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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7841FF81FF013BF4AC07EF55" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Granitobolus andohahelensis Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="D022AF86-E072-4090-B604-F7D5A3C8F425" docUuidSource="ZooBank" docVersion="12" lastPageNumber="72" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="71" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791429" ID-GBIF-Taxon="163747632" ID-Zenodo-Dep="3791429" LSID="urn:lsid:zoobank.org:act:D022AF86-E072-4090-B604-F7D5A3C8F425" httpUri="http://treatment.plazi.org/id/1772122E7841FF81FF013BF4AC07EF55" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<subSubSection box="[140,671,1152,1179]" pageId="70" pageNumber="71" type="nomenclature">
<paragraph blockId="70.[140,966,1152,1214]" box="[140,671,1152,1179]" pageId="70" pageNumber="71">
<heading box="[140,671,1152,1179]" fontSize="11" level="2" pageId="70" pageNumber="71" reason="3">
<emphasis bold="true" box="[140,671,1152,1179]" pageId="70" pageNumber="71">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,594,1152,1179]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,485,1152,1179]" italics="true" pageId="70" pageNumber="71">Granitobolus andohahelensis</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[606,671,1152,1178]" pageId="70" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,966,1187,1214]" pageId="70" pageNumber="71" type="description">
<paragraph blockId="70.[140,966,1152,1214]" box="[140,966,1187,1214]" pageId="70" pageNumber="71">
<uri box="[140,966,1187,1214]" pageId="70" pageNumber="71">
urn:lsid:zoobank.org:act:
<uuid box="[421,966,1187,1214]" pageId="70" pageNumber="71">D022AF86-E072-4090-B604-F7D5A3C8F425</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="70" pageNumber="71" type="materials_examined">
<paragraph blockId="70.[140,1108,1257,1672]" pageId="70" pageNumber="71">
<emphasis bold="true" box="[140,379,1257,1283]" pageId="70" pageNumber="71">
<emphasis box="[140,243,1257,1283]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818317" collectingDate="1995-02-06" collectionCode="FMMC" collectorName="K. Emberton" country="Madagascar" county="Province" elevation="500" latitude="-24.921944" location="Mt. Vasiha" longLatPrecision="21" longitude="46.745834" municipality="Andohahela" pageId="70" pageNumber="71" specimenCode="FMMC 13603" specimenCount="39" specimenCount-female="7" stateProvince="Toliara" typeStatus="holotype">
4 ♂,
<specimenCount box="[460,510,1258,1284]" pageId="70" pageNumber="71" type="female">7 ♀</specimenCount>
,
<specimenCount box="[529,634,1258,1284]" pageId="70" pageNumber="71" type="generic">32 imm.</specimenCount>
<emphasis box="[646,750,1258,1284]" italics="true" pageId="70" pageNumber="71">
<typeStatus box="[646,744,1258,1284]" pageId="70" pageNumber="71">Holotype</typeStatus>
:
</emphasis>
1 ♂ (
<quantity box="[834,924,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="70" pageNumber="71" unit="mm" value="33.0">33 mm</quantity>
long),
<specimenCode collectionCode="FMMC" pageId="70" pageNumber="71">FMMC 13603</specimenCode>
,
<collectingCountry box="[233,369,1293,1319]" name="Madagascar" pageId="70" pageNumber="71">Madagascar</collectingCountry>
,
<collectingCounty box="[384,483,1293,1319]" pageId="70" pageNumber="71">Province</collectingCounty>
<collectingRegion box="[492,575,1293,1319]" country="Madagascar" name="Toliara" pageId="70" pageNumber="71">Toliara</collectingRegion>
,
<collectingMunicipality box="[590,730,1293,1319]" pageId="70" pageNumber="71">Andohahela</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7841FF81FF013BF4AC07EF55:9A04F5E37841FF80FD673A7AAC34EF76" box="[746,877,1293,1320]" country="Madagascar" county="Province" latitude="-24.921944" longLatPrecision="21" longitude="46.745834" municipality="Andohahela" name="Mt. Vasiha" pageId="70" pageNumber="71" stateProvince="Toliara">Mt. Vasiha</location>
,
<quantity box="[893,975,1293,1320]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">
<elevation box="[893,975,1293,1320]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<geoCoordinate degrees="24" direction="south" minutes="55" orientation="latitude" pageId="70" pageNumber="71" precision="15" seconds="19" value="-24.921944">24°5519” S</geoCoordinate>
,
<geoCoordinate box="[171,317,1328,1355]" degrees="46" direction="east" minutes="44" orientation="longitude" pageId="70" pageNumber="71" precision="15" seconds="45" value="46.745834">46°4445” E</geoCoordinate>
, station 12, on 15° slope, E of summit, no granite outcrops, leg.
<collectorName pageId="70" pageNumber="71">K. Emberton</collectorName>
,
<date box="[274,387,1363,1390]" pageId="70" pageNumber="71" value="1995-02-06">
<collectingDate box="[274,387,1363,1390]" pageId="70" pageNumber="71" value="1995-02-06">6.II.1995</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608818208" collectingDate="2007-05-30" collectionCode="FMMC" collectorName="Wesener" country="Madagascar" latitude="-24.930166" location="Malio" longLatPrecision="1" longitude="46.772385" municipality="Parc National de Andohahela" pageId="70" pageNumber="71" specimenCode="W045" specimenCount="9" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis box="[402,511,1364,1389]" italics="true" pageId="70" pageNumber="71">
<typeStatus box="[402,505,1364,1389]" pageId="70" pageNumber="71">Paratypes</typeStatus>
:
</emphasis>
1 ♂,
<specimenCount box="[586,633,1364,1390]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<specimenCount box="[651,741,1363,1390]" pageId="70" pageNumber="71" type="generic">8 imm.</specimenCount>
,
<collectionCode box="[756,846,1364,1390]" pageId="70" pageNumber="71">FMMC</collectionCode>
<specimenCode box="[857,933,1363,1390]" collectionCode="FMMC" pageId="70" pageNumber="71">W045</specimenCode>
,
<collectingMunicipality pageId="70" pageNumber="71">Parc National de Andohahela</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7841FF81FF013BF4AC07EF55:9A04F5E37841FF80FEC13A03AECAEFC0" box="[332,403,1399,1425]" country="Madagascar" latitude="-24.930166" longLatPrecision="1" longitude="46.772385" municipality="Parc National de Andohahela" name="Malio" pageId="70" pageNumber="71" stateProvince="Toliara">Malio</location>
, rainforest,
<geoCoordinate box="[543,698,1398,1425]" degrees="24" direction="south" minutes="55.810" orientation="latitude" pageId="70" pageNumber="71" precision="1" value="-24.930166">24°55.810 S</geoCoordinate>
,
<geoCoordinate box="[713,870,1399,1425]" degrees="46" direction="east" minutes="46.343" orientation="longitude" pageId="70" pageNumber="71" precision="1" value="46.772385">46°46.343 E</geoCoordinate>
, leg.
<collectorName box="[935,1031,1399,1425]" pageId="70" pageNumber="71">Wesener</collectorName>
et al.,
<date box="[140,263,1434,1460]" pageId="70" pageNumber="71" value="2007-05-30">
<collectingDate box="[140,263,1434,1460]" pageId="70" pageNumber="71" value="2007-05-30">30.V.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818323" box="[278,853,1434,1461]" collectingDate="2007-05-30" collectionCode="ZMUC" collectorName="Wesener" country="Madagascar" latitude="-24.930166" location="Malio" longLatPrecision="1" longitude="46.772385" municipality="Parc National de Andohahela" pageId="70" pageNumber="71" specimenCode="W045" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
1 ♂,
<specimenCount box="[343,390,1434,1460]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<collectionCode box="[407,495,1434,1460]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="70" pageNumber="71">ZMUC</collectionCode>
<specimenCode box="[506,582,1434,1461]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="70" pageNumber="71">W045</specimenCode>
, same data as previous
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818233" collectingDate="2007-05-30" collectionCode="ZMH" collectorName="Wesener" country="Madagascar" isTailEnumeration="true" latitude="-24.930166" location="Malio" longLatPrecision="1" longitude="46.772385" municipality="Parc National de Andohahela" pageId="70" pageNumber="71" specimenCode="W045" specimenCount="3" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
<specimenCount box="[868,916,1434,1460]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<specimenCount box="[933,1023,1434,1460]" pageId="70" pageNumber="71" type="generic">2 imm.</specimenCount>
,
<collectionCode box="[1038,1108,1434,1460]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="70" pageNumber="71" type="Museum">ZMH</collectionCode>
<specimenCode box="[140,216,1469,1496]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="70" pageNumber="71" type="Museum">W045</specimenCode>
, same data as previous
</materialsCitation>
.
</paragraph>
<paragraph blockId="70.[140,1108,1257,1672]" pageId="70" pageNumber="71">
<emphasis bold="true" box="[185,498,1504,1530]" pageId="70" pageNumber="71">
<emphasis box="[185,257,1504,1530]" inLineHeading="true" pageId="70" pageNumber="71" reason="1">Other</emphasis>
material examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818333" location="University" pageId="70" pageNumber="71" specimenCount="13" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount box="[509,555,1504,1531]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<specimenCount box="[573,677,1504,1530]" pageId="70" pageNumber="71" type="generic">12 imm.</specimenCount>
, W045, same data as previous material, but stored as vouchers at
<location LSID="urn:lsid:plazi:treatment:1772122E7841FF81FF013BF4AC07EF55:9A04F5E37841FF80FE613977AD39EC4C" box="[492,608,1539,1565]" name="University" pageId="70" pageNumber="71" stateProvince="Antananarivo">University</location>
<collectingRegion box="[619,775,1539,1565]" country="Madagascar" name="Antananarivo" pageId="70" pageNumber="71">Antananarivo</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818214" collectingDate="2007-06-06" collectionCode="FMMC" collectorName="Wesener &amp; Schutte" latitude="-24.988094" location="Manantantely" longLatPrecision="1" longitude="46.92443" municipality="Province" pageId="70" pageNumber="71" specimenCode="W078" specimenCount="5" specimenCount-female="1" stateProvince="Toliara">
1 ♂,
<specimenCount box="[852,897,1540,1566]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<specimenCount box="[915,1003,1539,1566]" pageId="70" pageNumber="71" type="generic">4 imm.</specimenCount>
,
<collectionCode box="[1018,1108,1540,1566]" pageId="70" pageNumber="71">FMMC</collectionCode>
<specimenCode box="[140,215,1575,1601]" collectionCode="FMMC" pageId="70" pageNumber="71">W078</specimenCode>
,
<collectingMunicipality box="[229,327,1575,1601]" pageId="70" pageNumber="71">Province</collectingMunicipality>
<collectingRegion box="[335,417,1575,1601]" country="Madagascar" name="Toliara" pageId="70" pageNumber="71">Toliara</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7841FF81FF013BF4AC07EF55:9A04F5E37841FF80FE223953AD17EC10" box="[431,590,1575,1601]" latitude="-24.988094" longLatPrecision="1" longitude="46.92443" municipality="Province" name="Manantantely" pageId="70" pageNumber="71" stateProvince="Toliara">Manantantely</location>
,
<geoCoordinate box="[603,782,1574,1601]" degrees="24" direction="south" minutes="59" orientation="latitude" pageId="70" pageNumber="71" precision="1" seconds="17.14" value="-24.988094">24°5917.14” S</geoCoordinate>
,
<geoCoordinate box="[797,979,1574,1601]" degrees="46" direction="east" minutes="55" orientation="longitude" pageId="70" pageNumber="71" precision="1" seconds="27.95" value="46.92443">46°5527.95” E</geoCoordinate>
, rainforest, leg.
<collectorName box="[186,281,1610,1636]" pageId="70" pageNumber="71">Wesener</collectorName>
et
<collectorName box="[315,404,1610,1636]" pageId="70" pageNumber="71">Schütte</collectorName>
,
<date box="[415,550,1610,1636]" pageId="70" pageNumber="71" value="2007-06-06">
<collectingDate box="[415,550,1610,1636]" pageId="70" pageNumber="71" value="2007-06-06">06.VI.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818346" box="[562,1025,1610,1636]" collectingDate="2007-06-06" collectionCode="ZMH" collectorName="Wesener &amp; Schutte" latitude="-24.988094" location="Manantantely" longLatPrecision="1" longitude="46.92443" municipality="Province" pageId="70" pageNumber="71" specimenCode="W078" specimenCount="1" specimenCount-female="1" stateProvince="Toliara">
<specimenCount box="[562,605,1610,1636]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<collectionCode box="[619,688,1610,1636]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="70" pageNumber="71" type="Museum">ZMH</collectionCode>
<specimenCode box="[695,769,1610,1636]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="70" pageNumber="71" type="Museum">W078</specimenCode>
, same data as previous
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818283" collectingDate="2007-06-06" collectionCode="ZMH" collectorName="Wesener &amp; Schutte" latitude="-24.988094" location="University" longLatPrecision="1" longitude="46.92443" municipality="Province" pageId="70" pageNumber="71" specimenCount="7" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount box="[1036,1078,1610,1636]" pageId="70" pageNumber="71" type="female">1 ♀</specimenCount>
,
<specimenCount pageId="70" pageNumber="71" type="generic">6 imm.</specimenCount>
, W078, same data as previous,
<location LSID="urn:lsid:plazi:treatment:1772122E7841FF81FF013BF4AC07EF55:9A04F5E37841FF80FDA23919ADFAECD6" box="[559,675,1645,1671]" name="University" pageId="70" pageNumber="71" stateProvince="Antananarivo">University</location>
<collectingRegion box="[682,839,1645,1671]" country="Madagascar" name="Antananarivo" pageId="70" pageNumber="71">Antananarivo</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="diagnosis">
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis bold="true" box="[185,453,166,192]" pageId="71" pageNumber="72">
<emphasis box="[185,324,166,192]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[462,660,167,193]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis box="[462,660,167,193]" italics="true" pageId="71" pageNumber="72">G. andohahelensis</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[814,963,167,193]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis box="[814,963,167,193]" italics="true" pageId="71" pageNumber="72">G. endemicus</emphasis>
</taxonomicName>
in a slightly different colour pattern and differently shaped telopodites of the anterior (
<figureCitation box="[999,1099,202,228]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38F</figureCitation>
) and posterior gonopods (
<figureCitation box="[423,530,237,263]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="description">
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis bold="true" box="[185,335,272,298]" pageId="71" pageNumber="72">
<emphasis box="[185,330,272,298]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[341,493,273,298]" italics="true" pageId="71" pageNumber="72">Measurements</emphasis>
: males with 37 or 38 rings, circa
<quantity box="[862,988,272,299]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="71" pageNumber="72" unit="mm" value="33.0" valueMax="35.0" valueMin="31.0">3135 mm</quantity>
long (broken),
<quantity box="[205,346,307,334]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="71" pageNumber="72" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.33.5 mm</quantity>
wide. Females with 38 rings,
<quantity box="[684,811,308,334]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="71" pageNumber="72" unit="mm" value="40.0" valueMax="43.0" valueMin="37.0">3743 mm</quantity>
long,
<quantity box="[883,1024,308,334]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="71" pageNumber="72" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.74.1 mm</quantity>
wide.
</paragraph>
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis box="[185,300,343,369]" italics="true" pageId="71" pageNumber="72">Coloration</emphasis>
affected by alcohol. Anterior part of head and appendages white to reddish (
<figureCitation box="[204,309,378,404]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38A</figureCitation>
). Mesozona and anterior part of metazona dark grey, posterior part of metazona dark orange (
<figureCitation box="[401,484,413,439]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Figs 38</figureCitation>
AC).
<emphasis box="[561,664,413,439]" italics="true" pageId="71" pageNumber="72">Antennae</emphasis>
very short, in males protruding back to ring 4 (
<figureCitation box="[222,324,448,474]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38A</figureCitation>
), in females to ring 2.
<emphasis box="[575,734,448,474]" italics="true" pageId="71" pageNumber="72">Preanal process</emphasis>
well-rounded, extending beyond anal valves (
<figureCitation box="[274,378,483,510]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38C</figureCitation>
).
</paragraph>
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis box="[185,375,519,545]" italics="true" pageId="71" pageNumber="72">Anterior gonopod</emphasis>
sternite elongated into a short, well-rounded lobe (
<figureCitation box="[984,1095,519,545]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38D</figureCitation>
). Coxite large, inconspicuous, with neither processes, nor projections (
<figureCitation box="[922,1030,554,580]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38D</figureCitation>
). Telopodite visible on anterior side, on posterior side apically with a rounded process (
<figureCitation captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38F</figureCitation>
). Telopodite process starting at mesal margin, laterally curved, slightly extending beyond coxite (
<figureCitation box="[313,413,659,686]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38F</figureCitation>
).
</paragraph>
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis box="[185,383,695,721]" italics="true" pageId="71" pageNumber="72">Posterior gonopods</emphasis>
: telopodite apex not erect, covered by a membranous area (
<figureCitation captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38G</figureCitation>
). A membranous area present between apex and beak, with a short process projecting above margin (
<figureCitation box="[391,493,765,791]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 17, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="71" pageNumber="72">Fig. 38E</figureCitation>
).
</paragraph>
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis bold="true" box="[185,456,800,826]" pageId="71" pageNumber="72">
<emphasis box="[185,333,800,826]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Intraspecific</emphasis>
variation:
</emphasis>
the specimens from Manantantely display, in comparison with the
<typeStatus box="[239,338,835,861]" pageId="71" pageNumber="72">holotype</typeStatus>
from Vasiha, small differences on the beak of the posterior gonopod. These differences are now regarded as intraspecific variation. No variation at all was observed between the
<typeStatus box="[391,490,906,932]" pageId="71" pageNumber="72">holotype</typeStatus>
collected in Vasiha 1995 and those collected few hundred meters East in Malio 2007 (
<figureCitation box="[524,610,941,967]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="71" pageNumber="72">Fig. 30</figureCitation>
). Like in
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[723,871,941,968]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis box="[723,871,941,968]" italics="true" pageId="71" pageNumber="72">G. endemicus</emphasis>
</taxonomicName>
, females of different size classes are present.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="distribution">
<paragraph blockId="71.[140,1108,166,1284]" pageId="71" pageNumber="72">
<emphasis bold="true" box="[185,494,1011,1037]" pageId="71" pageNumber="72">
<emphasis box="[185,336,1011,1037]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Distribution</emphasis>
and ecology:
</emphasis>
known from the rainforest of Malio, as well as
<quantity box="[1033,1107,1011,1038]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="71" pageNumber="72" unit="km" value="20.0">20 km</quantity>
east from the forest of Manantantely (
<figureCitation box="[574,659,1047,1073]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="71" pageNumber="72">Fig. 30</figureCitation>
). It is currently unclear if similar specimens collected from Ebosika, the Isaka-Ivondro corridor and Ivorona also belong to
<taxonomicName authority=", Single" authorityName="Single" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis italics="true" pageId="71" pageNumber="72">G. andohahelensis</emphasis>
, Single
</taxonomicName>
specimens taken from the littoral forests of Mandena and Sainte Luce show a uniformly black coloration and most likely belong to a yet undescribed species. All specimens of
<taxonomicName box="[427,623,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis box="[427,623,1187,1214]" italics="true" pageId="71" pageNumber="72">G. andohahelensis</emphasis>
</taxonomicName>
were collected in thin root layers covering large granitic outcrops.
</paragraph>
</subSubSection>
<subSubSection box="[185,862,1258,1284]" pageId="71" pageNumber="72" type="etymology">
<paragraph blockId="71.[140,1108,166,1284]" box="[185,862,1258,1284]" pageId="71" pageNumber="72">
<emphasis bold="true" box="[185,323,1258,1284]" pageId="71" pageNumber="72">
<emphasis box="[185,318,1258,1284]" inLineHeading="true" pageId="71" pageNumber="72" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName box="[330,490,1258,1284]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species">
<emphasis box="[330,490,1258,1284]" italics="true" pageId="71" pageNumber="72">andohahelensis</emphasis>
</taxonomicName>
, adjective, after the
<typeStatus box="[719,767,1258,1284]" pageId="71" pageNumber="72">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>