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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E784CFF8AFF013FD2AEDFEE05" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Caprobolus andringitra Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="331040CB-492C-483D-8D12-41B8265EB54D" docUuidSource="ZooBank" docVersion="11" lastPageNumber="77" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="76" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791441" ID-GBIF-Taxon="163747634" ID-Zenodo-Dep="3791441" LSID="urn:lsid:zoobank.org:act:331040CB-492C-483D-8D12-41B8265EB54D" httpUri="http://treatment.plazi.org/id/1772122E784CFF8AFF013FD2AEDFEE05" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
<subSubSection box="[140,608,166,193]" pageId="75" pageNumber="76" type="nomenclature">
<paragraph blockId="75.[140,976,166,229]" box="[140,608,166,193]" pageId="75" pageNumber="76">
<heading box="[140,608,166,193]" fontSize="11" level="2" pageId="75" pageNumber="76" reason="3">
<emphasis bold="true" box="[140,608,166,193]" pageId="75" pageNumber="76">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,531,166,193]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,423,166,193]" italics="true" pageId="75" pageNumber="76">Caprobolus andringitra</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[543,608,167,193]" pageId="75" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,976,202,229]" pageId="75" pageNumber="76" type="description">
<paragraph blockId="75.[140,976,166,229]" box="[140,976,202,229]" pageId="75" pageNumber="76">
<uri box="[140,976,202,229]" pageId="75" pageNumber="76">
urn:lsid:zoobank.org:act:
<uuid box="[422,976,202,229]" pageId="75" pageNumber="76">331040CB-492C-483D-8D12-41B8265EB54D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="75" pageNumber="76" type="materials_examined">
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis bold="true" box="[140,372,272,298]" pageId="75" pageNumber="76">
<emphasis box="[140,241,272,298]" inLineHeading="true" pageId="75" pageNumber="76" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818229" collectingDate="1993-12-05" collectingDateMax="1993-12-06" collectingDateMin="1993-12-05" collectorName="S Ambalavao &amp; S. Goodman" country="Madagascar" county="Province" elevation="1210" latitude="-22.222778" location="Volotsangana River" longLatPrecision="21" longitude="46.97167" municipality="Andringitra reserve" pageId="75" pageNumber="76" specimenCode="FMNH-INS 13601" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
6 ♂,
<specimenCount box="[440,524,272,298]" pageId="75" pageNumber="76" type="generic">1 imm.</specimenCount>
<emphasis box="[532,635,272,298]" italics="true" pageId="75" pageNumber="76">
<typeStatus box="[532,629,272,298]" pageId="75" pageNumber="76">Holotype</typeStatus>
:
</emphasis>
1 ♂ (
<quantity box="[705,789,272,299]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="75" pageNumber="76" unit="mm" value="41.0">41 mm</quantity>
long),
<specimenCode box="[873,1103,272,298]" collectionCode="FMNH-INS" pageId="75" pageNumber="76">FMNH-INS 13601</specimenCode>
,
<collectingCountry box="[140,274,307,333]" name="Madagascar" pageId="75" pageNumber="76">Madagascar</collectingCountry>
,
<collectingCounty box="[285,383,307,333]" pageId="75" pageNumber="76">Province</collectingCounty>
<collectingRegion box="[389,535,307,333]" country="Madagascar" name="Fianarantsoa" pageId="75" pageNumber="76">Fianarantsoa</collectingRegion>
,
<collectingMunicipality box="[546,766,307,334]" pageId="75" pageNumber="76">Andringitra reserve</collectingMunicipality>
, camp 3, 40 km
<collectorName box="[953,1103,307,334]" pageId="75" pageNumber="76">S Ambalavao</collectorName>
, along
<location LSID="urn:lsid:plazi:treatment:1772122E784CFF8AFF013FD2AEDFEE05:9A04F5E3784CFF8DFF423E23AEF3EB20" box="[207,426,343,369]" country="Madagascar" county="Province" latitude="-22.222778" longLatPrecision="21" longitude="46.97167" municipality="Andringitra reserve" name="Volotsangana River" pageId="75" pageNumber="76" stateProvince="Fianarantsoa">Volotsangana River</location>
, rainforest,
<quantity box="[556,647,343,369]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="75" pageNumber="76" unit="m" value="1210.0">
<elevation box="[556,647,343,369]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="75" pageNumber="76" unit="m" value="1210.0">1210 m</elevation>
</quantity>
,
<geoCoordinate box="[657,796,343,369]" degrees="22" direction="south" minutes="13" orientation="latitude" pageId="75" pageNumber="76" precision="15" seconds="22" value="-22.222778">22°1322 S</geoCoordinate>
,
<geoCoordinate box="[807,949,342,369]" degrees="46" direction="east" minutes="58" orientation="longitude" pageId="75" pageNumber="76" precision="15" seconds="18" value="46.97167">46°5818 E</geoCoordinate>
, leg.
<collectorName pageId="75" pageNumber="76">S. Goodman</collectorName>
,
<date box="[206,366,378,405]" pageId="75" pageNumber="76" value="1993-12-05" valueMax="1993-12-06" valueMin="1993-12-05">
<collectingDate box="[206,366,378,405]" pageId="75" pageNumber="76" value="1993-12-05" valueMax="1993-12-06" valueMin="1993-12-05">56.XII.1993</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608818343" collectingDate="1993-12-05" collectingDateMax="1993-12-06" collectingDateMin="1993-12-05" collectorName="S Ambalavao &amp; S. Goodman" country="Madagascar" county="Province" elevation="1210" latitude="-22.222778" location="Volotsangana River" longLatPrecision="21" longitude="46.97167" municipality="Andringitra reserve" pageId="75" pageNumber="76" specimenCode="FMNH-INS 13600" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis box="[378,485,379,404]" italics="true" pageId="75" pageNumber="76">
<typeStatus box="[378,480,379,404]" pageId="75" pageNumber="76">Paratypes</typeStatus>
:
</emphasis>
3 ♂,
<specimenCount box="[553,641,378,404]" pageId="75" pageNumber="76" type="generic">1 imm.</specimenCount>
,
<specimenCode box="[653,884,378,404]" collectionCode="FMNH-INS" pageId="75" pageNumber="76">FMNH-INS 13600</specimenCode>
, same data as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818271" box="[202,741,413,440]" collectingDate="1993-12-05" collectingDateMax="1993-12-06" collectingDateMin="1993-12-05" collectorName="S Ambalavao &amp; S. Goodman" country="Madagascar" county="Province" elevation="1210" latitude="-22.222778" location="Volotsangana River" longLatPrecision="21" longitude="46.97167" municipality="Andringitra reserve" pageId="75" pageNumber="76" specimenCode="FMNH-INS 13611" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
1 ♂,
<specimenCode box="[257,485,413,439]" collectionCode="FMNH-INS" pageId="75" pageNumber="76">FMNH-INS 13611</specimenCode>
, same data as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818294" collectingDate="1997-10-07" collectingDateMax="1997-10-12" collectingDateMin="1997-10-07" collectionCode="FMMC" collectorName="S. Goodman" country="Madagascar" elevation="900" latitude="-22.416666" location="Ivohibe" longLatPrecision="1260" longitude="46.933334" pageId="75" pageNumber="76" specimenCode="FMMC 8219" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
2 ♂,
<specimenCode box="[808,965,413,439]" collectionCode="FMMC" pageId="75" pageNumber="76">FMMC 8219</specimenCode>
, RS
<location LSID="urn:lsid:plazi:treatment:1772122E784CFF8AFF013FD2AEDFEE05:9A04F5E3784CFF8DFC7A3EE9AB16EBE6" box="[1015,1103,413,439]" country="Madagascar" latitude="-22.416666" longLatPrecision="1260" longitude="46.933334" name="Ivohibe" pageId="75" pageNumber="76" stateProvince="Fianarantsoa">Ivohibe</location>
,
<quantity box="[140,217,448,475]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="75" pageNumber="76" unit="m" value="900.0">
<elevation box="[140,217,448,475]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="75" pageNumber="76" unit="m" value="900.0">900 m</elevation>
</quantity>
, rainforest,
<geoCoordinate box="[350,449,448,475]" degrees="22" direction="south" minutes="25" orientation="latitude" pageId="75" pageNumber="76" precision="925" value="-22.416666">22°25 S</geoCoordinate>
,
<geoCoordinate box="[461,562,448,475]" degrees="46" direction="east" minutes="56" orientation="longitude" pageId="75" pageNumber="76" precision="925" value="46.933334">46°56 E</geoCoordinate>
, leg.
<collectorName box="[621,769,448,474]" pageId="75" pageNumber="76">S. Goodman</collectorName>
,
<date box="[781,935,448,474]" pageId="75" pageNumber="76" value="1997-10-07" valueMax="1997-10-12" valueMin="1997-10-07">
<collectingDate box="[781,935,448,474]" pageId="75" pageNumber="76" value="1997-10-07" valueMax="1997-10-12" valueMin="1997-10-07">712.X.1997</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis bold="true" box="[185,472,483,509]" pageId="75" pageNumber="76">
<emphasis box="[185,254,483,509]" inLineHeading="true" pageId="75" pageNumber="76" reason="1">Other</emphasis>
material examined
</emphasis>
: 1 ♂,
<collectionCode box="[540,650,484,510]" pageId="75" pageNumber="76">CASENT</collectionCode>
9032814, Province
<collectingRegion box="[868,1009,483,509]" country="Madagascar" name="Fianarantsoa" pageId="75" pageNumber="76">Fianarantsoa</collectingRegion>
, Forêt de Vevembe,
<quantity box="[253,347,519,545]" metricMagnitude="4" metricUnit="m" metricValue="6.659999999999999" pageId="75" pageNumber="76" unit="km" value="66.6">66.6 km</quantity>
293°WNW Farafangana, rainforest in transition to montane rainforest,
<quantity box="[182,256,554,580]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="75" pageNumber="76" unit="m" value="600.0">600 m</quantity>
,
<geoCoordinate box="[267,402,554,580]" degrees="22" direction="south" minutes="47" orientation="latitude" pageId="75" pageNumber="76" precision="15" seconds="28" value="-22.791111">22°4728” S</geoCoordinate>
,
<geoCoordinate box="[413,550,554,581]" degrees="47" direction="east" minutes="10" orientation="longitude" pageId="75" pageNumber="76" precision="15" seconds="55" value="47.181946">47°1055” E</geoCoordinate>
, leg. B. L. Fisher et al., yellow pan trap,
<date box="[982,1107,554,580]" pageId="75" pageNumber="76" value="2006-04-23">23.IV.2006</date>
</paragraph>
</subSubSection>
<subSubSection lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="description">
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis bold="true" box="[185,335,589,615]" pageId="75" pageNumber="76">
<emphasis box="[185,330,589,615]" inLineHeading="true" pageId="75" pageNumber="76" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[344,496,590,615]" italics="true" pageId="75" pageNumber="76">Measurements</emphasis>
: male
<typeStatus box="[574,673,589,615]" pageId="75" pageNumber="76">holotype</typeStatus>
with 44 rings, circa
<quantity box="[912,996,589,616]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="75" pageNumber="76" unit="mm" value="41.0">41 mm</quantity>
long,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="75" pageNumber="76" unit="mm" value="3.2">3.2 mm</quantity>
wide. Male
<typeStatus box="[327,433,625,651]" pageId="75" pageNumber="76">paratypes</typeStatus>
with 43 or 44 rings (specimen from Vevembe with 46), up to
<quantity box="[170,253,660,686]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="75" pageNumber="76" unit="mm" value="44.0">44 mm</quantity>
long, up to
<quantity box="[393,483,659,686]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="75" pageNumber="76" unit="mm" value="3.5">3.5 mm</quantity>
wide.
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,300,695,721]" italics="true" pageId="75" pageNumber="76">Coloration</emphasis>
faded in alcohol. Head, legs and antennae orange-red (
<figureCitation box="[949,1035,695,721]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Figs 39</figureCitation>
AD). Anal valves and subanal scale reddish, preanal process greyish (
<figureCitation box="[845,950,730,756]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39C</figureCitation>
). Metazonites red, remaining parts of body rings up to ozopore openings orange-reddish, more dorsally greyish. Specimens dorsally with a slender, orange line.
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,244,835,861]" italics="true" pageId="75" pageNumber="76">Head</emphasis>
: each eye with circa 2428 ocelli arranged in 6 or 7 vertical rows (
<figureCitation box="[987,1095,835,862]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39D</figureCitation>
). Labrum with standard three irregular teeth and a single row of 1012 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennae long, protruding back to ring 6. Relative lengths of antennomeres: 1&lt;&lt;2=3=4=5=6, first antennomere very short (
<figureCitation box="[274,382,976,1002]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40A</figureCitation>
). Terminal antennomere with four large sensory cones located together inside a membranous area (
<figureCitation box="[544,645,1011,1038]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40B</figureCitation>
). Antennomere 5 latero-apically with five rows (
<figureCitation box="[208,312,1047,1073]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40C</figureCitation>
), antennomere 6 with two rows (
<figureCitation box="[687,786,1047,1073]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40B</figureCitation>
) of sensilla basiconica.
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,373,1082,1108]" italics="true" pageId="75" pageNumber="76">Gnathochilarium</emphasis>
lamellae linguales each with two standard setae located behind one another (
<figureCitation box="[291,394,1117,1143]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39E</figureCitation>
). Stipites each with three apical setae. Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spine-like structures (
<figureCitation captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40D</figureCitation>
). Central pads of endochilarium separated into two areas, a group of 68 sensilla located apically (
<figureCitation box="[380,482,1223,1249]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40E</figureCitation>
). Close to endochilarium with a large group of sensilla, located towards hypopharyngeal crest. Mentum basally subdivided by a membranous suture (
<figureCitation box="[225,327,1293,1319]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39E</figureCitation>
). Stipites each with a sclerotized ledge (
<figureCitation box="[771,871,1293,1319]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39E</figureCitation>
)
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,291,1328,1354]" italics="true" pageId="75" pageNumber="76">Mandible</emphasis>
external tooth simple, rounded; mesal tooth with three cusps (
<figureCitation box="[995,1096,1328,1354]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40F</figureCitation>
). Seven to nine dense pectinate lamellae (
<figureCitation box="[580,686,1363,1390]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40G</figureCitation>
). Intermediate area between pectinate lamellae and molar plate almost completely absent. Molar plate short, only slightly longer than pectinate lamellae, with only six or seven transverse furrows (
<figureCitation box="[964,1065,1434,1460]" captionStart="Figure 40" captionStartId="73.[140,212,1526,1548]" captionTargetBox="[154,1094,165,1507]" captionTargetId="figure@73.[154,1095,165,1507]" captionTargetPageId="73" captionText="Figure 40. Caprobolus andringitra sp. n. male paratype, SEM. A antenna, overview B antenna, apical view on disc C antennomere 5 with sensilla basiconica D gnathochilarium, ventral view E endochilarium detail F left mandible, mesal view G mandible, detail of pectinate lamella. 3iT = 3-combed internal tooth; cp = central pads of endochilarium; Endo = endochilarium; eT = external tooth; L = ledge; LT = lateral tooth; Me = mentum; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica; ST = stipites." figureDoi="http://doi.org/10.5281/zenodo.3767328" httpUri="https://zenodo.org/record/3767328/files/figure.png" pageId="75" pageNumber="76">Fig. 40F</figureCitation>
).
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" box="[185,916,1469,1496]" pageId="75" pageNumber="76">
<emphasis box="[185,274,1469,1495]" italics="true" pageId="75" pageNumber="76">Collum:</emphasis>
smooth, not protruding as far as body ring 2 (
<figureCitation box="[800,904,1469,1495]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39A</figureCitation>
).
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,303,1504,1530]" italics="true" pageId="75" pageNumber="76">Body rings:</emphasis>
mostly smooth, with some irregular punctation and some transverse impressions on mesozona. Ozopores starting at ring 6, touching suture between mesozona and metazona. Ozopore on ring 6 located more ventrally than subsequent ozopores.
</paragraph>
<paragraph blockId="75.[140,1108,272,1672]" pageId="75" pageNumber="76">
<emphasis box="[185,251,1610,1636]" italics="true" pageId="75" pageNumber="76">Telson</emphasis>
: anal valves with well-developed lips and micropunctation. Preanal ring and subanal process well-rounded, not extending beyond anal valves (
<figureCitation box="[876,980,1645,1671]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="75" pageNumber="76">Fig. 39C</figureCitation>
).
</paragraph>
<paragraph blockId="76.[140,1108,167,1108]" pageId="76" pageNumber="77">
<emphasis box="[185,232,167,192]" italics="true" pageId="76" pageNumber="77">Legs</emphasis>
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 410 ventral/mesal setae. Length of midbody legs circa 1.0 times body diameter in males (
<figureCitation box="[663,768,237,263]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="76" pageNumber="77">Fig. 39F</figureCitation>
). Each podomere with a pair of setae ventrally. Coxae 3 and beyond of cylindrical shape. Tarsus with a stout dorso-apical seta and three pairs of setae, apical pair stout, others successively smaller (
<figureCitation box="[148,259,343,369]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="76" pageNumber="77">Fig. 39G</figureCitation>
).
</paragraph>
<paragraph blockId="76.[140,1108,167,1108]" pageId="76" pageNumber="77">
<emphasis box="[185,430,378,404]" italics="true" pageId="76" pageNumber="77">Male sexual characters</emphasis>
: tarsal pads absent (
<figureCitation box="[665,773,378,404]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="76" pageNumber="77">Fig. 39G</figureCitation>
). Coxae 3 and 4 each with a short, conical coxal process. Process protruding posteriorly (
<figureCitation box="[816,918,413,439]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., AC male holotype, DG male paratype A head, lateral view B legs 17, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="76" pageNumber="77">Fig. 39B</figureCitation>
).
</paragraph>
<paragraph blockId="76.[140,1108,167,1108]" pageId="76" pageNumber="77">
<emphasis box="[185,375,448,475]" italics="true" pageId="76" pageNumber="77">Anterior gonopod</emphasis>
sternite elongated into a wide, almost rectangular lobe. Mesal process of coxite prominent, longer than telopodite, but shorter than telopodite process (
<figureCitation box="[189,295,519,545]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41A</figureCitation>
). Telopodite on anterior side with a circular opening for the posterior gonopods (
<figureCitation box="[268,374,554,580]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41A</figureCitation>
). Telopodite process very long, slender, slightly curved inwards. Shape resembling a sable (
<figureCitation box="[434,536,589,615]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41B</figureCitation>
), basally projecting to lateral side into a long, slen- der, well-rounded, retrorse process. Retrorse process laterally extending beyond telopodite (
<figureCitation box="[227,330,659,686]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41B</figureCitation>
).
</paragraph>
<paragraph blockId="76.[140,1108,167,1108]" pageId="76" pageNumber="77">
<emphasis box="[185,388,695,721]" italics="true" pageId="76" pageNumber="77">Posterior gonopods</emphasis>
unique (
<figureCitation box="[501,590,695,721]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Figs 41</figureCitation>
CE). Gonopod pair connected via a wellsclerotized sternite. Coxites and telopodites positioned parallel to those of opposite gonopod. Coxite wide, well-sclerotized, with a single groove (
<figureCitation box="[851,958,765,791]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41C</figureCitation>
). Coxite and telopodite separated by a suture. Telopodite massive, longer than and as wide as coxite (
<figureCitation box="[148,252,835,862]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41C</figureCitation>
). Telopodite basally divided into two branches (
<figureCitation box="[785,926,835,862]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Figs 41D, E</figureCitation>
). Both branches of similar width and almost similar length, resembling a U (
<figureCitation box="[811,893,871,897]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Figs 41</figureCitation>
CE). Sperm canal protruding at base of U, running free but somehow attached to lateral main branch (
<figureCitation box="[148,252,941,967]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41E</figureCitation>
), tip prolonged into flagellum (
<figureCitation box="[622,732,941,967]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41D</figureCitation>
). Mesal branch of telopodite on anterior side medially with a finger-shaped process (
<figureCitation box="[725,832,976,1002]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="76" pageNumber="77">Fig. 41D</figureCitation>
).
</paragraph>
<paragraph blockId="76.[140,1108,167,1108]" box="[185,380,1011,1037]" pageId="76" pageNumber="77">
<emphasis box="[185,380,1011,1037]" italics="true" pageId="76" pageNumber="77">Female unknown.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="76" pageNumber="77" type="etymology">
<paragraph blockId="76.[140,1108,167,1108]" pageId="76" pageNumber="77">
<emphasis bold="true" box="[185,323,1047,1073]" pageId="76" pageNumber="77">
<emphasis box="[185,318,1047,1073]" inLineHeading="true" pageId="76" pageNumber="77" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[330,454,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species">
<emphasis box="[330,454,1047,1073]" italics="true" pageId="76" pageNumber="77">andringitra</emphasis>
</taxonomicName>
, noun in apposition, after the
<typeStatus box="[803,851,1047,1073]" pageId="76" pageNumber="77">type</typeStatus>
locality, the Andringitra mountain massive.
</paragraph>
</subSubSection>
</treatment>
</document>