatmentCitationGroup id="EFCBC452005CFFD9FF182FB7175E32F7" box="[151,579,687,711]" pageId="22" pageNumber="481">
<treatmentCitation id="4E7AC56D005CFFD9FF182FB7172332F7" author="LeConte, J. L." box="[151,574,687,711]" page="211" pageId="22" pageNumber="481" year="1852">
<taxonomicName id="08DB98FF005CFFD9FF182FB7172332F7" ID-CoL="3N6CY" authority="LeConte 1852: 211" authorityName="LeConte. The" authorityPageNumber="211" authorityYear="1852" box="[151,574,687,711]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="variolatus">
<emphasis id="FDAF3F6E005CFFD9FF182FB7146E32F7" box="[151,371,687,711]" italics="true" pageId="22" pageNumber="481">Hydrochus variolatus</emphasis>
<bibRefCitation id="AB4A9E8D005CFFD9FEF62FB7172332F7" author="LeConte, J. L." box="[377,574,687,711]" pageId="22" pageNumber="481" pagination="125 - 216" refId="ref24696" refString="LeConte, J. L. (1852) Descriptions of new species of Coleoptera, from California (1851. Annals of the Lyceum of natural History of New York, 5, 125 - 216." type="journal article" year="1852">LeConte 1852: 211</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CF64E37C005CFFD9FF182FD617A632D5" blockId="22.[151,699,687,742]" box="[151,699,717,742]" pageId="22" pageNumber="481">
<treatmentCitationGroup id="EFCBC452005CFFD9FF182FD617A632D5" box="[151,699,717,742]" pageId="22" pageNumber="481">
<treatmentCitation id="4E7AC56D005CFFD9FF182FD6171932D6" author="Makhan, D." box="[151,516,718,742]" page="67" pageId="22" pageNumber="481" year="1993">
<taxonomicName id="08DB98FF005CFFD9FF182FD6171932D6" ID-CoL="6MJX8" authority="Makhan 1993: 67" authorityName="Makhan (HNHM" authorityPageNumber="67" authorityYear="1993" box="[151,516,718,742]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="otvosi">
<emphasis id="FDAF3F6E005CFFD9FF182FD6145432D5" box="[151,329,718,742]" italics="true" pageId="22" pageNumber="481">Hydrochus otvosi</emphasis>
<bibRefCitation id="AB4A9E8D005CFFD9FEC02FD6171932D6" author="Makhan, D." box="[335,516,718,742]" pageId="22" pageNumber="481" pagination="65 - 70" refId="ref24761" refString="Makhan, D. (1993) Nine new Hydrochus species from South and Central America. Annales historico-naturales musei nationalis hungarici, 85, 65 - 70." type="journal article" year="1993">Makhan 1993: 67</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="FDAF3F6E005CFFD9FD812FD617A632D5" bold="true" box="[526,699,717,742]" pageId="22" pageNumber="481">New Synonymy.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005CFFD9FF182E20106F34A6" pageId="22" pageNumber="481" type="materials_examined">
<paragraph id="CF64E37C005CFFD9FF182E20142F338E" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">
<emphasis id="FDAF3F6E005CFFD9FF182E2014583362" bold="true" box="[151,325,824,850]" pageId="22" pageNumber="481">Type material.</emphasis>
<materialsCitation id="7FB3E921005CFFD9FEC32E201433338E" ID-GBIF-Occurrence="3128194508" country="Mexico" county="California" location="Images" municipality="San Diego" pageId="22" pageNumber="481" specimenCount="1" typeStatus="holotype">
There is only
<specimenCount id="D9DD28F5005CFFD9FE682E21179D3362" box="[487,640,824,851]" pageId="22" pageNumber="481" type="generic">one specimen</specimenCount>
of
<taxonomicName id="08DB98FF005CFFD9FD2A2E20111F3362" authority="LeConte" authorityName="LeConte. The" authorityYear="1852" box="[677,1026,824,850]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="variolatus">
<emphasis id="FDAF3F6E005CFFD9FD2A2E2016853362" box="[677,920,824,850]" italics="true" pageId="22" pageNumber="481">Hydrochus variolatus</emphasis>
LeConte
</taxonomicName>
, a male, in the LeConte collection at the MCZ. It is therefore the
<typeStatus id="10605DDE005CFFD9FE5B2E4417243346" box="[468,569,860,886]" pageId="22" pageNumber="481" type="holotype">holotype</typeStatus>
, by monotypy.
<location id="CA04B5A7005CFFD9FD692E45162A3347" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FD692E45162A3347" box="[742,823,861,887]" country="Mexico" county="California" municipality="San Diego" name="Images" pageId="22" pageNumber="481">Images</location>
of the habitus, male genitalia, and labels of this specimen given herein (
<figureCitation id="57E0FFF9005CFFD9FEE12E9814D033AA" box="[366,461,896,922]" captionStart="FIGURES 17–18" captionStartId="30.[151,264,1922,1946]" captionTargetBox="[170,1430,188,1882]" captionTargetId="figure-22@30.[151,1435,181,1899]" captionTargetPageId="30" captionText="FIGURES 17–18. Habitus and male genitalia. 17. Hydrochus variolatus LeConte, non-type. 18. Hydrochus variolatus LeConte, holotype." figureDoi="http://doi.org/10.5281/zenodo.4777877" httpUri="https://zenodo.org/record/4777877/files/figure.png" pageId="22" pageNumber="481">Figs. 18</figureCitation>
,
<figureCitation id="57E0FFF9005CFFD9FE542E9814E633AB" box="[475,507,896,923]" captionStart="FIGURES 25–27. 25" captionStartId="34.[151,264,1944,1968]" captionTargetBox="[170,1427,194,1905]" captionTargetId="figure-22@34.[151,1435,181,1920]" captionTargetPageId="34" captionText="FIGURES 25–27. 25. Hydrochus aeruginosus n. sp., holotype. 26. Hydrochus variolatus LeConte, type labels. 27. Hydrochus vagus LeConte, type labels" figureDoi="http://doi.org/10.5281/zenodo.4777900" httpUri="https://zenodo.org/record/4777900/files/figure.png" pageId="22" pageNumber="481">26</figureCitation>
).
<location id="CA04B5A7005CFFD9FD9F2E98179E33AA" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FD9F2E98179E33AA" box="[528,643,896,922]" country="Mexico" county="California" municipality="San Diego" name="Originally" pageId="22" pageNumber="481">Originally</location>
described from
<collectingMunicipality id="2F007906005CFFD9FCB02E9816A433AA" box="[831,953,896,922]" pageId="22" pageNumber="481">San Diego</collectingMunicipality>
,
<collectingCounty id="26059BF0005CFFD9FC492E98112733AA" box="[966,1082,896,922]" pageId="22" pageNumber="481">California</collectingCounty>
, the following new records are from
<collectingCountry id="B7CCA3EC005CFFD9FF5A2EBC1433338E" box="[213,302,932,958]" name="Mexico" pageId="22" pageNumber="481">Mexico</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="CF64E37C005CFFD9FF482ED0106F34A6" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">
<emphasis id="FDAF3F6E005CFFD9FF482ED014F033D2" bold="true" box="[199,493,968,994]" pageId="22" pageNumber="481">Material examined (19):</emphasis>
<materialsCitation id="7FB3E921005CFFD9FE792ED0142A3436" ID-GBIF-Occurrence="3128194565" collectingDate="1938-06-14" collectionCode="CAS" collectorName="Michelbacher &amp; Ross" country="Mexico" location="Ensenada" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Baja California">
<emphasis id="FDAF3F6E005CFFD9FE792ED0175033D2" bold="true" box="[502,589,968,994]" pageId="22" pageNumber="481">
<collectingCountry id="B7CCA3EC005CFFD9FE792ED0175033D2" box="[502,589,968,994]" name="Mexico" pageId="22" pageNumber="481">Mexico</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E005CFFD9FDD22ED0161333D2" box="[605,782,968,994]" country="Mexico" name="Baja California" pageId="22" pageNumber="481">Baja California</collectingRegion>
,
<quantity id="08234E99005CFFD9FC942ED0167833D2" box="[795,869,968,994]" metricMagnitude="4" metricUnit="m" metricValue="2.7358848" pageId="22" pageNumber="481" unit="mi" value="17.0">17 mi.</quantity>
S. of
<location id="CA04B5A7005CFFD9FC212ED0110133D2" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FC212ED0110133D2" box="[942,1052,968,994]" country="Mexico" name="Ensenada" pageId="22" pageNumber="481" stateProvince="Baja California">Ensenada</location>
,
<date id="BB65C5BC005CFFD9FBA62ED011B933D2" box="[1065,1188,968,994]" pageId="22" pageNumber="481" value="1938-06-14">
<collectingDate id="AB213C54005CFFD9FBA62ED011B933D2" box="[1065,1188,968,994]" pageId="22" pageNumber="481" value="1938-06-14">14.vi.1938</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FB6C2ED0106033D2" box="[1251,1405,968,994]" pageId="22" pageNumber="481">Michelbacher</collectorName>
&amp;
<collectorName id="622E86AA005CFFD9FF182EF515D03437" box="[151,205,1005,1031]" pageId="22" pageNumber="481">Ross</collectorName>
(9
<collectionCode id="A9CA7BB9005CFFD9FF7B2EF414323436" box="[244,303,1004,1030]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="481" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FECC2EF51404341A" ID-GBIF-Occurrence="3128194460" collectingDate="1963-04-23" collectionCode="CAS" collectorName="H. B. Leech &amp; P. H. Arnaud Jr." country="Mexico" location="Arr. Santo Domingo" municipality="Norte" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Baja California">
<collectingMunicipality id="2F007906005CFFD9FECC2EF5149B3437" box="[323,390,1005,1031]" pageId="22" pageNumber="481">Norte</collectingMunicipality>
,
<location id="CA04B5A7005CFFD9FE1F2EF417653436" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FE1F2EF417653436" box="[400,632,1004,1030]" country="Mexico" municipality="Norte" name="Arr. Santo Domingo" pageId="22" pageNumber="481" stateProvince="Baja California">Arr. Santo Domingo</location>
,
<quantity id="08234E99005CFFD9FD0C2EF417CE3436" box="[643,723,1004,1031]" metricMagnitude="3" metricUnit="m" metricValue="9.1732608" pageId="22" pageNumber="481" unit="mi" value="5.7">5.7 mi.</quantity>
<collectorName id="622E86AA005CFFD9FD552EF516A83436" box="[730,949,1004,1031]" pageId="22" pageNumber="481">E. Hamilton Ranch</collectorName>
,
<date id="BB65C5BC005CFFD9FC4E2EF411243436" box="[961,1081,1004,1030]" pageId="22" pageNumber="481" value="1963-04-23">
<collectingDate id="AB213C54005CFFD9FC4E2EF411243436" box="[961,1081,1004,1030]" pageId="22" pageNumber="481" value="1963-04-23">23.iv.1963</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FBF92EF5101C3436" box="[1142,1281,1004,1031]" pageId="22" pageNumber="481">H. B. Leech</collectorName>
,
<collectorName id="622E86AA005CFFD9FA822EF510853436" box="[1293,1432,1004,1031]" pageId="22" pageNumber="481">P.H. Arnaud</collectorName>
,
<collectorName id="622E86AA005CFFD9FF18290915AC341B" box="[151,177,1041,1067]" pageId="22" pageNumber="481">Jr.</collectorName>
(4
<collectionCode id="A9CA7BB9005CFFD9FF592908140C341A" box="[214,273,1040,1066]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="481" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FEAA2909107A341A" ID-GBIF-Occurrence="3128194404" box="[293,1383,1040,1067]" collectingDate="1970-07-19" collectionCode="MCZ" collectorName="S. Laguna Hanson &amp; P. D. Perkins" country="Mexico" elevation="1524" location="Sa. Juarez" municipality="Norte" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Baja California">
<collectingMunicipality id="2F007906005CFFD9FEAA29091475341B" box="[293,360,1041,1067]" pageId="22" pageNumber="481">Norte</collectingMunicipality>
,
<location id="CA04B5A7005CFFD9FEFC290814FA341B" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FEFC290814FA341B" box="[371,487,1040,1067]" country="Mexico" municipality="Norte" name="Sa. Juarez" pageId="22" pageNumber="481" stateProvince="Baja California">Sa. Juarez</location>
,
<quantity id="08234E99005CFFD9FE7C29081730341A" box="[499,557,1040,1066]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="22" pageNumber="481" unit="mi" value="1.0">1 mi.</quantity>
<collectorName id="622E86AA005CFFD9FDBC29081619341B" box="[563,772,1040,1067]" pageId="22" pageNumber="481">S. Laguna Hanson</collectorName>
, elev.
<quantity id="08234E99005CFFD9FCC429081683341A" box="[843,926,1040,1067]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="481" unit="ft" value="5000.0">
<elevation id="44F6044F005CFFD9FCC429081683341A" box="[843,926,1040,1067]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="481" unit="ft" value="5000.0">5000 ft</elevation>
</quantity>
,
<date id="BB65C5BC005CFFD9FC2629081136341A" box="[937,1067,1040,1066]" pageId="22" pageNumber="481" value="1970-07-19">
<collectingDate id="AB213C54005CFFD9FC2629081136341A" box="[937,1067,1040,1066]" pageId="22" pageNumber="481" value="1970-07-19">19.vii.1970</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FBE8290911E5341A" box="[1127,1272,1040,1067]" pageId="22" pageNumber="481">P. D. Perkins</collectorName>
(1
<collectionCode id="A9CA7BB9005CFFD9FA9229081042341A" box="[1309,1375,1040,1066]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="481" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FAFD2908168D347E" ID-GBIF-Occurrence="3128194377" collectingDate="1961-06-15" collectionCode="TAMU" collectorName="E. L. Sleeper" country="Mexico" location="San Jose del Castillo" pageId="22" pageNumber="481" specimenCount="1" specimenCount-male="1" stateProvince="Baja California">
<location id="CA04B5A7005CFFD9FAFD2908144E347E" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FAFD2908144E347E" country="Mexico" name="San Jose del Castillo" pageId="22" pageNumber="481" stateProvince="Baja California">San Jose del Castillo</location>
,
<date id="BB65C5BC005CFFD9FED1292C14C5347F" box="[350,472,1076,1103]" pageId="22" pageNumber="481" value="1961-06-15">
<collectingDate id="AB213C54005CFFD9FED1292C14C5347F" box="[350,472,1076,1103]" pageId="22" pageNumber="481" value="1961-06-15">15.vi.1961</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FD9A292D17B5347E" box="[533,680,1076,1103]" pageId="22" pageNumber="481">E. L. Sleeper</collectorName>
-
<specimenCount id="D9DD28F5005CFFD9FD4E292C1611347E" box="[705,780,1076,1102]" pageId="22" pageNumber="481" type="male">1 male</specimenCount>
(1
<collectionCode id="A9CA7BB9005CFFD9FCBD292C1695347E" box="[818,904,1076,1102]" pageId="22" pageNumber="481">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FC14292C14283443" ID-GBIF-Occurrence="3128194305" collectingDate="1962-05-18" collectionCode="NMNH" collectorName="F. Islas" country="Mexico" location="Escarcega" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Campeche">
<collectingRegion id="0D1F2D9E005CFFD9FC14292C1109347E" box="[923,1044,1076,1102]" country="Mexico" name="Campeche" pageId="22" pageNumber="481">Campeche</collectingRegion>
,
<location id="CA04B5A7005CFFD9FBAF292D1189347F" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FBAF292D1189347F" box="[1056,1172,1077,1103]" country="Mexico" name="Escarcega" pageId="22" pageNumber="481" stateProvince="Campeche">Escarcega</location>
,
<date id="BB65C5BC005CFFD9FB2F292C100D347E" box="[1184,1296,1076,1102]" pageId="22" pageNumber="481" value="1962-05-18">
<collectingDate id="AB213C54005CFFD9FB2F292C100D347E" box="[1184,1296,1076,1102]" pageId="22" pageNumber="481" value="1962-05-18">18.v.1962</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FAC2292D1081347E" box="[1357,1436,1076,1103]" pageId="22" pageNumber="481">F. Islas</collectorName>
S. (1
<collectionCode id="A9CA7BB9005CFFD9FF5D294114303443" box="[210,301,1113,1139]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="481" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FEB02940115B3442" ID-GBIF-Occurrence="3128194407" box="[319,1094,1112,1138]" collectingDate="1989-06-04" collectionCode="UNAM" collectorName="G. Ortega" country="Mexico" location="Rio Chamela" municipality="Chamela" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Jalisco">
<collectingRegion id="0D1F2D9E005CFFD9FEB0294014933442" box="[319,398,1112,1138]" country="Mexico" name="Jalisco" pageId="22" pageNumber="481">Jalisco</collectingRegion>
,
<collectingMunicipality id="2F007906005CFFD9FE16294014E23442" box="[409,511,1112,1138]" pageId="22" pageNumber="481">Chamela</collectingMunicipality>
,
<location id="CA04B5A7005CFFD9FD85294017823442" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FD85294017823442" box="[522,671,1112,1138]" country="Mexico" municipality="Chamela" name="Rio Chamela" pageId="22" pageNumber="481" stateProvince="Jalisco">Rio Chamela</location>
,
<date id="BB65C5BC005CFFD9FD252940160B3442" box="[682,790,1112,1138]" pageId="22" pageNumber="481" value="1989-06-04">
<collectingDate id="AB213C54005CFFD9FD252940160B3442" box="[682,790,1112,1138]" pageId="22" pageNumber="481" value="1989-06-04">4.vi.1989</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FCDE294016A33442" box="[849,958,1112,1138]" pageId="22" pageNumber="481">G. Ortega</collectorName>
(1
<collectionCode id="A9CA7BB9005CFFD9FC6C294011233442" box="[995,1086,1112,1138]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="22" pageNumber="481">UNAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FBDF2941178734A7" ID-GBIF-Occurrence="3128194386" collectingDate="1965-08-27" collectionCode="NMNH" collectorName="W. Poza Rica &amp; Paul J. Spangler" country="Mexico" location="Veracruz" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingRegion id="0D1F2D9E005CFFD9FBDF294111AA3443" box="[1104,1207,1113,1139]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="22" pageNumber="481">Veracruz</collectingRegion>
,
<quantity id="08234E99005CFFD9FB4E294011E73442" box="[1217,1274,1112,1138]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="22" pageNumber="481" unit="mi" value="9.0">9 mi.</quantity>
<collectorName id="622E86AA005CFFD9FA8F294110853442" box="[1280,1432,1112,1139]" pageId="22" pageNumber="481">W. Poza Rica</collectorName>
,
<date id="BB65C5BC005CFFD9FF182964143C34A7" box="[151,289,1148,1175]" pageId="22" pageNumber="481" value="1965-08-27">
<collectingDate id="AB213C54005CFFD9FF182964143C34A7" box="[151,289,1148,1175]" pageId="22" pageNumber="481" value="1965-08-27">27.viii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FED22964170F34A6" box="[349,530,1148,1175]" pageId="22" pageNumber="481">Paul J. Spangler</collectorName>
(1
<collectionCode id="A9CA7BB9005CFFD9FDB82965178F34A7" box="[567,658,1149,1175]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="481" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005CFFD9FD2A2964107334A6" ID-GBIF-Occurrence="3128194433" box="[677,1390,1148,1175]" collectingDate="1948-12-17" collectionCode="CAS" collectorName="El Fortin &amp; H. B. Leech" country="Mexico" location="Rio Metlao" pageId="22" pageNumber="481" specimenCount="1" stateProvince="Veracruz">
<location id="CA04B5A7005CFFD9FD2A2964163E34A6" LSID="urn:lsid:plazi:treatment:4772526A005CFFD9FF182F5E11ED3772:CA04B5A7005CFFD9FD2A2964163E34A6" box="[677,803,1148,1174]" country="Mexico" name="Rio Metlao" pageId="22" pageNumber="481" stateProvince="Veracruz">Rio Metlao</location>
nr.
<collectorName id="622E86AA005CFFD9FCC1296416A834A6" box="[846,949,1148,1174]" pageId="22" pageNumber="481">El Fortin</collectorName>
,
<date id="BB65C5BC005CFFD9FC4E2964115E34A6" box="[961,1091,1148,1174]" pageId="22" pageNumber="481" value="1948-12-17">
<collectingDate id="AB213C54005CFFD9FC4E2964115E34A6" box="[961,1091,1148,1174]" pageId="22" pageNumber="481" value="1948-12-17">17.xii.1948</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005CFFD9FBF02965101834A6" box="[1151,1285,1148,1175]" pageId="22" pageNumber="481">H. B. Leech</collectorName>
(1
<collectionCode id="A9CA7BB9005CFFD9FAA42964107B34A6" box="[1323,1382,1148,1174]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="481" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005CFFD9FF4829B811523682" pageId="22" pageNumber="481" type="diagnosis">
<paragraph id="CF64E37C005CFFD9FF4829B8146B355E" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">
<emphasis id="FDAF3F6E005CFFD9FF4829B814D0348A" bold="true" box="[199,461,1184,1210]" pageId="22" pageNumber="481">Differential diagnosis.</emphasis>
Among Central American species, recognized by the combination of moderate size (ca.
<quantity id="08234E99005CFFD9FF1829DC145D34EF" box="[151,320,1220,1247]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" metricValueMax="3.46" metricValueMin="3.1" pageId="22" pageNumber="481" unit="mm" value="3.28" valueMax="3.46" valueMin="3.1">3.10–3.46 mm</quantity>
), the deep pronotal depressions, the well raised and relatively impunctate pronotal reliefs, and the male genitalia (
<figureCitation id="57E0FFF9005CFFD9FECA29F014D33532" box="[325,462,1256,1282]" captionStart="FIGURES 17–18" captionStartId="30.[151,264,1922,1946]" captionTargetBox="[170,1430,188,1882]" captionTargetId="figure-22@30.[151,1435,181,1899]" captionTargetPageId="30" captionText="FIGURES 17–18. Habitus and male genitalia. 17. Hydrochus variolatus LeConte, non-type. 18. Hydrochus variolatus LeConte, holotype." figureDoi="http://doi.org/10.5281/zenodo.4777877" httpUri="https://zenodo.org/record/4777877/files/figure.png" pageId="22" pageNumber="481">Figs. 17, 18</figureCitation>
; described below). This species has a generalized
<taxonomicName id="08DB98FF005CFFD9FB8E29F011663532" authorityName="Leach" authorityYear="1817" box="[1025,1147,1256,1282]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="FDAF3F6E005CFFD9FB8E29F011663532" box="[1025,1147,1256,1282]" italics="true" pageId="22" pageNumber="481">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The male genitalia have a very large adbl, and a unique character: the adbl and the continuing shaft of the aedeagus are on the same plane for about 1/3 the length of the aedeagus; this is best seen in lateral view.
</paragraph>
<paragraph id="CF64E37C005CFFD9FF48286016463676" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">
Male genitalia (
<figureCitation id="57E0FFF9005CFFD9FEF52860171835A2" box="[378,517,1400,1426]" captionStart="FIGURES 17–18" captionStartId="30.[151,264,1922,1946]" captionTargetBox="[170,1430,188,1882]" captionTargetId="figure-22@30.[151,1435,181,1899]" captionTargetPageId="30" captionText="FIGURES 17–18. Habitus and male genitalia. 17. Hydrochus variolatus LeConte, non-type. 18. Hydrochus variolatus LeConte, holotype." figureDoi="http://doi.org/10.5281/zenodo.4777877" httpUri="https://zenodo.org/record/4777877/files/figure.png" pageId="22" pageNumber="481">Figs. 17, 18</figureCitation>
). General characters: paramere tips slender and pointed, narrower in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, then weakly arcuate away from midline, plsa absent or very slight; aedeagus in dorsal/ventral views moderately wide, parallelsided or nearly so; in lateral view wider, especially basally, adbl and continuing shaft of aedeagus on same plane for about 1/3 length of aedeagus, after which shaft dorsal margin is sinuate; lateral margins of aedeagus sclerotized; basal piece shorter than parameres, orifice strongly sclerotized.
</paragraph>
<paragraph id="CF64E37C005CFFD9FF482B48118736BE" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">Dorsal surface: pdmm sinuate, slightly overlapping aedeagus; adbl very large, ca. ½ as wide as total genitalia width at that level; adtl at about midlength of shaft; paramere tips without sclerotized ridge.</paragraph>
<paragraph id="CF64E37C005CFFD9FF482B8011523682" blockId="22.[151,1437,824,1859]" box="[199,1103,1688,1715]" pageId="22" pageNumber="481">Ventral surface: pvmm approximating lateral margin of aedeagus; avtl near base.</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005CFFD9FF482BA411ED3772" pageId="22" pageNumber="481" type="discussion">
<paragraph id="CF64E37C005CFFD9FF482BA411ED3772" blockId="22.[151,1437,824,1859]" pageId="22" pageNumber="481">
<emphasis id="FDAF3F6E005CFFD9FF482BA4142936E6" bold="true" box="[199,308,1724,1750]" pageId="22" pageNumber="481">Remarks</emphasis>
. I have examined the
<typeStatus id="10605DDE005CFFD9FDA12BA4178D36E6" box="[558,656,1724,1750]" pageId="22" pageNumber="481" type="holotype">holotype</typeStatus>
of
<taxonomicName id="08DB98FF005CFFD9FD392BA4114D36E6" authority="Makhan (HNHM)" authorityName="Makhan (HNHM" authorityYear="1993" box="[694,1104,1724,1750]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="otvosi">
<emphasis id="FDAF3F6E005CFFD9FD392BA4166636E6" box="[694,891,1724,1750]" italics="true" pageId="22" pageNumber="481">Hydrochus otvosi</emphasis>
Makhan (HNHM)
</taxonomicName>
, including the male genitalia, and have determined that it is a synonym of
<taxonomicName id="08DB98FF005CFFD9FD152BF8112E36CA" authority="LeConte. The" authorityName="LeConte. The" authorityYear="1852" box="[666,1075,1760,1786]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="variolatus">
<emphasis id="FDAF3F6E005CFFD9FD152BF8169236CA" box="[666,911,1760,1786]" italics="true" pageId="22" pageNumber="481">Hydrochus variolatus</emphasis>
LeConte. The
</taxonomicName>
<taxonomicName id="08DB98FF005CFFD9FBB32BF911BE36CA" authorityName="Makhan (HNHM" authorityYear="1993" box="[1084,1187,1761,1786]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="otvosi">
<emphasis id="FDAF3F6E005CFFD9FBB32BF911BE36CA" box="[1084,1187,1761,1786]" italics="true" pageId="22" pageNumber="481">H. otvosi</emphasis>
</taxonomicName>
<typeStatus id="10605DDE005CFFD9FB232BF8101336CA" box="[1196,1294,1760,1786]" pageId="22" pageNumber="481" type="holotype">holotype</typeStatus>
specimen is supposedly from
<collectingCountry id="B7CCA3EC005CFFD9FED42A1C14C4372E" box="[347,473,1796,1822]" name="Costa Rica" pageId="22" pageNumber="481">Costa Rica</collectingCountry>
. The non-type specimen of
<taxonomicName id="08DB98FF005CFFD9FC9D2A1D16BA372E" authorityName="LeConte. The" authorityYear="1852" box="[786,935,1796,1822]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="variolatus">
<emphasis id="FDAF3F6E005CFFD9FC9D2A1D16BA372E" box="[786,935,1796,1822]" italics="true" pageId="22" pageNumber="481">H. variolatus</emphasis>
</taxonomicName>
imaged (
<figureCitation id="57E0FFF9005CFFD9FB9F2A1C117F372E" box="[1040,1122,1796,1822]" captionStart="FIGURES 17–18" captionStartId="30.[151,264,1922,1946]" captionTargetBox="[170,1430,188,1882]" captionTargetId="figure-22@30.[151,1435,181,1899]" captionTargetPageId="30" captionText="FIGURES 17–18. Habitus and male genitalia. 17. Hydrochus variolatus LeConte, non-type. 18. Hydrochus variolatus LeConte, holotype." figureDoi="http://doi.org/10.5281/zenodo.4777877" httpUri="https://zenodo.org/record/4777877/files/figure.png" pageId="22" pageNumber="481">Fig. 17</figureCitation>
) has this locality data: Baja California, Norte, Sa. Juarez,
<quantity id="08234E99005CFFD9FE6B2A3017033772" box="[484,542,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="22" pageNumber="481" unit="mi" value="1.0">1 mi.</quantity>
S. Laguna Hanson, elev.
<quantity id="08234E99005CFFD9FCB42A3016933772" box="[827,910,1832,1859]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="481" unit="ft" value="5000.0">5000 ft</quantity>
,
<date id="BB65C5BC005CFFD9FC152A3011013772" box="[922,1052,1832,1858]" pageId="22" pageNumber="481" value="1970-07-19">19.vii.1970</date>
, leg. P. D. Perkins.
</paragraph>
</subSubSection>
</treatment>
</document>
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