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<document id="4D9A87A977676E9DBC6184781EAA8F58" ID-CLB-Dataset="36901" ID-DOI="10.11646/zootaxa.3765.1.1" ID-GBIF-Dataset="dc48dc34-f0b6-4715-8044-1aa2d5851737" ID-ISSN="1175-5326" ID-Zenodo-Dep="285551" ID-ZooBank="4776DD1C-9FCC-4F05-8FDF-66AA14B959FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460384113075" checkinUser="plazi" docAuthor="Kataev, Boris M., Wrase, David W. &amp; Schmidt, Joachim" docDate="2014" docId="232FAA28FFC9021AFF53FD42FD23F850" docLanguage="en" docName="zt03765p028.pdf" docOrigin="Zootaxa 3765 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chydaeus putaoensis, sp. n." docType="treatment" docVersion="14" lastPageNumber="18" masterDocId="DF16D250FFC6020BFFC4FF95FF9EF910" masterDocTitle="New species of the genus Chydaeus from China, Nepal, Myanmar, and Thailand, with remarks on species previously described (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="16" updateTime="1698297231703" updateUser="ExternalLinkService">
<mods:mods id="B6427A0D36FDE1DD3A74896BF3543F2A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0AB5F0F64EBCABD6287782F181B17A9">
<mods:title id="362E652DC872601C28358F7435201748">New species of the genus Chydaeus from China, Nepal, Myanmar, and Thailand, with remarks on species previously described (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
<mods:name id="809FB861457E6F292AF19ED5F0B648C5" type="personal">
<mods:role id="ADD5B923176DC90F68843E598154343C">
<mods:roleTerm id="4C6C5BD3FE5C1FF4F5D13775BB20C807">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="472E93D2D4F4C3CA3EE7A60743A3522A">Kataev, Boris M.</mods:namePart>
<mods:affiliation id="F5B705EB17D661F4B48AF3F79D68B94C">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
<mods:nameIdentifier id="12255DCC230FA4C69AB4B6766C944284" type="email">harpal@zin.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="C4F98AAF4D49525CE0D0A002ACC3D773" type="personal">
<mods:role id="4183D77F9394545089942EB08FAEEE7B">
<mods:roleTerm id="9E694B0B3C00E4462DD37A5F636FED01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF2CFD1BB066F361929B8AD12E5D1A15">Wrase, David W.</mods:namePart>
<mods:affiliation id="A4C4157AD60FC5132080E9D9A5D8FDE7">Dunckerstrasse 78, D- 10437 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="24A9FA4152289FD546F0BA0DC48958E7" type="email">carterus@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="56E7CAFB7E1F68BC7BD876AE8BE3B373" type="personal">
<mods:role id="2511A6754A66F9069513A5BF6BE6E575">
<mods:roleTerm id="85386143F131FEFDAE5A3176637CDB5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35374358262E45D4ABAFD2772AF04C30">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="BBEC1B3863515269AD80F098A14B0AEF">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitätsplatz 2, D – 18055 Rostock, Germany.</mods:affiliation>
<mods:nameIdentifier id="DF40759F0E225D0178FDA40D44A41B20" type="email">schmidt@agonum.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="FF1510A97441D264EC520D3FB4A6B77D">text</mods:typeOfResource>
<mods:relatedItem id="EE76B1D208F9B10313C80DB81DE3072D" type="host">
<mods:titleInfo id="F5935E56E11748FA5A0988D3F3AE54BF">
<mods:title id="A08495D2257520289AF4B72B8D93380B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DFC8972C17CDFD9F95CCC3F614AC4A85">
<mods:date id="C8F8E81A50D46A3058AADFB7AC6CABCE">2014</mods:date>
<mods:detail id="F7314FD411C445D5C47E3009E996D271" type="pubDate">
<mods:number id="242135CDDEAA6C3B360A8868797FE320">2014-02-14</mods:number>
</mods:detail>
<mods:detail id="13C5DD7B1A43C4AEB59CA42C52D00EE7" type="volume">
<mods:number id="5639488607358426CC79AE31E86B471C">3765</mods:number>
</mods:detail>
<mods:detail id="B430A0E965DFDA3D44373ECD9E1EEC13" type="issue">
<mods:number id="53EEECE6E126BC47A1819D192E4EF9C9">1</mods:number>
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<mods:start id="3133C2F09BDEA08F5A43A5676260023F">1</mods:start>
<mods:end id="C9E9265CD2972E74724D6E8B4BFBBF79">28</mods:end>
</mods:extent>
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<mods:classification id="936F111DFF16DE0BF384BBD7C74B056B">journal article</mods:classification>
<mods:identifier id="33AFE72B4F9E3B71F781360B924C8A64" type="CLB-Dataset">36901</mods:identifier>
<mods:identifier id="43FAD342750D0DD34B41CDD05233FC7B" type="DOI">10.11646/zootaxa.3765.1.1</mods:identifier>
<mods:identifier id="1E7C57507D2339DD9C1A337B0586787D" type="GBIF-Dataset">dc48dc34-f0b6-4715-8044-1aa2d5851737</mods:identifier>
<mods:identifier id="9FF07E2D24144C5388E6D2A5447099EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B6D57AC8AD0F1432A8258CDE6F985F85" type="Zenodo-Dep">285551</mods:identifier>
<mods:identifier id="E258310EDCA47DFE310BE2CDED014E35" type="ZooBank">4776DD1C-9FCC-4F05-8FDF-66AA14B959FC</mods:identifier>
</mods:mods>
<treatment id="232FAA28FFC9021AFF53FD42FD23F850" ID-DOI="http://doi.org/10.5281/zenodo.6138884" ID-GBIF-Taxon="127667022" ID-Zenodo-Dep="6138884" LSID="urn:lsid:plazi:treatment:232FAA28FFC9021AFF53FD42FD23F850" httpUri="http://treatment.plazi.org/id/232FAA28FFC9021AFF53FD42FD23F850" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="E39C48B5FFC90204FF53FD42FE41FBE2" box="[151,479,727,754]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="AB391B3EFFC90204FF53FD42FE41FBE2" blockId="15.[151,479,727,787]" box="[151,479,727,754]" pageId="15" pageNumber="16">
<heading id="F071AC52FFC90204FF53FD42FE41FBE2" bold="true" box="[151,479,727,754]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="99F2C72CFFC90204FF53FD42FE41FBE2" bold="true" box="[151,479,727,754]" pageId="15" pageNumber="16">
<taxonomicName id="6C8660BDFFC90204FF53FD42FE0BFBE1" ID-CoL="V44S" authority="KATAEV ET AL." box="[151,405,727,753]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFC90204FF53FD42FE0BFBE1" bold="true" box="[151,405,727,753]" italics="true" pageId="15" pageNumber="16">Chydaeus putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFC90204FE58FD4DFE41FBE2" box="[412,479,728,754]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E39C48B5FFC90204FF53FD6FFEDDFA03" box="[151,323,762,787]" pageId="15" pageNumber="16" type="description">
<paragraph id="AB391B3EFFC90204FF53FD6FFEDDFA03" blockId="15.[151,479,727,787]" box="[151,323,762,787]" pageId="15" pageNumber="16">
(
<figureCitation id="33BD07BBFFC90204FF5BFD6FFF79FA03" box="[159,231,762,787]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,263,1971,1994]" captionTargetBox="[211,1370,549,1955]" captionTargetId="figure@3.[206,1381,541,1964]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Chydaeus, dorsal habitus (holotypes). 1, C. dissimilis sp. n.; 2, C. shaanxiensis sp. n.; 3, C. kabaki sp. n.; 4, C. weishanensis sp. n.; 5, C. belousovi sp. n.; 6, C. putaoensis sp. n. Scales = 2.0 mm." httpUri="https://zenodo.org/record/285552/files/figure.png" pageId="15" pageNumber="16">Figs 6</figureCitation>
,
<figureCitation id="33BD07BBFFC90204FF37FD6EFEA5FA03" box="[243,315,763,787]" captionStart="FIGURES 44 – 48" captionStartId="16.[151,263,1264,1287]" captionTargetBox="[218,1368,449,1242]" captionTargetId="figure@16.[213,1373,445,1243]" captionTargetPageId="16" captionText="FIGURES 44 – 48. Chydaeus putaoensis sp. n. (44 – 46, holotype; 47 – 48, paratype). 44, left metepisternum; 45 – 46, median lobe of aedeagus; 47, laterotergite IX and stylus; 48, stylus. 44, 47, ventral aspect; 45, 48, lateral aspect; 46, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/285557/files/figure.png" pageId="15" pageNumber="16">44–48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E39C48B5FFC90204FF53FCD7FCEBFA6F" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="AB391B3EFFC90204FF53FCD7FAD5FA4B" blockId="15.[151,1437,834,2011]" box="[151,1355,834,859]" pageId="15" pageNumber="16">
<emphasis id="99F2C72CFFC90204FF53FCD7FEA3FA4B" bold="true" box="[151,317,834,859]" pageId="15" pageNumber="16">
<typeStatus id="743DA59CFFC90204FF53FCD7FF4CFA4B" box="[151,210,834,859]" pageId="15" pageNumber="16">Type</typeStatus>
material
</emphasis>
.
<materialsCitation id="1BEE1163FFC90204FE88FCD7FAD5FA4B" ID-GBIF-Occurrence="3467093359" box="[332,1355,834,859]" collectingDate="2006-12-08" collectorName="N BURMA" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="743DA59CFFC90204FE88FCD7FE27FA4B" box="[332,441,834,859]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BD80D0B7FFC90204FE07FCD6FE44FA4A" box="[451,474,835,858]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
, &quot;
<collectorName id="06737EE8FFC90204FE2BFCD6FDECFA4B" box="[495,626,835,859]" pageId="15" pageNumber="16">N BURMA</collectorName>
leg
<locationDeviation id="BA8B021CFFC90204FD67FCD6FBC1FA4B" box="[675,1119,834,859]" pageId="15" pageNumber="16">
Reuter ca.
<quantity id="6C7EB6DBFFC90204FCDFFCD7FCFCFA4B" box="[795,866,834,859]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="km" value="60.0">60 km</quantity>
NW Putao 2200mNN
</locationDeviation>
, 7+
<date id="DF383DFEFFC90204FB42FCD7FB64FA4B" box="[1158,1274,834,859]" pageId="15" pageNumber="16" value="2006-12-08">
<collectingDate id="CF7CC416FFC90204FB42FCD7FB64FA4B" box="[1158,1274,834,859]" pageId="15" pageNumber="16" value="2006-12-08">12.8.2006</collectingDate>
</date>
&quot; (cJS).
</materialsCitation>
</paragraph>
<paragraph id="AB391B3EFFC90204FF03FCF2FCEBFA6F" blockId="15.[151,1437,834,2011]" box="[199,885,870,895]" pageId="15" pageNumber="16">
<materialsCitation id="1BEE1163FFC90204FF03FCF2FCEBFA6F" ID-GBIF-Occurrence="3467093328" box="[199,885,870,895]" collectingDate="2006-12-08" collectionCode="ZIN" collectorName="N BURMA" pageId="15" pageNumber="16" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="743DA59CFFC90204FF03FCF2FEA6FA6F" box="[199,312,871,895]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BD80D0B7FFC90204FE87FCF2FEF0FA6E" box="[323,366,871,895]" pageId="15" pageNumber="16" type="male">2 ♂</specimenCount>
,
<specimenCount id="BD80D0B7FFC90204FEBFFCF3FE3AFA6E" box="[379,420,870,895]" pageId="15" pageNumber="16" type="female">5 ♀</specimenCount>
, same data as holotype (cJS, cWR,
<collectionCode id="CD9783FBFFC90204FCF2FCF2FCF7FA6F" box="[822,873,871,895]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="15" pageNumber="16">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E39C48B5FFC9021BFF03FC1FFD69F898" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="AB391B3EFFC90204FF03FC1FFDC7FAB3" blockId="15.[151,1437,834,2011]" box="[199,601,906,931]" pageId="15" pageNumber="16">
<emphasis id="99F2C72CFFC90204FF03FC1FFECFFAB3" bold="true" box="[199,337,906,931]" pageId="15" pageNumber="16">Description</emphasis>
. Dorsal habitus:
<figureCitation id="33BD07BBFFC90204FDD5FC1FFDCBFAB3" box="[529,597,906,931]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,263,1971,1994]" captionTargetBox="[211,1370,549,1955]" captionTargetId="figure@3.[206,1381,541,1964]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Chydaeus, dorsal habitus (holotypes). 1, C. dissimilis sp. n.; 2, C. shaanxiensis sp. n.; 3, C. kabaki sp. n.; 4, C. weishanensis sp. n.; 5, C. belousovi sp. n.; 6, C. putaoensis sp. n. Scales = 2.0 mm." httpUri="https://zenodo.org/record/285552/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03FC3BFB48FAD7" blockId="15.[151,1437,834,2011]" box="[199,1238,942,967]" pageId="15" pageNumber="16">
Size. Body length 9.0–
<quantity id="6C7EB6DBFFC90204FE0CFC3AFD81FAD6" box="[456,543,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="16" unit="mm" value="9.8">9.8 mm</quantity>
, width
<quantity id="6C7EB6DBFFC90204FDB5FC3AFD65FAD6" box="[625,763,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.9" metricValueMin="4.4" pageId="15" pageNumber="16" unit="mm" value="4.65" valueMax="4.9" valueMin="4.4">4.4–4.9 mm</quantity>
; in
<typeStatus id="743DA59CFFC90204FCE7FC3BFC1BFAD7" box="[803,901,942,967]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
9.1 and
<quantity id="6C7EB6DBFFC90204FC22FC3AFBA3FAD6" box="[998,1085,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="15" pageNumber="16" unit="mm" value="4.4">4.4 mm</quantity>
, respectively.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03FC47FBD6FAFB" blockId="15.[151,1437,834,2011]" box="[199,1096,978,1003]" pageId="15" pageNumber="16">
Morphological characters as described for
<taxonomicName id="6C8660BDFFC90204FD61FC46FCAEFAFB" box="[677,816,979,1003]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFC90204FD61FC46FCAEFAFB" box="[677,816,979,1003]" italics="true" pageId="15" pageNumber="16">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFC90204FCFCFC47FCE9FAFB" box="[824,887,978,1003]" pageId="15" pageNumber="16" rank="species">
<emphasis id="99F2C72CFFC90204FCFCFC47FCE9FAFB" bold="true" box="[824,887,978,1003]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
except as follows:
</paragraph>
<paragraph id="AB391B3EFFC90204FF03FC63FD1CFD23" blockId="15.[151,1437,834,2011]" pageId="15" pageNumber="16">Color. Generally slightly darker, often antennomeres partly, tibiae at least apically, tarsi throughout and occasionally also palpi somewhat infuscate.</paragraph>
<paragraph id="AB391B3EFFC90204FF03FBABFAF3FD6B" blockId="15.[151,1437,834,2011]" pageId="15" pageNumber="16">Microsculpture. Elytra in both sexes with distinct microsculpture consisting of isodiametric meshes along basal border and along sides, also on intervals 8 and 9, and of weakly transverse meshes on remaining surface.</paragraph>
<paragraph id="AB391B3EFFC90204FF03FB13FB36FC3F" blockId="15.[151,1437,834,2011]" pageId="15" pageNumber="16">
Head. Comparatively large (HWmax/PWmax = 0.65–0.69 and HWmin/PWmax = 0.57–0.61), more convex on clypeus and frons than in
<taxonomicName id="6C8660BDFFC90204FE7CFB3EFDDAFDD3" box="[440,580,1195,1219]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFC90204FE7CFB3EFDDAFDD3" box="[440,580,1195,1219]" italics="true" pageId="15" pageNumber="16">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFC90204FD8FFB3FFD15FDD3" box="[587,651,1194,1219]" pageId="15" pageNumber="16" rank="species">
<emphasis id="99F2C72CFFC90204FD8FFB3FFD15FDD3" bold="true" box="[587,651,1194,1219]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
, with deeper frontal foveae and clypeo-frontal suture. Eyes (HWmax/ HWmin = 0.10–0.19) slightly smaller. Labrum more shallowly emarginate apically. Antennae longer, in male surpassing pronotal basal margin by approximately one apical antennomere, in female only reaching to pronotal basal edge; in both sexes with antennomeres 5 to 7 each about 2.0–2.3 times as long as wide.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03FAAEFF72FF3B" blockId="15.[151,1437,834,2011]" pageId="15" pageNumber="16">
Pronotum. More transverse (PWmax/PL = 1.56–1.65), widest at beginning of second third, with slightly wider base (PWmax/PWmin = 1.19–1.25) than in
<taxonomicName id="6C8660BDFFC90204FD42FACAFC8CFC66" box="[646,786,1374,1398]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFC90204FD42FACAFC8CFC66" box="[646,786,1374,1398]" italics="true" pageId="15" pageNumber="16">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFC90204FCDEFACBFCC4FC67" box="[794,858,1374,1399]" pageId="15" pageNumber="16" rank="species">
<emphasis id="99F2C72CFFC90204FCDEFACBFCC4FC67" bold="true" box="[794,858,1374,1399]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
; basal margin noticeably wider than apical margin. Sides usually more distinctly sinuate in posterior quarter. Basal angles less obtuse, slightly more than 90°, lacking denticle at apex. Lateral depressions usually discernible from apical angles to basal margin. Basal foveae deeper and narrower, reaching pronotal basal margin; area at basal angles between basal foveae and lateral depressions somewhat convex. Pronotal surface rather densely punctate along margins; punctures coarsest and confluent laterobasally.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03F9A3FDE4FE13" blockId="15.[151,1437,834,2011]" pageId="15" pageNumber="16">
Elytra. Wider and shorter [EL/EW = 1.20–1.29 (
<quantity id="6C7EB6DBFFC90204FD2BF9A3FCE0FF5F" box="[751,894,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.2131" metricValueMax="3.2258" metricValueMin="3.2004" pageId="15" pageNumber="16" unit="in" value="1.265" valueMax="1.27" valueMin="1.26">1.26–1.27 in</quantity>
male and
<quantity id="6C7EB6DBFFC90204FC32F9A2FB1AFF5F" box="[1014,1156,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.1623" metricValueMax="3.2766" metricValueMin="3.048" pageId="15" pageNumber="16" unit="in" value="1.245" valueMax="1.29" valueMin="1.2">1.20–1.29 in</quantity>
female), EL/PL = 2.55– 2.66 (
<quantity id="6C7EB6DBFFC90204FF1CF9CFFEF6FF63" box="[216,360,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="6.5405" metricValueMax="6.604" metricValueMin="6.476999999999999" pageId="15" pageNumber="16" unit="in" value="2.575" valueMax="2.6" valueMin="2.55">2.55–2.60 in</quantity>
male and
<quantity id="6C7EB6DBFFC90204FE1BF9CFFDF3FF63" box="[479,621,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="6.6421" metricValueMax="6.7564" metricValueMin="6.5278" pageId="15" pageNumber="16" unit="in" value="2.615" valueMax="2.66" valueMin="2.57">2.57–2.66 in</quantity>
female), EW/PWmax = 1.26–1.34 (
<quantity id="6C7EB6DBFFC90204FBC3F9CFFB09FF63" box="[1031,1175,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="3.2384999999999997" metricValueMax="3.2766" metricValueMin="3.2004" pageId="15" pageNumber="16" unit="in" value="1.275" valueMax="1.29" valueMin="1.26">1.26–1.29 in</quantity>
male and
<quantity id="6C7EB6DBFFC90204FACAF9CFFA02FF63" box="[1294,1436,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" metricValueMax="3.4036" metricValueMin="3.2004" pageId="15" pageNumber="16" unit="in" value="1.3" valueMax="1.34" valueMin="1.26">1.26–1.34 in</quantity>
female)], fused along suture, each with a tiny humeral denticle at apex visible from behind. Basal borders slightly more markedly sinuate laterally, joined each with lateral margin at more distinct obtuse angle. In males striae slightly more impressed and intervals on disc slightly more convex than in males of
<taxonomicName id="6C8660BDFFC90204FB77F952FAD8FFCE" box="[1203,1350,1734,1758]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFC90204FB77F952FAD8FFCE" box="[1203,1350,1734,1758]" italics="true" pageId="15" pageNumber="16">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFC90204FA91F953FA02FFCF" box="[1365,1436,1734,1759]" pageId="15" pageNumber="16" rank="species">
<emphasis id="99F2C72CFFC90204FA91F953FA02FFCF" bold="true" box="[1365,1436,1734,1759]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
Parascutellar basal setigerous pores absent.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03F89BFB90FE37" blockId="15.[151,1437,834,2011]" box="[199,1038,1806,1831]" pageId="15" pageNumber="16">
Ventral side. Metepisterna (
<figureCitation id="33BD07BBFFC90204FE3FF89BFDD3FE37" box="[507,589,1806,1831]" captionStart="FIGURES 44 – 48" captionStartId="16.[151,263,1264,1287]" captionTargetBox="[218,1368,449,1242]" captionTargetId="figure@16.[213,1373,445,1243]" captionTargetPageId="16" captionText="FIGURES 44 – 48. Chydaeus putaoensis sp. n. (44 – 46, holotype; 47 – 48, paratype). 44, left metepisternum; 45 – 46, median lobe of aedeagus; 47, laterotergite IX and stylus; 48, stylus. 44, 47, ventral aspect; 45, 48, lateral aspect; 46, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/285557/files/figure.png" pageId="15" pageNumber="16">Fig. 44</figureCitation>
) transverse, noticeably wider than long.
</paragraph>
<paragraph id="AB391B3EFFC90204FF03F8A7FD17FE5B" blockId="15.[151,1437,834,2011]" box="[199,649,1842,1867]" pageId="15" pageNumber="16">
<collectingCountry id="D3915BAEFFC90204FF03F8A7FF61FE5B" box="[199,255,1842,1867]" name="India" pageId="15" pageNumber="16">Hind</collectingCountry>
wings reduced to very small stubs.
</paragraph>
<paragraph id="AB391B3EFFC9021BFF03F8C2FD97F9C5" blockId="15.[151,1437,834,2011]" lastBlockId="16.[151,1436,151,392]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Legs. Metacoxa each with additional posteromedial setigerous pore and without additional medial setigerous or unsetigerous pores. Tarsomere 5 with two, occasionally three pairs of lateroventral setae. Metatarsi in male about as long as and in female slightly shorter than HWmin. In male, protarsi markedly enlarged, with tarsomere 2 only slightly narrower than protibia at apical margin (tarsomeres 1–4 with adhesive vestiture ventrally); mesotarsi moderately enlarged, narrower than mesotibia at apical margin, with tarsomeres 2 and 3 about as long as wide and with adhesive vestiture ventrally.</paragraph>
<paragraph id="AB391B3EFFD6021BFF03FF4AFDBCF80C" blockId="16.[151,1436,151,392]" pageId="16" pageNumber="17">
Female genitalia (
<figureCitation id="33BD07BBFFD6021BFE50FF4AFD87F9E8" box="[404,537,223,248]" captionStart="FIGURES 44 – 48" captionStartId="16.[151,263,1264,1287]" captionTargetBox="[218,1368,449,1242]" captionTargetId="figure@16.[213,1373,445,1243]" captionTargetPageId="16" captionText="FIGURES 44 – 48. Chydaeus putaoensis sp. n. (44 – 46, holotype; 47 – 48, paratype). 44, left metepisternum; 45 – 46, median lobe of aedeagus; 47, laterotergite IX and stylus; 48, stylus. 44, 47, ventral aspect; 45, 48, lateral aspect; 46, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/285557/files/figure.png" pageId="16" pageNumber="17">Figs 47–48</figureCitation>
). Apical stylomere rather strongly widened in middle portion in lateral aspect, somewhat curved in ventral aspect.
</paragraph>
<paragraph id="AB391B3EFFD6021BFF03FEB2FD69F898" blockId="16.[151,1436,151,392]" pageId="16" pageNumber="17">
Aedeagus. Median lobe (
<figureCitation id="33BD07BBFFD6021BFE22FEB2FDF7F850" box="[486,617,295,320]" captionStart="FIGURES 44 – 48" captionStartId="16.[151,263,1264,1287]" captionTargetBox="[218,1368,449,1242]" captionTargetId="figure@16.[213,1373,445,1243]" captionTargetPageId="16" captionText="FIGURES 44 – 48. Chydaeus putaoensis sp. n. (44 – 46, holotype; 47 – 48, paratype). 44, left metepisternum; 45 – 46, median lobe of aedeagus; 47, laterotergite IX and stylus; 48, stylus. 44, 47, ventral aspect; 45, 48, lateral aspect; 46, dorsal aspect. Scale = 1.0 mm." httpUri="https://zenodo.org/record/285557/files/figure.png" pageId="16" pageNumber="17">Figs 45–46</figureCitation>
) robust, but narrower in dorsal aspect than in
<taxonomicName id="6C8660BDFFD6021BFB45FEBDFA91F850" box="[1153,1295,296,320]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFB45FEBDFA91F850" box="[1153,1295,296,320]" italics="true" pageId="16" pageNumber="17">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFADCFEB2FAC4F850" box="[1304,1370,295,320]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFADCFEB2FAC4F850" bold="true" box="[1304,1370,295,320]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
, with relatively smaller basal bulb; ventral side convex up to apex; terminal lamella markedly curved dorsad. Internal sac with several folded formations covered by tiny spines.
</paragraph>
</subSubSection>
<caption id="FFF94BB6FFD6021BFF53FB65FE9EFC54" httpUri="https://zenodo.org/record/285557/files/figure.png" pageId="16" pageNumber="17" targetBox="[218,1368,449,1242]" targetPageId="16">
<paragraph id="AB391B3EFFD6021BFF53FB65FE9EFC54" blockId="16.[151,1436,1264,1348]" pageId="16" pageNumber="17">
<emphasis id="99F2C72CFFD6021BFF53FB65FEC6FC16" bold="true" box="[151,344,1264,1287]" pageId="16" pageNumber="17">FIGURES 44–48.</emphasis>
<taxonomicName id="6C8660BDFFD6021BFEA4FB64FDA7FC17" box="[352,569,1265,1287]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFEA4FB64FDA7FC17" box="[352,569,1265,1287]" italics="true" pageId="16" pageNumber="17">Chydaeus putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFD86FB64FDE3FC16" box="[578,637,1265,1286]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFD86FB64FDE3FC16" bold="true" box="[578,637,1265,1286]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
(44–46, holotype; 47–48, paratype). 44, left metepisternum; 45–46, median lobe of aedeagus; 47, laterotergite IX and stylus; 48, stylus. 44, 47, ventral aspect; 45, 48, lateral aspect; 46, dorsal aspect. Scale = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="E39C48B5FFD6021BFF03FAE4FEE3FCBD" pageId="16" pageNumber="17" type="etymology">
<paragraph id="AB391B3EFFD6021BFF03FAE4FEE3FCBD" blockId="16.[151,1436,1393,2030]" pageId="16" pageNumber="17">
<emphasis id="99F2C72CFFD6021BFF03FAE4FED8FC9A" bold="true" box="[199,326,1393,1418]" pageId="16" pageNumber="17">Etymology</emphasis>
. The specific name refers to Putao, a city in
<collectingCountry id="D3915BAEFFD6021BFCF3FAE7FC38FC9A" box="[823,934,1394,1418]" name="Myanmar" pageId="16" pageNumber="17">Myanmar</collectingCountry>
, in which mountainous environment the
<typeStatus id="743DA59CFFD6021BFAA8FAE7FA02FC9A" box="[1388,1436,1394,1418]" pageId="16" pageNumber="17">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="E39C48B5FFD6021AFF03FA2CFAE2F9E8" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="AB391B3EFFD6021BFF03FA2CFA02FEFC" blockId="16.[151,1436,1393,2030]" pageId="16" pageNumber="17">
<emphasis id="99F2C72CFFD6021BFF03FA2CFE4EFCC2" bold="true" box="[199,464,1465,1490]" pageId="16" pageNumber="17">Comparative remarks</emphasis>
. This new species is very similar and apparently closely related to
<taxonomicName id="6C8660BDFFD6021BFB0CFA2CFACAFCC1" box="[1224,1364,1465,1489]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFB0CFA2CFACAFCC1" box="[1224,1364,1465,1489]" italics="true" pageId="16" pageNumber="17">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFA98FA2CFA02FCC2" box="[1372,1436,1465,1490]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFA98FA2CFA02FCC2" bold="true" box="[1372,1436,1465,1490]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
described above. Both the species share many morphological features, including an almost acute, only slightly blunted apex of left mandible and an unusual position of the supraorbital setigerous pores which are removed rather far from the supraorbital furrows.
<taxonomicName id="6C8660BDFFD6021BFDDDF9B0FC99FF2D" box="[537,775,1573,1597]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFDDDF9B0FC99FF2D" box="[537,775,1573,1597]" italics="true" pageId="16" pageNumber="17">Chydaeus putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFCCBF9B0FCCEFF2E" box="[783,848,1573,1598]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFCCBF9B0FCCEFF2E" bold="true" box="[783,848,1573,1598]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
distinctly differs from
<taxonomicName id="6C8660BDFFD6021BFB92F9B0FB7CFF2D" box="[1110,1250,1573,1597]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFB92F9B0FB7CFF2D" box="[1110,1250,1573,1597]" italics="true" pageId="16" pageNumber="17">C. belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFB2EF9B0FAB5FF2E" box="[1258,1323,1573,1598]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFB2EF9B0FAB5FF2E" bold="true" box="[1258,1323,1573,1598]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
in having the antennae longer, the frontal foveae deeper, the pronotum and elytra wider, the parascutellar setigerous pore absent, the pro- and mesotarsi in male more enlarged, and the median lobe of the aedeagus with apex distinctly curved dorsad. Additional distinctive features of
<taxonomicName id="6C8660BDFFD6021BFD08F904FCF8FFB9" box="[716,870,1681,1705]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFD08F904FCF8FFB9" box="[716,870,1681,1705]" italics="true" pageId="16" pageNumber="17">C. putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFCB5F904FC2DFFBA" box="[881,947,1681,1706]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFCB5F904FC2DFFBA" bold="true" box="[881,947,1681,1706]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
are listed in the description. Although we tentatively include this new species in the
<taxonomicName id="6C8660BDFFD6021BFD52F920FC98FFDD" box="[662,774,1717,1741]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kasaharai">
<emphasis id="99F2C72CFFD6021BFD52F920FC98FFDD" box="[662,774,1717,1741]" italics="true" pageId="16" pageNumber="17">kasaharai</emphasis>
</taxonomicName>
group, in fact it is morphologically and geographically intermediate between this less specialized group of species distributed in
<collectingCountry id="D3915BAEFFD6021BFC26F94CFBBBFFE2" box="[994,1061,1753,1778]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
and the more specialized
<taxonomicName id="6C8660BDFFD6021BFA97F94FFA02FFE1" box="[1363,1436,1754,1777]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="irvinei">
<emphasis id="99F2C72CFFD6021BFA97F94FFA02FFE1" box="[1363,1436,1754,1777]" italics="true" pageId="16" pageNumber="17">irvinei</emphasis>
</taxonomicName>
species group which is adapted to the high altitude environments of the Himalaya. Like all members of the
<taxonomicName id="6C8660BDFFD6021BFA97F96BFA02FE05" box="[1363,1436,1790,1813]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="irvinei">
<emphasis id="99F2C72CFFD6021BFA97F96BFA02FE05" box="[1363,1436,1790,1813]" italics="true" pageId="16" pageNumber="17">irvinei</emphasis>
</taxonomicName>
species group,
<taxonomicName id="6C8660BDFFD6021BFE88F8B4FE72FE29" box="[332,492,1825,1849]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFE88F8B4FE72FE29" box="[332,492,1825,1849]" italics="true" pageId="16" pageNumber="17">C. putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFE3EF8B4FDDFFE2A" box="[506,577,1825,1850]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFE3EF8B4FDDFFE2A" bold="true" box="[506,577,1825,1850]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
lacks a parascutellar setigerous pore; however, this new species has a parascutellar striole which is present in all members of the
<taxonomicName id="6C8660BDFFD6021BFCECF8D0FC06FE4D" box="[808,920,1861,1885]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kasaharai">
<emphasis id="99F2C72CFFD6021BFCECF8D0FC06FE4D" box="[808,920,1861,1885]" italics="true" pageId="16" pageNumber="17">kasaharai</emphasis>
</taxonomicName>
species group and absent or only suggested in the members of the
<taxonomicName id="6C8660BDFFD6021BFEBFF8FFFE5AFE91" box="[379,452,1898,1921]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="irvinei">
<emphasis id="99F2C72CFFD6021BFEBFF8FFFE5AFE91" box="[379,452,1898,1921]" italics="true" pageId="16" pageNumber="17">irvinei</emphasis>
</taxonomicName>
group. Interestingly,
<taxonomicName id="6C8660BDFFD6021BFD73F8FCFCCFFE91" box="[695,849,1897,1921]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD6021BFD73F8FCFCCFFE91" box="[695,849,1897,1921]" italics="true" pageId="16" pageNumber="17">C. putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD6021BFC9EF8FCFC04FE92" box="[858,922,1897,1922]" pageId="16" pageNumber="17" rank="species">
<emphasis id="99F2C72CFFD6021BFC9EF8FCFC04FE92" bold="true" box="[858,922,1897,1922]" pageId="16" pageNumber="17">sp. n.</emphasis>
</taxonomicNameLabel>
shares some characters with some species of the
<taxonomicName id="6C8660BDFFD6021BFF06F81BFE95FEB5" box="[194,267,1934,1957]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="irvinei">
<emphasis id="99F2C72CFFD6021BFF06F81BFE95FEB5" box="[194,267,1934,1957]" italics="true" pageId="16" pageNumber="17">irvinei</emphasis>
</taxonomicName>
group (for example,
<taxonomicName id="6C8660BDFFD6021BFDC4F818FC1EFEB6" authority="Kataev &amp; Schmidt" authorityName="Kataev &amp; Schmidt" box="[512,896,1933,1958]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="99F2C72CFFD6021BFDC4F818FD3AFEB5" box="[512,676,1933,1957]" italics="true" pageId="16" pageNumber="17">C. ovalipennis</emphasis>
Kataev &amp; Schmidt
</taxonomicName>
and
<taxonomicName id="6C8660BDFFD6021BFC7FF818FA82FEB6" authority="Kataev &amp; Schmidt" authorityName="Kataev &amp; Schmidt" box="[955,1308,1933,1958]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="99F2C72CFFD6021BFC7FF818FBA3FEB5" box="[955,1085,1933,1957]" italics="true" pageId="16" pageNumber="17">C. minimus</emphasis>
Kataev &amp; Schmidt
</taxonomicName>
) which are unknown outside the
<taxonomicName id="6C8660BDFFD6021BFE48F827FE4BFED9" box="[396,469,1970,1993]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="irvinei">
<emphasis id="99F2C72CFFD6021BFE48F827FE4BFED9" box="[396,469,1970,1993]" italics="true" pageId="16" pageNumber="17">irvinei</emphasis>
</taxonomicName>
group: elytra fused along suture, tarsomere 5 usually with two pairs of lateroventral setae and apex of left mandible not truncate, only slightly blunted (the latter character is also present in
<emphasis id="99F2C72CFFD6021BFA46F840FA02FEFC" box="[1410,1436,2005,2028]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
</paragraph>
<paragraph id="AB391B3EFFD7021AFF53FF0DFAE2F9E8" blockId="17.[151,1436,151,320]" pageId="17" pageNumber="18">
<taxonomicName id="6C8660BDFFD7021AFF53FF0DFE9CF9A0" box="[151,258,152,176]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="belousovi" status="sp. nov.">
<emphasis id="99F2C72CFFD7021AFF53FF0DFE9CF9A0" box="[151,258,152,176]" italics="true" pageId="17" pageNumber="18">belousovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD7021AFECFFF02FED2F9A0" box="[267,332,151,176]" pageId="17" pageNumber="18" rank="species">
<emphasis id="99F2C72CFFD7021AFECFFF02FED2F9A0" bold="true" box="[267,332,151,176]" pageId="17" pageNumber="18">sp. n.</emphasis>
</taxonomicNameLabel>
of the
<taxonomicName id="6C8660BDFFD7021AFE5AFF0DFD90F9A0" box="[414,526,152,176]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kasaharai">
<emphasis id="99F2C72CFFD7021AFE5AFF0DFD90F9A0" box="[414,526,152,176]" italics="true" pageId="17" pageNumber="18">kasaharai</emphasis>
</taxonomicName>
group). Both
<taxonomicName id="6C8660BDFFD7021AFD6BFF0DFCCCF9A0" box="[687,850,152,176]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="99F2C72CFFD7021AFD6BFF0DFCCCF9A0" box="[687,850,152,176]" italics="true" pageId="17" pageNumber="18">C. ovalipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="6C8660BDFFD7021AFC48FF0DFB93F9BF" box="[908,1037,152,175]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="99F2C72CFFD7021AFC48FF0DFB93F9BF" box="[908,1037,152,175]" italics="true" pageId="17" pageNumber="18">C. minimus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="6C8660BDFFD7021AFB5FFF0DFAAAF9BF" box="[1179,1332,152,175]" class="Insecta" family="Carabidae" genus="Chydaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="putaoensis" status="sp. nov.">
<emphasis id="99F2C72CFFD7021AFB5FFF0DFAAAF9BF" box="[1179,1332,152,175]" italics="true" pageId="17" pageNumber="18">C. putaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82C17A57FFD7021AFAF9FF02FAE0F9A0" box="[1341,1406,151,176]" pageId="17" pageNumber="18" rank="species">
<emphasis id="99F2C72CFFD7021AFAF9FF02FAE0F9A0" bold="true" box="[1341,1406,151,176]" pageId="17" pageNumber="18">sp. n.</emphasis>
</taxonomicNameLabel>
in having the body smaller (
<quantity id="6C7EB6DBFFD7021AFE0BFF29FDFDF9C4" box="[463,611,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.6" metricValueMin="5.5" pageId="17" pageNumber="18" unit="mm" value="6.55" valueMax="7.6" valueMin="5.5">5.5–7.6 mm</quantity>
), the parascutellar striole absent or rudimentary and the supraorbital setigerous pore on head situated at a distance of about two or three diameters of pore from supraorbital furrows.
</paragraph>
</subSubSection>
<subSubSection id="E39C48B5FFD7021AFF03FE91FD23F850" pageId="17" pageNumber="18" type="distribution">
<paragraph id="AB391B3EFFD7021AFF03FE91FD23F850" blockId="17.[151,1436,151,320]" pageId="17" pageNumber="18">
<emphasis id="99F2C72CFFD7021AFF03FE91FEC7F80D" bold="true" box="[199,345,260,285]" pageId="17" pageNumber="18">Distribution</emphasis>
. Known only from the
<typeStatus id="743DA59CFFD7021AFD41FE90FD2BF80D" box="[645,693,261,285]" pageId="17" pageNumber="18">type</typeStatus>
locality (about
<quantity id="6C7EB6DBFFD7021AFCB8FE91FC55F80D" box="[892,971,260,285]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="km" value="60.0">60 km</quantity>
north-west of Putao,
<quantity id="6C7EB6DBFFD7021AFB1EFE91FAA4F80C" box="[1242,1338,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="m" value="2200.0">2200 m</quantity>
), in the northernmost part of the Kachin State,
<collectingCountry id="D3915BAEFFD7021AFD8FFEBDFD24F850" box="[587,698,296,320]" name="Myanmar" pageId="17" pageNumber="18">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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