342 lines
35 KiB
XML
342 lines
35 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E786BFFA8FF013D5DACF4E870" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Hylekobolus goodmani Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="C097B4BE-0F42-4C7D-B087-4C7B4396C826" docUuidSource="ZooBank" docVersion="10" lastPageNumber="111" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="109" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791477" ID-GBIF-Taxon="163747608" ID-Zenodo-Dep="3791477" LSID="urn:lsid:zoobank.org:act:C097B4BE-0F42-4C7D-B087-4C7B4396C826" httpUri="http://treatment.plazi.org/id/1772122E786BFFA8FF013D5DACF4E870" lastPageId="110" lastPageNumber="111" pageId="108" pageNumber="109">
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||
<subSubSection box="[140,595,553,580]" pageId="108" pageNumber="109" type="nomenclature">
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||
<paragraph blockId="108.[140,975,553,615]" box="[140,595,553,580]" pageId="108" pageNumber="109">
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||
<heading box="[140,595,553,580]" fontSize="11" level="2" pageId="108" pageNumber="109" reason="3">
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<emphasis bold="true" box="[140,595,553,580]" pageId="108" pageNumber="109">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,518,553,580]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,410,553,580]" italics="true" pageId="108" pageNumber="109">Hylekobolus goodmani</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[530,595,554,580]" pageId="108" pageNumber="109" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,975,589,615]" pageId="108" pageNumber="109" type="description">
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<paragraph blockId="108.[140,975,553,615]" box="[140,975,589,615]" pageId="108" pageNumber="109">
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<uri box="[140,975,589,615]" pageId="108" pageNumber="109">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[422,975,589,615]" pageId="108" pageNumber="109">C097B4BE-0F42-4C7D-B087-4C7B4396C826</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="materials_examined">
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<paragraph blockId="108.[140,1108,659,757]" lastBlockId="109.[140,1108,166,757]" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
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<emphasis bold="true" box="[140,371,659,685]" pageId="108" pageNumber="109">
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<emphasis box="[140,241,659,685]" inLineHeading="true" pageId="108" pageNumber="109" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818315" collectingDate="1995-10-19" collectingDateMax="1995-10-28" collectingDateMin="1995-10-19" collectionCode="FMMC" collectorName="S. Goodman" elevation="440" latitude="-24.626667" location="Andohahela" longLatPrecision="125" longitude="46.765" municipality="Province" pageId="108" pageNumber="109" specimenCode="FMMC 7936" specimenCount="3" specimenCount-female="3" stateProvince="Toliara" typeStatus="holotype">
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6 ♂,
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<specimenCount box="[436,480,660,686]" pageId="108" pageNumber="109" type="female">3 ♀</specimenCount>
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<emphasis box="[487,590,659,685]" italics="true" pageId="108" pageNumber="109">
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<typeStatus box="[487,584,659,685]" pageId="108" pageNumber="109">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (hind end missing),
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<specimenCode box="[877,1036,659,686]" collectionCode="FMMC" pageId="108" pageNumber="109">FMMC 7936</specimenCode>
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,
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<collectingMunicipality pageId="108" pageNumber="109">Province</collectingMunicipality>
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<collectingRegion box="[192,274,695,721]" country="Madagascar" name="Toliara" pageId="108" pageNumber="109">Toliara</collectingRegion>
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, RNI
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<location LSID="urn:lsid:plazi:treatment:1772122E786BFFA8FF013D5DACF4E870:9A04F5E3786BFFAAFED43DC3AEB8E880" box="[345,481,695,721]" latitude="-24.626667" longLatPrecision="125" longitude="46.765" municipality="Province" name="Andohahela" pageId="108" pageNumber="109" stateProvince="Toliara">Andohahela</location>
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parcel I,
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<quantity box="[588,665,695,721]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="108" pageNumber="109" unit="m" value="440.0">
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<elevation box="[588,665,695,721]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="108" pageNumber="109" unit="m" value="440.0">440 m</elevation>
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</quantity>
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, rainforest,
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<geoCoordinate box="[799,920,695,721]" degrees="24" direction="south" minutes="37.6" orientation="latitude" pageId="108" pageNumber="109" precision="92" value="-24.626667">24°37.6’ S</geoCoordinate>
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,
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<geoCoordinate box="[932,1056,694,721]" degrees="46" direction="east" minutes="45.9" orientation="longitude" pageId="108" pageNumber="109" precision="92" value="46.765">46°45.9’ E</geoCoordinate>
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, leg.
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<collectorName box="[140,288,730,756]" pageId="108" pageNumber="109">S. Goodman</collectorName>
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,
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<date box="[299,468,730,757]" pageId="108" pageNumber="109" value="1995-10-19" valueMax="1995-10-28" valueMin="1995-10-19">
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<collectingDate box="[299,468,730,757]" pageId="108" pageNumber="109" value="1995-10-19" valueMax="1995-10-28" valueMin="1995-10-19">19–28.X.1995</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2608818247" collectingDate="1995-11-07" collectingDateMax="1995-11-17" collectingDateMin="1995-11-07" collectionCode="FMMC" collectorName="S. Goodman" elevation="1200" lastPageId="109" lastPageNumber="110" latitude="-24.583334" location="Andohahela" longLatPrecision="125" longitude="46.735" municipality="Province" pageId="108" pageNumber="109" specimenCode="FMMC 8190" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
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<emphasis box="[479,586,731,756]" italics="true" pageId="108" pageNumber="109">
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<typeStatus box="[479,581,731,756]" pageId="108" pageNumber="109">Paratypes</typeStatus>
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:
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</emphasis>
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2 ♂,
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<specimenCount box="[651,737,730,756]" pageId="108" pageNumber="109" type="generic">1 imm.</specimenCount>
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,
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<specimenCode box="[748,907,730,756]" collectionCode="FMMC" pageId="108" pageNumber="109">FMMC 8190</specimenCode>
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,
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<collectingMunicipality box="[918,1016,730,756]" pageId="108" pageNumber="109">Province</collectingMunicipality>
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<collectingRegion box="[1022,1104,730,756]" country="Madagascar" name="Toliara" pageId="108" pageNumber="109">Toliara</collectingRegion>
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, RNI
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<location LSID="urn:lsid:plazi:treatment:1772122E786BFFA8FF013D5DACF4E870:9A04F5E3786AFFABFF443FD3AE08EA90" box="[201,337,167,193]" latitude="-24.583334" longLatPrecision="125" longitude="46.735" municipality="Province" name="Andohahela" pageId="109" pageNumber="110" stateProvince="Toliara">Andohahela</location>
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parcel I,
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<quantity box="[448,542,167,193]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="109" pageNumber="110" unit="m" value="1200.0">
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<elevation box="[448,542,167,193]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="109" pageNumber="110" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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, rainforest,
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<geoCoordinate box="[678,801,166,193]" degrees="24" direction="south" minutes="35.0" orientation="latitude" pageId="109" pageNumber="110" precision="92" value="-24.583334">24°35.0’ S</geoCoordinate>
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,
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<geoCoordinate box="[815,940,167,193]" degrees="46" direction="east" minutes="44.1" orientation="longitude" pageId="109" pageNumber="110" precision="92" value="46.735">46°44.1’ E</geoCoordinate>
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, leg.
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<collectorName pageId="109" pageNumber="110">S. Goodman</collectorName>
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,
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<date box="[205,368,202,229]" pageId="109" pageNumber="110" value="1995-11-07" valueMax="1995-11-17" valueMin="1995-11-07">
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<collectingDate box="[205,368,202,229]" pageId="109" pageNumber="110" value="1995-11-07" valueMax="1995-11-17" valueMin="1995-11-07">7–17.XI.1995</collectingDate>
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</date>
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</materialsCitation>
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;
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3767354" ID-Zenodo-Dep="3767354" httpUri="https://zenodo.org/record/3767354/files/figure.png" pageId="108" pageNumber="109" startId="108.[140,212,1526,1548]" targetBox="[140,1108,793,1496]" targetPageId="108">
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<paragraph blockId="108.[140,1109,1526,1676]" pageId="108" pageNumber="109">
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<emphasis bold="true" box="[140,252,1526,1548]" pageId="108" pageNumber="109">Figure 58.</emphasis>
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[259,468,1526,1547]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[259,468,1526,1547]" italics="true" pageId="108" pageNumber="109">Hylekobolus albicollaris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[475,528,1527,1548]" pageId="108" pageNumber="109" rank="species">sp. n.</taxonomicNameLabel>
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, male holotype.
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<emphasis bold="true" box="[685,704,1526,1548]" pageId="108" pageNumber="109">A</emphasis>
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head, lateral view
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<emphasis bold="true" box="[879,896,1526,1548]" pageId="108" pageNumber="109">B</emphasis>
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male legs 1–7, ventral view
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<emphasis bold="true" box="[189,208,1558,1580]" pageId="108" pageNumber="109">C</emphasis>
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telson
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<emphasis bold="true" box="[275,294,1558,1580]" pageId="108" pageNumber="109">D</emphasis>
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anterior gonopod, anterior view
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<emphasis bold="true" box="[605,620,1558,1580]" pageId="108" pageNumber="109">E</emphasis>
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left posterior gonopod, anterior view
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<emphasis bold="true" box="[976,990,1558,1580]" pageId="108" pageNumber="109">F</emphasis>
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anterior gonopod, posterior view
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<emphasis bold="true" box="[351,371,1590,1612]" pageId="108" pageNumber="109">G</emphasis>
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left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
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</paragraph>
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</caption>
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<paragraph blockId="109.[140,1108,166,757]" pageId="109" pageNumber="110">
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<emphasis bold="true" box="[185,484,237,263]" pageId="109" pageNumber="110">
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<emphasis box="[185,257,237,263]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Other</emphasis>
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material examined
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</emphasis>
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:
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<materialsCitation ID-GBIF-Occurrence="2608818275" box="[498,1025,237,264]" collectionCode="FMMC" pageId="109" pageNumber="110" specimenCode="FMMC 8191" specimenCount="1" specimenCount-female="1">
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2 ♂,
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<specimenCount box="[555,597,237,264]" pageId="109" pageNumber="110" type="female">1 ♀</specimenCount>
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,
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<specimenCode box="[612,769,237,263]" collectionCode="FMMC" pageId="109" pageNumber="110">FMMC 8191</specimenCode>
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, same data as previous
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818215" collectingDate="1995-12-05" collectionCode="FMMC" collectorName="d' Andohahela & S. Goodman" latitude="-24.561666" longLatPrecision="125" longitude="46.72167" pageId="109" pageNumber="110" specimenCode="FMMC 8202" specimenCount="2" specimenCount-female="2">
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1 ♂,
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<specimenCount pageId="109" pageNumber="110" type="female">2 ♀</specimenCount>
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,
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<specimenCode box="[175,334,272,298]" collectionCode="FMMC" pageId="109" pageNumber="110">FMMC 8202</specimenCode>
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, RNI
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<collectorName box="[404,569,272,298]" pageId="109" pageNumber="110">d’ Andohahela</collectorName>
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parcel 1, 1875 m, montane rainforest,
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<geoCoordinate degrees="24" direction="south" minutes="33.7" orientation="latitude" pageId="109" pageNumber="110" precision="92" value="-24.561666">24°33.7’ S</geoCoordinate>
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||
,
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<geoCoordinate box="[169,293,307,334]" degrees="46" direction="east" minutes="43.3" orientation="longitude" pageId="109" pageNumber="110" precision="92" value="46.72167">46°43.3’ E</geoCoordinate>
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,
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||
<collectingMethod box="[304,436,307,334]" pageId="109" pageNumber="110">pitfall traps</collectingMethod>
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, leg.
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||
<collectorName box="[494,642,307,334]" pageId="109" pageNumber="110">S. Goodman</collectorName>
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||
,
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||
<date box="[653,784,307,334]" pageId="109" pageNumber="110" value="1995-12-05">
|
||
<collectingDate box="[653,784,307,334]" pageId="109" pageNumber="110" value="1995-12-05">5.XII.1995</collectingDate>
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||
</date>
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</materialsCitation>
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.
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="109" pageNumber="110" type="diagnosis">
|
||
<paragraph blockId="109.[140,1108,166,757]" pageId="109" pageNumber="110">
|
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<emphasis bold="true" box="[185,452,342,368]" pageId="109" pageNumber="110">
|
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<emphasis box="[185,324,342,368]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Differential</emphasis>
|
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diagnosis:
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</emphasis>
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the gonopods clearly identify this species (
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<figureCitation box="[946,1032,342,369]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Figs 59</figureCitation>
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D–G).
|
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,283,378,404]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species">
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||
<emphasis box="[140,283,378,404]" italics="true" pageId="109" pageNumber="110">H. goodmani</emphasis>
|
||
</taxonomicName>
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||
is quite similar in colour to the sympatric
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[771,914,379,404]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[771,914,379,404]" italics="true" pageId="109" pageNumber="110">H. montanus</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel box="[923,984,378,404]" pageId="109" pageNumber="110" rank="species">sp. n.</taxonomicNameLabel>
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The latter, however, is much larger and has completely different posterior gonopods.
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</paragraph>
|
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</subSubSection>
|
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<subSubSection lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110" type="description">
|
||
<paragraph blockId="109.[140,1108,166,757]" pageId="109" pageNumber="110">
|
||
<emphasis bold="true" box="[185,335,448,474]" pageId="109" pageNumber="110">
|
||
<emphasis box="[185,330,448,474]" inLineHeading="true" pageId="109" pageNumber="110" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[341,500,449,474]" italics="true" pageId="109" pageNumber="110">Measurements</emphasis>
|
||
: male
|
||
<typeStatus box="[578,681,448,474]" pageId="109" pageNumber="110">holotype</typeStatus>
|
||
with 49 rings,
|
||
<quantity box="[858,942,448,475]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="109" pageNumber="110" unit="mm" value="55.0">55 mm</quantity>
|
||
long,
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="109" pageNumber="110" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.7–4.9 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph blockId="109.[140,1108,166,757]" pageId="109" pageNumber="110">
|
||
<emphasis box="[185,308,519,545]" italics="true" pageId="109" pageNumber="110">Coloration:</emphasis>
|
||
head, median part of collum, anterior 3/4 of body rings and anterior half of anal segment dark grey (
|
||
<figureCitation box="[500,605,554,581]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59A</figureCitation>
|
||
). Posterior margin of body rings, telson and margins of collum light brown (
|
||
<figureCitation box="[494,597,589,616]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59C</figureCitation>
|
||
). Legs light brown, antennae darker grey (
|
||
<figureCitation captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59B</figureCitation>
|
||
).
|
||
<emphasis box="[211,254,625,650]" italics="true" pageId="109" pageNumber="110">Eyes</emphasis>
|
||
with 40–44 ocelli (
|
||
<figureCitation box="[476,580,624,651]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="109.[140,1108,166,757]" lastBlockId="110.[140,1108,166,545]" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
|
||
<emphasis box="[185,370,659,686]" italics="true" pageId="109" pageNumber="110">Anterior gonopod</emphasis>
|
||
sternite basally wide, apically elevated into a rectangular lobe with a well-rounded tip (
|
||
<figureCitation box="[371,482,694,721]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59D</figureCitation>
|
||
). Mesal process of coxite slender and protruding (
|
||
<figureCitation captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110">Fig. 59D</figureCitation>
|
||
). Apical process of telopodite large and well-rounded. Retrorse margin slender, widely projecting above telopodite margin (
|
||
<figureCitation box="[652,756,166,193]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59F</figureCitation>
|
||
), basally of apical process not projecting (
|
||
<figureCitation box="[270,370,202,229]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767356" ID-Zenodo-Dep="3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="109" pageNumber="110" startId="109.[140,212,1526,1548]" targetBox="[140,1108,850,1506]" targetPageId="109">
|
||
<paragraph blockId="109.[140,1108,1526,1676]" pageId="109" pageNumber="110">
|
||
<emphasis bold="true" box="[140,251,1526,1548]" pageId="109" pageNumber="110">Figure 59.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[257,457,1526,1547]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[257,457,1526,1547]" italics="true" pageId="109" pageNumber="110">Hylekobolus goodmani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[463,515,1527,1548]" pageId="109" pageNumber="110" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis bold="true" box="[669,688,1526,1548]" pageId="109" pageNumber="110">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[860,877,1526,1548]" pageId="109" pageNumber="110">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[1089,1108,1526,1548]" pageId="109" pageNumber="110">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[203,222,1558,1580]" pageId="109" pageNumber="110">D</emphasis>
|
||
anterior gonopod, anterior view
|
||
<emphasis bold="true" box="[538,553,1558,1580]" pageId="109" pageNumber="110">E</emphasis>
|
||
left posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[916,930,1558,1580]" pageId="109" pageNumber="110">F</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[278,298,1590,1612]" pageId="109" pageNumber="110">G</emphasis>
|
||
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="110.[140,1108,166,545]" pageId="110" pageNumber="111">
|
||
<emphasis box="[185,378,237,263]" italics="true" pageId="110" pageNumber="111">Posterior gonopod</emphasis>
|
||
telopodite bent 90° (
|
||
<figureCitation box="[631,740,237,264]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59G</figureCitation>
|
||
). Projection absent from lateral margin (
|
||
<emphasis box="[240,259,273,298]" italics="true" pageId="110" pageNumber="111">w</emphasis>
|
||
,
|
||
<figureCitation box="[276,380,272,299]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59E</figureCitation>
|
||
). Apical process (
|
||
<emphasis box="[584,596,273,298]" italics="true" pageId="110" pageNumber="111">x</emphasis>
|
||
) basally extraordinary slender, only twice as wide as sperm canal (
|
||
<figureCitation box="[374,475,307,334]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59E</figureCitation>
|
||
). Projection of sperm canal (
|
||
<emphasis box="[792,804,308,333]" italics="true" pageId="110" pageNumber="111">z</emphasis>
|
||
) long and slender, running parallel to apical process (
|
||
<figureCitation box="[429,535,342,369]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59G</figureCitation>
|
||
). Tip sharp-edged. Membranous lobe (
|
||
<emphasis box="[977,988,343,368]" italics="true" pageId="110" pageNumber="111">y</emphasis>
|
||
) large and well-rounded (
|
||
<figureCitation box="[306,408,378,405]" captionStart="Figure 59" captionStartId="109.[140,212,1526,1548]" captionTargetBox="[140,1108,850,1506]" captionTargetId="figure@109.[140,1108,1130,1506]" captionTargetPageId="109" captionText="Figure 59. Hylekobolus goodmani sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767356" httpUri="https://zenodo.org/record/3767356/files/figure.png" pageId="110" pageNumber="111">Fig. 59E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="110" pageNumber="111" type="distribution">
|
||
<paragraph blockId="110.[140,1108,166,545]" pageId="110" pageNumber="111">
|
||
<emphasis bold="true" box="[185,485,413,439]" pageId="110" pageNumber="111">
|
||
<emphasis box="[185,336,413,439]" inLineHeading="true" pageId="110" pageNumber="111" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: this species is only known from the Anosy Mountains, north of the Isaka-Ivondro corridor (
|
||
<figureCitation box="[555,639,448,475]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="110" pageNumber="111">Fig. 54</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="110" pageNumber="111" type="etymology">
|
||
<paragraph blockId="110.[140,1108,166,545]" pageId="110" pageNumber="111">
|
||
<emphasis bold="true" box="[185,323,483,509]" pageId="110" pageNumber="111">
|
||
<emphasis box="[185,318,483,509]" inLineHeading="true" pageId="110" pageNumber="111" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[331,437,483,509]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[331,437,483,509]" italics="true" pageId="110" pageNumber="111">goodmani</emphasis>
|
||
</taxonomicName>
|
||
, adjective, named after Dr. Steven M. Goodman who collected this and numerous other remarkable Malagasy millipede species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |