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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7858FF99FE253B29AEF0EFC0" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Spirobolellidae" docType="treatment" docVersion="4" lastPageNumber="96" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="96" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
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<mods:start>1</mods:start>
<mods:end>128</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791453" ID-GBIF-Taxon="164184216" ID-Zenodo-Dep="3791453" LSID="urn:lsid:zoobank.org:act:F5D60255-4786-4DB3-BA0E-DC3A49AF18EA" httpUri="http://treatment.plazi.org/id/1772122E7858FF99FE253B29AEF0EFC0" lastPageNumber="96" pageId="95" pageNumber="96">
<subSubSection box="[424,695,1116,1143]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph blockId="95.[140,695,1116,1143]" box="[424,695,1116,1143]" pageId="95" pageNumber="96">
<heading box="[424,695,1116,1143]" pageId="95" pageNumber="96">
Family
<taxonomicName authority="Brölemann, 1913" box="[514,695,1116,1142]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="discussion">
<paragraph blockId="95.[140,1108,1187,1425]" pageId="95" pageNumber="96">
Of special potential phylogenetic interest are the small species of the South American genus
<taxonomicName authority="Hoffman, 1979" authorityName="Hoffman" authorityYear="1979" box="[265,619,1223,1249]" class="Diplopoda" family="Pachybolidae" genus="Atlanticobolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirobolida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[265,423,1223,1249]" italics="true" pageId="95" pageNumber="96">Atlanticobolus</emphasis>
<bibRefCitation author="Hoffman" box="[433,619,1223,1249]" firstAuthor="Hoffman" pageId="95" pageNumber="96" pagination="930 - 934" refId="ref57195" refString="Hoffman RL (1979) Studies on spiroboloid millipeds. XII. The status of Spirobolus noronhensis Pocock, 1890, and some related species (Pachybolidae). Proceedings of the Biological Society of Washington 91 (4): 930 - 934." type="journal article" year="1979">Hoffman, 1979</bibRefCitation>
</taxonomicName>
, the West African genus
<taxonomicName authority="Keeton, 1964" authorityName="Keeton" authorityYear="1964" class="Diplopoda" genus="Amblybolus" kingdom="Animalia" order="Spirobolida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis box="[917,1045,1223,1249]" italics="true" pageId="95" pageNumber="96">Amblybolus</emphasis>
<bibRefCitation author="Keeton" firstAuthor="Keeton" pageId="95" pageNumber="96" pagination="5 - 6" refId="ref57322" refString="Keeton WT (1964) Amblybolus genus nov. (Diplopoda: Spirobolida: Trigoniuloidea). Pilot Register of Zoology, card Nos 5 - 6." type="book chapter" year="1964">Keeton, 1964</bibRefCitation>
</taxonomicName>
, as well as the here described Malagasy genus. In these three genera, the posterior gonopod coxites touch one another while a sclerotized sternite is absent. The gonopod pair is nevertheless connected by a membrane. At least the genus
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis italics="true" pageId="95" pageNumber="96">Hylekobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[229,307,1364,1390]" pageId="95" pageNumber="96" rank="genus">gen. n.</taxonomicNameLabel>
with its conspicuously elongated posterior gonopods seems to fit well with the
<taxonomicName box="[244,421,1399,1425]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="95" pageNumber="96" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>