555 lines
77 KiB
XML
555 lines
77 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7803FFC0FF013E47AB0AEEEF" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Spirobolida Cook 1895" docType="key" docVersion="7" lastPageNumber="7" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="5" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<subSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="key">
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<treatment ID-GBIF-Taxon="164088169" LSID="urn:lsid:plazi:treatment:1772122E7803FFC0FF013E47AB0AEEEF" httpUri="http://treatment.plazi.org/id/1772122E7803FFC0FF013E47AB0AEEEF" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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<subSubSection box="[140,980,307,333]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph blockId="4.[140,980,307,333]" box="[140,980,307,333]" pageId="4" pageNumber="5">
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<heading bold="true" box="[140,980,307,333]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis bold="true" box="[140,980,307,333]" pageId="4" pageNumber="5">
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Key to the Malagasy genera of
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[506,646,307,333]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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of the family
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<taxonomicName baseAuthorityName="Hoffman" baseAuthorityYear="1980" box="[813,975,307,333]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Pachybolidae</taxonomicName>
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:
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="key">
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<keyStep pageId="4" pageNumber="5">
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<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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<keyLead pageId="4" pageNumber="5">
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<specimenCount box="[140,303,378,404]" pageId="4" pageNumber="5" type="male">1 Male</specimenCount>
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coxa 5 with a process curved anteriorly and covering coxa 4................ .....................................................................
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<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[762,1107,412,439]" class="Diplopoda" family="Trigoniulidae" genus="Leptogoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[762,1107,412,439]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[762,929,412,439]" italics="true" pageId="4" pageNumber="5">Leptogoniulus</emphasis>
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<bibRefCitation author="Silvestri" box="[937,1107,412,439]" firstAuthor="Silvestri" pageId="4" pageNumber="5" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
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</emphasis>
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</taxonomicName>
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only
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<taxonomicName authority="(Butler, 1876)" baseAuthorityName="Butler" baseAuthorityYear="1876" box="[651,941,448,475]" class="Diplopoda" family="Trigoniulidae" genus="Leptogoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sorornus">
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<emphasis box="[651,771,449,474]" italics="true" pageId="4" pageNumber="5">L. sorornus</emphasis>
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(Butler, 1876)
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</taxonomicName>
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, introduced to
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<collectingCountry box="[637,770,483,509]" name="Madagascar" pageId="4" pageNumber="5">Madagascar</collectingCountry>
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(
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<bibRefCitation author="Shelley and Lehtinen" box="[784,1095,483,510]" firstAuthor="Shelley" pageId="4" pageNumber="5" pagination="1379 - 1401" refId="ref57656" refString="Shelley RM, Lehtinen PT (1999) Diagnoses, synonymies and occurrences of the pantropical millipeds, Leptogoniulus sorornus (Butler) and Trigoniulus corallinus (Gervais) (Spirobolida: Pachybolidae: Trigoniulinae). Journal of Natural History 33 (9): 1379 - 1401." type="journal article" year="1999">Shelley and Lehtinen 1999</bibRefCitation>
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).
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</keyLead>
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</paragraph>
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<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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<keyLead pageId="4" pageNumber="5">
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– Male coxa 5 lacking processes, or if processes present, then never curved anteriorly. ......................................................................................................
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<emphasis bold="true" box="[1092,1107,553,579]" pageId="4" pageNumber="5">2</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="4" pageNumber="5">
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<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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<keyLead pageId="4" pageNumber="5">
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2 Anterior gonopod box-like, coxite and telopodite reaching similar height, lacking any processes. Collum unmodified. Preanal process absent............... ...........................................................................
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<taxonomicName authority="Pocock, 1894 Only" authorityName="Pocock" authorityYear="1894" class="Diplopoda" family="Trigoniulidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[801,1107,659,686]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[801,938,659,686]" italics="true" pageId="4" pageNumber="5">Trigoniulus</emphasis>
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Pocock, 1894
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</emphasis>
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Only
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</taxonomicName>
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<taxonomicName authority="(Gervais, 1847)" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[621,941,695,721]" class="Diplopoda" family="Trigoniulidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="corallinus">
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<emphasis box="[621,753,695,721]" italics="true" pageId="4" pageNumber="5">T. corallinus</emphasis>
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(Gervais, 1847)
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</taxonomicName>
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, introduced to
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<collectingCountry box="[637,770,730,756]" name="Madagascar" pageId="4" pageNumber="5">Madagascar</collectingCountry>
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(
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<bibRefCitation author="Shelley and Lehtinen" box="[784,1095,730,756]" firstAuthor="Shelley" pageId="4" pageNumber="5" pagination="1379 - 1401" refId="ref57656" refString="Shelley RM, Lehtinen PT (1999) Diagnoses, synonymies and occurrences of the pantropical millipeds, Leptogoniulus sorornus (Butler) and Trigoniulus corallinus (Gervais) (Spirobolida: Pachybolidae: Trigoniulinae). Journal of Natural History 33 (9): 1379 - 1401." type="journal article" year="1999">Shelley and Lehtinen 1999</bibRefCitation>
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).
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</keyLead>
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</paragraph>
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<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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<keyLead pageId="4" pageNumber="5">
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– Anterior gonopods not box-like, or if box-shaped, then at least telopodite with a process, a well-developed preanal process and an enlarged collum (
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<figureCitation captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="4" pageNumber="5">Fig. 31D</figureCitation>
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)..........................................................................................................
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<emphasis bold="true" box="[1092,1107,835,861]" pageId="4" pageNumber="5">3</emphasis>
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep pageId="4" pageNumber="5">
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||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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||
<keyLead pageId="4" pageNumber="5">
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||
3
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<emphasis box="[246,424,871,897]" italics="true" pageId="4" pageNumber="5">Incisura lateralis</emphasis>
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closed. Each
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<emphasis box="[580,760,871,897]" italics="true" pageId="4" pageNumber="5">lamella lingualis</emphasis>
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on gnathochilarium with four or more setae, all located at apical margin. Vulva kidney shaped, both valves meet in sinuous fissure ....................
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<taxonomicName authority="Wesener & Enghoff, 2008 See Wesener et al. 2008" class="Diplopoda" family="Pachybolidae" genus="Madabolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[658,1107,940,967]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[658,791,940,967]" italics="true" pageId="4" pageNumber="5">Madabolus</emphasis>
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<bibRefCitation author="Wesener & Enghoff" box="[798,1107,941,967]" firstAuthor="Wesener" pageId="4" pageNumber="5" pagination="37 - 53" refId="ref58000" refString="Wesener T, Enghoff H, Wagele J-W (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics 22: 37 - 53. [http: // www. publish. csiro. au /? act = view _ file & file _ id = IS 07008. pdf]" type="journal article" year="2008">Wesener & Enghoff, 2008</bibRefCitation>
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</emphasis>
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||
See
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||
<bibRefCitation author="Wesener" box="[884,1108,976,1003]" etAl="et al." firstAuthor="Wesener" pageId="4" pageNumber="5" pagination="37 - 53" refId="ref58000" refString="Wesener T, Enghoff H, Wagele J-W (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics 22: 37 - 53. [http: // www. publish. csiro. au /? act = view _ file & file _ id = IS 07008. pdf]" type="journal article" year="2008">Wesener et al. 2008</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
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<keyLead pageId="4" pageNumber="5">
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–
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<emphasis box="[246,425,1011,1037]" italics="true" pageId="4" pageNumber="5">Incisura lateralis</emphasis>
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open (
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||
<figureCitation box="[507,598,1011,1038]" captionStart="Figure 1" captionStartId="8.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1469]" captionTargetId="figure@8.[140,1108,165,1472]" captionTargetPageId="8" captionText="Figure 1. Corallobolus cruentus sp. n., A–C, E, F, male holotype, D female paratype. A head B telson C legs 1–7, ventral view D second leg pair with vulva, posterior view E anterior gonopods, posterior view F posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = basal joints of mandible; Pre = preanal ring; s = suture on clypeus; sub = subanal scale; St = sternite; T = telopodite; Tp = telopodite process; Vu = vulva. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820785" httpUri="https://zenodo.org/record/3820785/files/figure.png" pageId="4" pageNumber="5">Figs 1A</figureCitation>
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,
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<figureCitation box="[612,664,1012,1038]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="4" pageNumber="5">13A</figureCitation>
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).
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<emphasis box="[685,874,1011,1037]" italics="true" pageId="4" pageNumber="5">Lamella lingualis</emphasis>
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each with two setae located behind one another. Vulva simple, bivalve-like (
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<figureCitation box="[871,936,1047,1073]" captionStart="Figure 4" captionStartId="11.[140,212,615,637]" captionTargetBox="[140,1108,165,595]" captionTargetId="figure@11.[140,1108,165,595]" captionTargetPageId="11" captionText="Figure 4. Corallobolus cruentus sp. n., female paratype, SEM, right vulva. Abbreviations: av = anterior valve; pv = posterior valve." figureDoi="http://doi.org/10.5281/zenodo.3767288" httpUri="https://zenodo.org/record/3767288/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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||
)...................
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||
<emphasis bold="true" box="[1092,1107,1046,1072]" pageId="4" pageNumber="5">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
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||
<keyStep pageId="4" pageNumber="5">
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||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
<specimenCount box="[140,310,1082,1108]" pageId="4" pageNumber="5" type="adult">4 Adult</specimenCount>
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||
males>
|
||
<quantity box="[411,494,1082,1109]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="5" unit="mm" value="95.0">95 mm</quantity>
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||
long and>
|
||
<quantity box="[632,700,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="mm" value="7.0">7 mm</quantity>
|
||
wide. Usually with blood-red/black colour pattern (
|
||
<figureCitation box="[422,507,1117,1143]" captionStart="Figure 1" captionStartId="8.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1469]" captionTargetId="figure@8.[140,1108,165,1472]" captionTargetPageId="8" captionText="Figure 1. Corallobolus cruentus sp. n., A–C, E, F, male holotype, D female paratype. A head B telson C legs 1–7, ventral view D second leg pair with vulva, posterior view E anterior gonopods, posterior view F posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = basal joints of mandible; Pre = preanal ring; s = suture on clypeus; sub = subanal scale; St = sternite; T = telopodite; Tp = telopodite process; Vu = vulva. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820785" httpUri="https://zenodo.org/record/3820785/files/figure.png" pageId="4" pageNumber="5">Fig. 1A</figureCitation>
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||
) .............................................................................
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||
<emphasis bold="true" box="[1092,1107,1117,1143]" pageId="4" pageNumber="5">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
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||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Adult males <
|
||
<quantity box="[420,506,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="mm" value="80.0">80 mm</quantity>
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||
long and <
|
||
<quantity box="[656,749,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="5" unit="mm" value="6.5">6.5 mm</quantity>
|
||
wide, colour never blood red/ pitchblack..................................................................................................
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||
<emphasis bold="true" box="[1092,1107,1187,1213]" pageId="4" pageNumber="5">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
5 Coxite of anterior gonopod with a slender, elongated process, longer than telopodite. Telopodite of posterior gonopod basally diverging into two slender branches, i.e. mesal main branch and lateral basal branch. Both branches curved towards one another, forming an ‘O’ or almost closed ‘C’. Sperm canal always on mesal main branch ................
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||
<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[610,967,1363,1390]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[610,967,1363,1390]" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[610,794,1363,1390]" italics="true" pageId="4" pageNumber="5">Aphistogoniulus</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[801,967,1363,1389]" firstAuthor="Silvestri" pageId="4" pageNumber="5" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(ten species) See
|
||
<bibRefCitation author="Wesener" box="[884,1108,1399,1425]" etAl="et al." firstAuthor="Wesener" pageId="4" pageNumber="5" pagination="15 - 52" refId="ref58142" refString="Wesener T, Enghoff H, Hoffman RL, Sierwald P, Wagele J-W (2009) Revision of the endemic giant fire millipedes of Madagascar, genus Aphistogoniulus (Diplopoda, Spirobolida, Pachybolidae). International Journal of Myriapodology 2 (1): 15 - 52." type="journal article" year="2009">Wesener et al. 2009</bibRefCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
– Process of coxite of anterior gonopod short and wide, or, if slender, then never as long as telopodite. Telopodite of posterior gonopod not like above ..........
|
||
<emphasis bold="true" box="[1092,1107,1469,1495]" pageId="4" pageNumber="5">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<paragraph blockId="4.[140,1108,378,1672]" pageId="4" pageNumber="5">
|
||
<keyLead pageId="4" pageNumber="5">
|
||
6 Telopodite of posterior gonopod large, with a blunt tip; branches or processes absent (
|
||
<figureCitation box="[331,419,1539,1566]" captionStart="Figure 2" captionStartId="9.[140,212,713,735]" captionTargetBox="[140,1108,168,694]" captionTargetId="figure@9.[140,1108,165,694]" captionTargetPageId="9" captionText="Figure 2. Corallobolus cruentus sp. n., male holotype. A anterior gonopods, anterior view B left posterior gonopod, posterior view C left posterior gonopod, anterior view. Cx coxite cp coxite process rp retrorse process St sternite T telopodite Tp telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767284" httpUri="https://zenodo.org/record/3767284/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
|
||
). Telopodite and coxite of equal length, fused, only suture visible (
|
||
<figureCitation box="[301,384,1575,1601]" captionStart="Figure 1" captionStartId="8.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1469]" captionTargetId="figure@8.[140,1108,165,1472]" captionTargetPageId="8" captionText="Figure 1. Corallobolus cruentus sp. n., A–C, E, F, male holotype, D female paratype. A head B telson C legs 1–7, ventral view D second leg pair with vulva, posterior view E anterior gonopods, posterior view F posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = basal joints of mandible; Pre = preanal ring; s = suture on clypeus; sub = subanal scale; St = sternite; T = telopodite; Tp = telopodite process; Vu = vulva. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820785" httpUri="https://zenodo.org/record/3820785/files/figure.png" pageId="4" pageNumber="5">Fig. 1F</figureCitation>
|
||
). Telopodite of anterior gonopod with a large, triangular, retrorse process (
|
||
<figureCitation box="[338,420,1610,1636]" captionStart="Figure 1" captionStartId="8.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1469]" captionTargetId="figure@8.[140,1108,165,1472]" captionTargetPageId="8" captionText="Figure 1. Corallobolus cruentus sp. n., A–C, E, F, male holotype, D female paratype. A head B telson C legs 1–7, ventral view D second leg pair with vulva, posterior view E anterior gonopods, posterior view F posterior gonopods, anterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = basal joints of mandible; Pre = preanal ring; s = suture on clypeus; sub = subanal scale; St = sternite; T = telopodite; Tp = telopodite process; Vu = vulva. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820785" httpUri="https://zenodo.org/record/3820785/files/figure.png" pageId="4" pageNumber="5">Fig. 1E</figureCitation>
|
||
). Coxite with a slender, elongated process (
|
||
<figureCitation box="[885,970,1610,1636]" captionStart="Figure 2" captionStartId="9.[140,212,713,735]" captionTargetBox="[140,1108,168,694]" captionTargetId="figure@9.[140,1108,165,694]" captionTargetPageId="9" captionText="Figure 2. Corallobolus cruentus sp. n., male holotype. A anterior gonopods, anterior view B left posterior gonopod, posterior view C left posterior gonopod, anterior view. Cx coxite cp coxite process rp retrorse process St sternite T telopodite Tp telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767284" httpUri="https://zenodo.org/record/3767284/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
|
||
). Male tarsal pads present only to midbody legs ..............
|
||
<emphasis bold="true" box="[728,962,1644,1671]" pageId="4" pageNumber="5">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[728,876,1644,1671]" class="Diplopoda" family="Pachybolidae" genus="Corallobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[728,876,1644,1671]" italics="true" pageId="4" pageNumber="5">Corallobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[883,962,1645,1671]" pageId="4" pageNumber="5" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(monotypic)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Posterior gonopod always divided into at least two branches. Telopodite and coxite clearly separated by a suture and a membranous area. Often with a stout, finger-shaped mesal coxite branch at suture between coxite and telopodite (
|
||
<figureCitation box="[358,450,272,299]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6C</figureCitation>
|
||
). Coxite far longer than telopodite (
|
||
<figureCitation box="[876,969,272,299]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6C</figureCitation>
|
||
). Process of anterior gonopod telopodite never retrorse (
|
||
<figureCitation box="[745,832,307,334]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6B</figureCitation>
|
||
), process of coxite short and wide (
|
||
<figureCitation box="[364,453,343,370]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6A</figureCitation>
|
||
). Male tarsal pads present on legs 3 and beyond ............
|
||
<emphasis bold="true" box="[1092,1107,343,369]" pageId="5" pageNumber="6">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
7 Telopodite process of anterior gonopod slender basally, proximally widely swollen (
|
||
<figureCitation box="[345,431,413,440]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6B</figureCitation>
|
||
). Telopodite of posterior gonopod is not divided into a lateral main branch and a mesal brance; no swollen area present (
|
||
<figureCitation box="[925,1014,448,475]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6C</figureCitation>
|
||
). Sperm canal running through lateral branch (
|
||
<figureCitation box="[679,768,483,510]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="5" pageNumber="6">Fig. 6D</figureCitation>
|
||
)............................................ ............................................................
|
||
<emphasis bold="true" box="[695,958,518,545]" pageId="5" pageNumber="6">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[695,870,518,545]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[695,870,518,545]" italics="true" pageId="5" pageNumber="6">Sanguinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[877,958,519,545]" pageId="5" pageNumber="6" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(monotypic)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Telopodite process of anterior gonopod slender (
|
||
<figureCitation box="[791,883,554,580]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="5" pageNumber="6">Fig. 9D</figureCitation>
|
||
). Telopodite of posterior gonopod divided into a mesal main branch and a lateral branch (
|
||
<figureCitation captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="5" pageNumber="6">Fig. 9H</figureCitation>
|
||
). Membranous swollen area located between both branches (
|
||
<figureCitation box="[1010,1093,624,650]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="5" pageNumber="6">Fig. 9I</figureCitation>
|
||
). Sperm canal running through mesal branch (
|
||
<figureCitation box="[754,836,659,686]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="5" pageNumber="6">Fig. 9F</figureCitation>
|
||
) .................................. .............................................................
|
||
<emphasis bold="true" box="[699,937,694,721]" pageId="5" pageNumber="6">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[699,849,694,721]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[699,849,694,721]" italics="true" pageId="5" pageNumber="6">Colossobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[856,937,695,721]" pageId="5" pageNumber="6" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(seven species)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
8 Head with an antennal groove. Antenna protruding back to only body ring 2. Head sclerite overlapping part of basal joints of mandible (
|
||
<figureCitation box="[916,1024,765,792]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="5" pageNumber="6">Fig. 25D</figureCitation>
|
||
). Mentum of gnathochilarium not subdivided by a suture (
|
||
<figureCitation box="[841,940,800,827]" captionStart="Figure 25" captionStartId="45.[140,212,1654,1676]" captionTargetBox="[162,1133,1244,1638]" captionTargetId="figure@45.[162,1130,1244,1634]" captionTargetPageId="45" captionText="Figure 25. Distribution of the genus Riotintobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820816" httpUri="https://zenodo.org/record/3820816/files/figure.png" pageId="5" pageNumber="6">Fig. 25E</figureCitation>
|
||
) .................... ...................................................
|
||
<taxonomicName authority="Golovatch & Korsos, 1992 On" authorityName="Golovatch & Korsos" authorityYear="1992" class="Diplopoda" family="Pachybolidae" genus="Dactylobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[626,1107,834,862]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[626,780,835,862]" italics="true" pageId="5" pageNumber="6">Dactylobolus</emphasis>
|
||
Golovatch & Korsós, 1992
|
||
</emphasis>
|
||
On
|
||
</taxonomicName>
|
||
<collectingCountry box="[365,498,871,897]" name="Madagascar" pageId="5" pageNumber="6">Madagascar</collectingCountry>
|
||
only
|
||
<taxonomicName authority="(Karsch, 1881)" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[562,886,871,897]" class="Diplopoda" family="Pachybolidae" genus="Dactylobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bivirgatus">
|
||
<emphasis box="[562,705,871,897]" italics="true" pageId="5" pageNumber="6">D. bivirgatus</emphasis>
|
||
(
|
||
<bibRefCitation author="Karsch" box="[721,878,871,897]" firstAuthor="Karsch" pageId="5" pageNumber="6" pagination="19 - 35" refId="ref57295" refString="Karsch F (1881) Zur Formenlehre der pentazonen Myriopoden. Archiv fur Naturgeschichte 47 (1): 19 - 35, pl. 2." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Species occurs also on the
|
||
<collectingCountry box="[381,489,906,932]" name="Comoros" pageId="5" pageNumber="6">Comoros</collectingCountry>
|
||
,
|
||
<collectingCountry box="[501,613,906,932]" name="Seychelles" pageId="5" pageNumber="6">Seychelles</collectingCountry>
|
||
and close to
|
||
<collectingCountry box="[762,865,906,932]" name="Tanzania" pageId="5" pageNumber="6">Zanzibar</collectingCountry>
|
||
. Probably introduced (
|
||
<bibRefCitation author="VandenSpiegel and Golovatch" box="[689,1099,941,967]" firstAuthor="VandenSpiegel" pageId="5" pageNumber="6" pagination="41 - 58" refId="ref57763" refString="VandenSpiegel D, Golovatch SI (2007) The millipedes of the Comoro Islands (Myriopoda: Diplopoda). Journal of Afrotropical Zoology 3: 41 - 58." type="journal article" year="2007">VandenSpiegel and Golovatch 2007</bibRefCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Antennal groove absent (
|
||
<figureCitation box="[527,632,976,1003]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="5" pageNumber="6">Fig. 26C</figureCitation>
|
||
). Antenna at least protruding back to ring 3, often longer. Especially first basal joint of mandible swollen, projecting towards head, but never above ocelli (
|
||
<figureCitation box="[679,786,1047,1074]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="5" pageNumber="6">Fig. 26C</figureCitation>
|
||
). Mentum often subdivided by a suture (
|
||
<figureCitation box="[385,482,1082,1109]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="5" pageNumber="6">Fig. 26F</figureCitation>
|
||
).................................................................................
|
||
<emphasis bold="true" box="[1092,1107,1081,1107]" pageId="5" pageNumber="6">9</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
9 Sperm canal projecting above lateral margin of telopodite of posterior gonopod (
|
||
<figureCitation box="[305,409,1152,1178]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="5" pageNumber="6">Fig. 22G</figureCitation>
|
||
)....................................
|
||
<emphasis bold="true" box="[684,958,1152,1179]" pageId="5" pageNumber="6">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[684,870,1152,1179]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[684,870,1152,1179]" italics="true" pageId="5" pageNumber="6">Zehntnerobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[877,958,1152,1178]" pageId="5" pageNumber="6" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(monotypic)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Sperm canal never projecting above margins of telopodite of posterior gonopod, or, if so, then never at lateral, but at mesal margin ...........................
|
||
<emphasis bold="true" box="[1077,1108,1222,1248]" pageId="5" pageNumber="6">10</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
10 Telopodite, coxite and sternite of anterior gonopod mesally noticeably elongated into slender processes. Mentum of gnathochilarium not subdivided ... ...........................
|
||
<emphasis bold="true" box="[448,1108,1328,1354]" pageId="5" pageNumber="6">
|
||
<taxonomicName authority="de Saussure & Zehntner, 1901" authorityName="de Saussure & Zehntner" authorityYear="1901" box="[448,961,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[448,592,1328,1354]" italics="true" pageId="5" pageNumber="6">Spiromimus</emphasis>
|
||
<bibRefCitation author="de Saussure & Zehntner" box="[599,961,1328,1354]" firstAuthor="de Saussure" pageId="5" pageNumber="6" pagination="429 - 460" refId="ref56960" refString="de Saussure H, Zehntner L (1901) Myriopoden aus Madagaskar und Zansibar, gesammelt von Dr. A. Voeltzkow. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 26: 429 - 460." type="journal article" year="1901">de Saussure & Zehntner, 1901</bibRefCitation>
|
||
</taxonomicName>
|
||
(13 species)
|
||
</emphasis>
|
||
See
|
||
<bibRefCitation author="Wesener and Enghoff" box="[791,1108,1363,1390]" firstAuthor="Wesener" pageId="5" pageNumber="6" pagination="15 - 52" refId="ref58142" refString="Wesener T, Enghoff H, Hoffman RL, Sierwald P, Wagele J-W (2009) Revision of the endemic giant fire millipedes of Madagascar, genus Aphistogoniulus (Diplopoda, Spirobolida, Pachybolidae). International Journal of Myriapodology 2 (1): 15 - 52." type="journal article" year="2009">Wesener and Enghoff, 2009</bibRefCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
– Telopodite, coxite and sternite of anterior gonopod never altogether elongated into slender processes. Mentum of gnathochilarium basally subdivided (with two exceptions in
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[519,662,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[519,662,1469,1495]" italics="true" pageId="5" pageNumber="6">Riotintobolus</emphasis>
|
||
</taxonomicName>
|
||
) by a suture (
|
||
<figureCitation box="[818,921,1469,1496]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="5" pageNumber="6">Fig. 26G</figureCitation>
|
||
)...................
|
||
<emphasis bold="true" box="[1077,1108,1469,1495]" pageId="5" pageNumber="6">11</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<paragraph blockId="5.[140,1108,167,1602]" pageId="5" pageNumber="6">
|
||
<keyLead pageId="5" pageNumber="6">
|
||
11 Telopodite of posterior gonopod strongly elongated into a simple flagellum (
|
||
<figureCitation box="[254,355,1539,1566]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="5" pageNumber="6">Fig. 23E</figureCitation>
|
||
). Telopodite of anterior gonopod not apically, but at mid-length (!) with a slender, retrorse process (
|
||
<figureCitation box="[605,709,1575,1601]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="5" pageNumber="6">Fig. 23D</figureCitation>
|
||
)
|
||
<emphasis bold="true" box="[718,959,1574,1601]" pageId="5" pageNumber="6">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[718,873,1574,1601]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[718,873,1574,1601]" italics="true" pageId="5" pageNumber="6">Flagellobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[879,959,1575,1601]" pageId="5" pageNumber="6" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(monotypic)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Telopodite of posterior gonopod never shaped like a simple flagellum. Telopodite of anterior gonopods never at mid-length with a slender, retrorse process .....................................................................................................
|
||
<emphasis bold="true" box="[1077,1108,237,263]" pageId="6" pageNumber="7">12</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
12 Telopodite of posterior gonopod apically always with a flag-like membranous extension (
|
||
<figureCitation box="[368,476,307,334]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="6" pageNumber="7">Fig. 26M</figureCitation>
|
||
) ............................
|
||
<emphasis bold="true" box="[701,950,307,334]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[701,862,307,334]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[701,862,307,334]" italics="true" pageId="6" pageNumber="7">Riotintobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[869,950,307,333]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(four species)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" box="[140,1108,342,369]" pageId="6" pageNumber="7">
|
||
<keyLead box="[140,1108,342,369]" pageId="6" pageNumber="7">
|
||
– Posterior gonopod without a flag-like membranous extension..................
|
||
<emphasis bold="true" box="[1077,1108,342,368]" pageId="6" pageNumber="7">13</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
13 Preanal process present, protruding (
|
||
<figureCitation box="[655,757,378,405]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, I–M male holotype, C–H female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="6" pageNumber="7">Fig. 26B</figureCitation>
|
||
). Collum greatly enlarged, ventrally longer than body ring 2 (
|
||
<figureCitation box="[618,724,413,439]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="6" pageNumber="7">Fig. 31A</figureCitation>
|
||
). Posterior margin of body rings with rectangular extensions (
|
||
<figureCitation box="[579,687,448,474]" captionStart="Figure 33" captionStartId="60.[140,212,1558,1580]" captionTargetBox="[170,1078,339,1541]" captionTargetId="figure@60.[170,1079,339,1541]" captionTargetPageId="60" captionText="Figure 33. Pseudocentrobolus aureus sp. n. A–C male, D female paratype, SEM. A left mandible, mesal view B midbody ring, overview C midbody ring, detail of metazona D right vulva, lateral view. 3iT = 3-combed internal tooth; av = anterior valve; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; re = rectangular excavations." figureDoi="http://doi.org/10.5281/zenodo.3767314" httpUri="https://zenodo.org/record/3767314/files/figure.png" pageId="6" pageNumber="7">Fig. 33C</figureCitation>
|
||
). Rings moniliform, caudal margins uplifted (
|
||
<figureCitation box="[350,450,483,510]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="6" pageNumber="7">Fig. 31A</figureCitation>
|
||
) ...................................................................................
|
||
<emphasis bold="true" box="[1077,1108,483,509]" pageId="6" pageNumber="7">14</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Preanal process absent (
|
||
<figureCitation box="[517,623,519,545]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., A–C male holotype, D–G male paratype A head, lateral view B legs 1–7, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="6" pageNumber="7">Fig. 39C</figureCitation>
|
||
). Collum of usual shape (
|
||
<figureCitation box="[921,1027,519,545]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., A–C male holotype, D–G male paratype A head, lateral view B legs 1–7, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="6" pageNumber="7">Fig. 39A</figureCitation>
|
||
). Body rings without extensions, not moniliform (
|
||
<figureCitation box="[728,830,554,580]" captionStart="Figure 39" captionStartId="72.[140,212,1526,1548]" captionTargetBox="[140,1108,927,1507]" captionTargetId="figure@72.[140,1108,927,1506]" captionTargetPageId="72" captionText="Figure 39. Caprobolus andringitra sp. n., A–C male holotype, D–G male paratype A head, lateral view B legs 1–7, ventral view C telson D head, detail of eye and mandible E gnathochilarium, ventral view F right midbody leg, overview G tarsus of midbody leg, SEM. av = anal valves; Co = collum; Gn = gnathochilarium; L = ledge; LL = lamella lingualis; Md = mandible; Me = mentum; Pre = preanal ring; re = rectangular extension; ST = stipites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767326" httpUri="https://zenodo.org/record/3767326/files/figure.png" pageId="6" pageNumber="7">Fig. 39C</figureCitation>
|
||
) ...............................
|
||
<emphasis bold="true" box="[1077,1108,553,579]" pageId="6" pageNumber="7">15</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
14 Telopodite of posterior gonopods at mesal margin with a rounded disc carrying numerous small spines (
|
||
<figureCitation box="[557,662,624,650]" captionStart="Figure 31" captionStartId="57.[140,212,1494,1516]" captionTargetBox="[198,1049,450,1475]" captionTargetId="figure@57.[198,1051,450,1477]" captionTargetPageId="57" captionText="Figure 31. Pseudocentrobolus aureus sp. n., A–D, G, H male paratype; E, F female paratype. A head, lateral view B legs 1–7, ventral view C telson D head, antenna and collum, lateral view E second leg with vulva F female midbody ring with legs G posterior gonopods, anterior view H posterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; mem-p = membranous process; Pre = preanal ring; St = sternite; St-Cx = coxosternite; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820820" httpUri="https://zenodo.org/record/3820820/files/figure.png" pageId="6" pageNumber="7">Figs 31A</figureCitation>
|
||
, 34A, B), and a membranous process at mid-length (
|
||
<figureCitation box="[387,492,659,686]" captionStart="Figure 34" captionStartId="61.[140,212,641,663]" captionTargetBox="[140,1108,165,621]" captionTargetId="figure@61.[140,1108,165,621]" captionTargetPageId="61" captionText="Figure 34. Pseudocentrobolus aureus sp. n., male holotype, anterior gonopod A anterior view B posterior view. Cx = coxite; St = sternite; T = telopodite. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767316" httpUri="https://zenodo.org/record/3767316/files/figure.png" pageId="6" pageNumber="7">Fig. 34D</figureCitation>
|
||
) ...................
|
||
<emphasis bold="true" box="[648,955,659,686]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[648,867,659,686]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[648,867,659,686]" italics="true" pageId="6" pageNumber="7">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[874,955,659,685]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(two species)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Telopodite of posterior gonopods without a disc, apically with a beak-shaped process (
|
||
<figureCitation box="[342,453,730,756]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="6" pageNumber="7">Fig. 37M</figureCitation>
|
||
), at mid-length without a process .................................... ..............................................................
|
||
<emphasis bold="true" box="[709,955,764,791]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[709,867,764,791]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[709,867,764,791]" italics="true" pageId="6" pageNumber="7">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[874,955,765,791]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(two species)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
15 Telopodite of anterior gonopod with a very long, thin, sabre-like process (
|
||
<figureCitation captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="6" pageNumber="7">Fig. 41B</figureCitation>
|
||
). Telopodite of posterior gonopod basally divided into two long, slender branches of similar length and width, resembling a U (
|
||
<figureCitation box="[862,967,871,897]" captionStart="Figure 41" captionStartId="74.[140,212,1526,1548]" captionTargetBox="[232,1016,176,1498]" captionTargetId="figure@74.[223,1024,165,1506]" captionTargetPageId="74" captionText="Figure 41. Caprobolus andringitra sp. n., male holotype. A anterior gonopod, anterior view B anterior gonopod, posterior view C right posterior gonopod, posterior view D right posterior gonopod, telopodite, posterior view E right posterior gonopod, telopodite, anterior view. cp = internal coxite process; Cx = coxite; rp = retrorse process; SC = free sperm canal; St = sternite; T = telopodite; T-ibr = inner branch of telopodite; T-mbr =main branch of telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820823" httpUri="https://zenodo.org/record/3820823/files/figure.png" pageId="6" pageNumber="7">Fig. 41D</figureCitation>
|
||
)................. ..................................................................
|
||
<emphasis bold="true" box="[734,958,905,932]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[734,870,905,932]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[734,870,905,932]" italics="true" pageId="6" pageNumber="7">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[877,958,906,932]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(monotypic)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" box="[140,1108,941,968]" pageId="6" pageNumber="7">
|
||
<keyLead box="[140,1108,941,968]" pageId="6" pageNumber="7">
|
||
– Anterior and posterior gonopods not shaped like above ...........................
|
||
<emphasis bold="true" box="[1077,1108,941,967]" pageId="6" pageNumber="7">16</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="6" pageNumber="7">
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
16 Process of anterior gonopod telopodite basally slender, apically wide and slightly swollen (
|
||
<figureCitation box="[440,545,1011,1038]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="6" pageNumber="7">Fig. 44E</figureCitation>
|
||
). Posterior gonopod without a swollen membranous area. Sperm canal discharging on lateral margin (
|
||
<figureCitation box="[858,963,1047,1073]" captionStart="Figure 44" captionStartId="78.[140,212,695,717]" captionTargetBox="[140,1108,165,676]" captionTargetId="figure@78.[140,1109,165,676]" captionTargetPageId="78" captionText="Figure 44. Alluviobolus laticlavius sp. n. male holotype. A head, lateral view B legs 1–7, ventral view C telson D gnathochilarium, ventral view E anterior gonopod, anterior view F right posterior gonopod, anterior view G anterior gonopod, posterior view H right posterior gonopod, telopodite, posterior view I posterior gonopods, coxite and sternite, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3820827" httpUri="https://zenodo.org/record/3820827/files/figure.png" pageId="6" pageNumber="7">Fig. 44H</figureCitation>
|
||
) ................. ................................................................
|
||
<emphasis bold="true" box="[721,941,1081,1108]" pageId="6" pageNumber="7">
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[721,869,1081,1108]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[721,869,1081,1108]" italics="true" pageId="6" pageNumber="7">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
sp. n.
|
||
</emphasis>
|
||
(three species)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,167,1214]" pageId="6" pageNumber="7">
|
||
<keyLead pageId="6" pageNumber="7">
|
||
– Anterior gonopod apically never with a slender process. Posterior telopodite apically with a swollen membranous area (
|
||
<figureCitation box="[720,814,1152,1178]" captionStart="Figure 48" captionStartId="86.[140,212,1494,1516]" captionTargetBox="[209,1039,164,1474]" captionTargetId="figure@86.[208,1039,164,1087]" captionTargetPageId="86" captionText="Figure 48. Ostinobolus rufus sp. n., A, B, D, G–J male holotype; C, E, F male paratype. A head, lateral view B legs 1–7, ventral view C living specimen D telson E gnathochilarium F midbody ring with leg G anterior gonopod, anterior view H anterior gonopod, posterior view I left posterior gonopod, anterior view Į left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; L = ledge; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767338" httpUri="https://zenodo.org/record/3767338/files/figure.png" pageId="6" pageNumber="7">Fig. 48J</figureCitation>
|
||
). Sperm canal discharging at mesal margin........................................
|
||
<emphasis bold="true" box="[720,950,1187,1214]" pageId="6" pageNumber="7">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[720,862,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis bold="true" box="[720,862,1187,1214]" italics="true" pageId="6" pageNumber="7">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[869,950,1187,1213]" pageId="6" pageNumber="7" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(four species)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |