treatments-xml/data/17/60/61/1760610960073A0E56CF87F6BDDFF822.xml

<document id="6F98270F6C589B5D3A2E8469F6610F3A" ID-CLB-Dataset="46498" ID-DOI="10.11646/zootaxa.3764.1.6" ID-GBIF-Dataset="36fe0b2b-664f-42f2-88ae-e15601dc1650" ID-ISSN="1175-5326" ID-Zenodo-Dep="227486" ID-ZooBank="436036B6-3707-42CB-B056-AC3DE0BEE7E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460380638178" checkinUser="plazi" docAuthor="Chen, Szu-Lung &amp; Yeh, Wen-Chi" docDate="2014" docId="1760610960073A0E56CF87F6BDDFF822" docLanguage="en" docName="zt03764p100.pdf" docOrigin="Zootaxa 3764 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sarasaeschana chiangchinlii Chen &amp; Yeh, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="97" masterDocId="EB59197160063A0B56588608BC69FFD2" masterDocTitle="Description of a new species of the genus Sarasaeschna Karube &amp; Yeh, with a key to the species of Taiwan (Odonata: Anisoptera: Aeshnidae)" masterLastPageNumber="100" masterPageNumber="92" pageNumber="93" updateTime="1698296803953" updateUser="ExternalLinkService">
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<mods:title id="3BD6E9074F1B45142827750AE590CACF">Description of a new species of the genus Sarasaeschna Karube &amp; Yeh, with a key to the species of Taiwan (Odonata: Anisoptera: Aeshnidae)</mods:title>
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<mods:namePart id="AB5651BCAAD455EF115CA7EF082A6C61">Chen, Szu-Lung</mods:namePart>
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<mods:namePart id="4009EBF64362247710DDB4ACDE3F4857">Yeh, Wen-Chi</mods:namePart>
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<treatment id="1760610960073A0E56CF87F6BDDFF822" ID-DOI="http://doi.org/10.5281/zenodo.6128097" ID-GBIF-Taxon="119414128" ID-Zenodo-Dep="6128097" LSID="urn:lsid:plazi:treatment:1760610960073A0E56CF87F6BDDFF822" httpUri="http://treatment.plazi.org/id/1760610960073A0E56CF87F6BDDFF822" lastPageId="5" lastPageNumber="97" pageId="1" pageNumber="93">
<subSubSection id="D7D3839460073A0A56CF87F6BE31FDE9" pageId="1" pageNumber="93" type="nomenclature">
<paragraph id="9F76D01F60073A0A56CF87F6BE2EFDC5" blockId="1.[151,600,510,571]" box="[151,583,510,535]" pageId="1" pageNumber="93">
<heading id="C43E677360073A0A56CF87F6BE2EFDC5" bold="true" box="[151,583,510,535]" fontSize="10" level="2" pageId="1" pageNumber="93" reason="2">
<emphasis id="ADBD0C0D60073A0A56CF87F6BE2EFDC5" bold="true" box="[151,583,510,535]" pageId="1" pageNumber="93">
<taxonomicName id="58C9AB9C60073A0A56CF87F6BD8FFDC5" box="[151,486,510,535]" class="Insecta" family="Aeshnidae" genus="Sarasaeschana" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="93" phylum="Arthropoda" rank="species" species="chiangchinlii" status="sp. nov.">
<emphasis id="ADBD0C0D60073A0A56CF87F6BD8FFDC5" bold="true" box="[151,486,510,535]" italics="true" pageId="1" pageNumber="93">Sarasaeschana chiangchinlii</emphasis>
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<taxonomicNameLabel id="B68EB17660073A0A57B587F6BE2EFDC5" box="[493,583,510,535]" pageId="1" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="9F76D01F60073A0A56CF842ABE31FDE9" blockId="1.[151,600,510,571]" box="[151,600,546,571]" pageId="1" pageNumber="93">Figures 1–7, 15–17, 21, 25, 29, 32 &amp; 33</paragraph>
</subSubSection>
<subSubSection id="D7D3839460073A0A56CF8462B8F6FD75" pageId="1" pageNumber="93" type="etymology">
<paragraph id="9F76D01F60073A0A56CF8462B8F6FD75" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
<emphasis id="ADBD0C0D60073A0A56CF8462BD77FD51" bold="true" box="[151,286,618,643]" pageId="1" pageNumber="93">Etymology.</emphasis>
The new species is dedicated to Mr. Chin-Li Chiang for his continuous effort and hard work in searching for many rare dragonfly species of 
<collectingCountry id="E7DE908F60073A0A54CB8486BE8CFD75" box="[659,741,654,679]" name="Taiwan" pageId="1" pageNumber="93">Taiwan</collectingCountry>
and also discovery of this new species.
</paragraph>
</subSubSection>
<subSubSection id="D7D3839460073A0A569F84BAB972FCC1" pageId="1" pageNumber="93" type="diagnosis">
<paragraph id="9F76D01F60073A0A569F84BAB972FCC1" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
<emphasis id="ADBD0C0D60073A0A569F84BABD28FD19" bold="true" box="[199,321,690,715]" pageId="1" pageNumber="93">Diagnosis.</emphasis>
<taxonomicName id="58C9AB9C60073A0A571384BBBD97FD19" box="[331,510,691,715]" class="Insecta" family="Aeshnidae" genus="Sarasaeschana" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="93" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60073A0A571384BBBD97FD19" box="[331,510,691,715]" italics="true" pageId="1" pageNumber="93">S. chiangchinlii</emphasis>
</taxonomicName>
is a medium and black aeshnid with yellow markings. It differs from the other known 
<taxonomicName id="58C9AB9C60073A0A56B284DEBDEFFD3C" box="[234,390,726,750]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="ADBD0C0D60073A0A56B284DEBDEFFD3C" box="[234,390,726,750]" italics="true" pageId="1" pageNumber="93">Sarasaeschna</emphasis>
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mainly in having the male cercus resemble a sickle. Male has hastate and female has sagittate AD spots on S2. All wings of female colored with pale brown entirely and more intensely at basal half.
</paragraph>
</subSubSection>
<subSubSection id="D7D3839460073A0E569F8516BDDDF8EE" lastPageId="5" lastPageNumber="97" pageId="1" pageNumber="93" type="description">
<paragraph id="9F76D01F60073A0A569F8516B848FBE1" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
<emphasis id="ADBD0C0D60073A0A569F8516BD6DFCE5" bold="true" box="[199,260,798,823]" pageId="1" pageNumber="93">Male</emphasis>
(
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<emphasis id="ADBD0C0D60073A0A574C8516BD12FCE5" bold="true" box="[276,379,798,823]" pageId="1" pageNumber="93">holotype</emphasis>
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, Figs 1, 3). HEAD: labium with medial lobe yellow and lateral lobes dark brown. Face black on lower part, with faint yellow basal spots on mandibles and labrum. Anteclypeus black, postclypeus yellow with lower margin black. Antefrons black and wrinkled, lateral sides and lower 1/5 smooth and yellow, upper margin straight (Fig. 5). Postfrons depressed medially and black, laterally invaded by a pair of transverse yellow spots originating from yellow area of antefrons, and forming a T-mark (
<figureCitation id="07F2CC9A60073A0A55C485A6BF9AFC15" box="[924,1011,942,967]" captionStart="FIGURES 18 – 29" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[248,1345,206,1889]" captionTargetId="figure@4.[236,1350,193,1894]" captionTargetPageId="4" captionText="FIGURES 18 – 29. Different characters of the Sarasaeschna of Taiwan. 18 – 21, male pterothorax, dorsal; 22 – 25, male cerci, dorsal; 26 – 29, male cerci, lateral; 18, 22 &amp; 26, S. pyanan; 19, 23 &amp; 27, S. lieni; 20, 24 &amp; 28, S. tsaopiensis; 21, 25 &amp; 29, S. chiangchinlii." httpUri="https://zenodo.org/record/227488/files/figure.png" pageId="1" pageNumber="93">Fig. 21</figureCitation>
). Vertex, antenna and occiput black; vertex about 1/3 width of frons, tumid, with pair of small faint yellow spots on top. Compound eyes dull green in life, interorbital suture short, about 2/3 width of vertex. Black hairs present on upper half of head, denser and longer on frons, vertex and occiput, and lacking from wrinkled area of antefrons.
</paragraph>
<paragraph id="9F76D01F60073A0A569F8237BE49FAD9" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
PTEROTHORAX: Dorsal stripes cigar 
<emphasis id="ADBD0C0D60073A0A54DB8236BEE5FB85" bold="true" box="[643,652,1086,1111]" pageId="1" pageNumber="93">-</emphasis>
shaped and a little divergent downward from each other, lower ends obtusely attenuate and upper ends connected above with pair of round spots (
<figureCitation id="07F2CC9A60073A0A527B826AB813FBA9" box="[1059,1146,1122,1147]" captionStart="FIGURES 18 – 29" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[248,1345,206,1889]" captionTargetId="figure@4.[236,1350,193,1894]" captionTargetPageId="4" captionText="FIGURES 18 – 29. Different characters of the Sarasaeschna of Taiwan. 18 – 21, male pterothorax, dorsal; 22 – 25, male cerci, dorsal; 26 – 29, male cerci, lateral; 18, 22 &amp; 26, S. pyanan; 19, 23 &amp; 27, S. lieni; 20, 24 &amp; 28, S. tsaopiensis; 21, 25 &amp; 29, S. chiangchinlii." httpUri="https://zenodo.org/record/227488/files/figure.png" pageId="1" pageNumber="93">Fig. 21</figureCitation>
). Antealar sinuses black. Stripes of mesepimeron roundly obtuse at upper ends, broadening and touching humeral suture at lower half. Upper spot of metepisternum small and triangular, posterior 3/4 of metepimeron yellow. Mesinfraepistermun entirely yellow, metinfraepisternum yellow at ventral margin and posterior half. Brown long hairs densely present on front and less so on sides of pterothorax.
</paragraph>
<paragraph id="9F76D01F60073A0A569F831FBFABFAFD" blockId="1.[151,1437,618,1978]" box="[199,962,1302,1327]" pageId="1" pageNumber="93">LEGS: Black, apex of hind femur reaching to anterior margin of S2.</paragraph>
<paragraph id="9F76D01F60073A0A569F8333BFFFF9D5" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
WINGS: hyaline, slightly tinged with pale brown at basal extremity, veins black. Pterostigma dark brown, braced in all wings and underlain by 1.5–2.0 cells. Nodal index 6:16:17: 
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forewings and 7:11:13: 
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hindwings. One cubito-anal crossvein in all wings. Triangle a little longer in forewings than in hindwings and two- or threecelled. Hypertriangles crossed in left wings and uncrossed in right wings, subtriangles uncrossed. Anal loop compact and three- or four-celled, anal triangle three-celled. Tornus roundly angulate; membranule well developed and pale brownish, reaching downward to upper 1/3 of anal triangle.
</paragraph>
<paragraph id="9F76D01F60073A0A569F801BBD33F868" blockId="1.[151,1437,618,1978]" pageId="1" pageNumber="93">
ABDOMEN: S1 and S2 swollen, S3 constricted at anterior 1/3; posterior half of S3–7 widened and weakly spindle-shaped, S8–10 narrow and parallel-sided. Dorsally, S2 with a hastate AD spot and a pair of round and closely adjoining PD spots (
<figureCitation id="07F2CC9A60073A0A578B8052BE4FF9A1" box="[467,550,1626,1651]" captionStart="FIGURES 8 – 17" captionStartId="3.[151,264,1940,1963]" captionTargetBox="[268,1317,200,1887]" captionTargetId="figure@3.[256,1333,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 17. Dorsal maculations of abdominal S 2 in the Sarasaeschna of Taiwan. 8, S. pyanan, male; 9, S. pyanan, female; 10, S. lieni, male; 11, S. lieni, female; 12, S. tsaopiensis, male; 13 &amp; 14, S. tsaopiensis, females, 13, normal form, 14, dark form; 15, S. chiangchinlii, holotype male; 16 &amp; 17, S. chiangchinlii, paratype females." httpUri="https://zenodo.org/record/227487/files/figure.png" pageId="1" pageNumber="93">Fig. 15</figureCitation>
), S3–8 with paired and closely adjoining PD spots, about equal sized on S3–6, smaller on S7 and tiny on S8. Laterally, S1 yellowish, S2 with large and square AL spot, broadly crescent PL spot and small spot on posteroventral corner (Fig. 32), AL spot covering auricle; S3–5 with AL spots, triangular on S3, small and transverse on S4 and S5. Ventrally, bases of S4–6 with pair of faint spots. Auricles semicircular in shape, protruding backward and armed with row of dark denticles at posterior margin. Mid-dorsal carina well defined on S3–7 and basal half of S8. Cercus about 2-1/3 as long as S10, in dorsal view sickle-shaped, with basal 1/3 narrow and apical 2/3 expanded and broadly ridged dorsally, apex strongly incurved and blunt (
<figureCitation id="07F2CC9A60073A0A52DE813DB8B5F89C" box="[1158,1244,1845,1870]" captionStart="FIGURES 18 – 29" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[248,1345,206,1889]" captionTargetId="figure@4.[236,1350,193,1894]" captionTargetPageId="4" captionText="FIGURES 18 – 29. Different characters of the Sarasaeschna of Taiwan. 18 – 21, male pterothorax, dorsal; 22 – 25, male cerci, dorsal; 26 – 29, male cerci, lateral; 18, 22 &amp; 26, S. pyanan; 19, 23 &amp; 27, S. lieni; 20, 24 &amp; 28, S. tsaopiensis; 21, 25 &amp; 29, S. chiangchinlii." httpUri="https://zenodo.org/record/227488/files/figure.png" pageId="1" pageNumber="93">Fig. 25</figureCitation>
); in lateral view, cercus with wide and deep basal notch (
<figureCitation id="07F2CC9A60073A0A54028151BEC6F8A0" box="[602,687,1881,1906]" captionStart="FIGURES 18 – 29" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[248,1345,206,1889]" captionTargetId="figure@4.[236,1350,193,1894]" captionTargetPageId="4" captionText="FIGURES 18 – 29. Different characters of the Sarasaeschna of Taiwan. 18 – 21, male pterothorax, dorsal; 22 – 25, male cerci, dorsal; 26 – 29, male cerci, lateral; 18, 22 &amp; 26, S. pyanan; 19, 23 &amp; 27, S. lieni; 20, 24 &amp; 28, S. tsaopiensis; 21, 25 &amp; 29, S. chiangchinlii." httpUri="https://zenodo.org/record/227488/files/figure.png" pageId="1" pageNumber="93">Fig. 29</figureCitation>
). Epiproct a little shorter than 1/2 of cercus, apex upcurved gently and notched deeply with two forks markedly diverging outward, depth of the apical notch a little longer than half its width (Fig. 6).
</paragraph>
<paragraph id="9F76D01F60043A0956CF816EBED4F849" blockId="2.[151,1436,1894,1947]" pageId="2" pageNumber="94">
<emphasis id="ADBD0C0D60043A0956CF816EBD50F8AE" bold="true" box="[151,313,1894,1917]" pageId="2" pageNumber="94">FIGURES 1–7.</emphasis>
<taxonomicName id="58C9AB9C60043A095767816EBE3EF8AE" box="[319,599,1894,1916]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="94" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60043A095767816EBE3EF8AE" box="[319,599,1894,1916]" italics="true" pageId="2" pageNumber="94">Sarasaeschna chiangchinlii</emphasis>
</taxonomicName>
. 1, 3, 5 &amp; 6, 
<typeStatus id="40726EBD60043A095481816EBF5BF8AE" box="[729,818,1894,1916]" pageId="2" pageNumber="94" type="holotype">holotype</typeStatus>
male; 2, 4 &amp; 7, 
<typeStatus id="40726EBD60043A09558B816FB843F8AE" box="[979,1066,1895,1916]" pageId="2" pageNumber="94" type="paratype">paratype</typeStatus>
female; 1 &amp; 2, dorsal; 3 &amp; 4, lateral; 5, face; 6, male epiproct, ventral; 7, ovipositor, lateral.
</paragraph>
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<paragraph id="9F76D01F60053A0856CF819CBFADF83A" blockId="3.[151,1436,1940,2024]" pageId="3" pageNumber="95">
<emphasis id="ADBD0C0D60053A0856CF819CBD2FF878" bold="true" box="[151,326,1940,1963]" pageId="3" pageNumber="95">FIGURES 8–17.</emphasis>
Dorsal maculations of abdominal S2 in the 
<taxonomicName id="58C9AB9C60053A0854A7819DBFE5F879" box="[767,908,1941,1963]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="ADBD0C0D60053A0854A7819DBFE5F879" box="[767,908,1941,1963]" italics="true" pageId="3" pageNumber="95">Sarasaeschna</emphasis>
</taxonomicName>
of Taiwan. 8, 
<taxonomicName id="58C9AB9C60053A085243819EB814F879" box="[1051,1149,1942,1963]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="pyanan">
<emphasis id="ADBD0C0D60053A085243819EB814F879" box="[1051,1149,1942,1963]" italics="true" pageId="3" pageNumber="95">S. pyanan</emphasis>
</taxonomicName>
, male; 9, 
<taxonomicName id="58C9AB9C60053A0852B9819EB92DF879" box="[1249,1348,1942,1963]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="pyanan">
<emphasis id="ADBD0C0D60053A0852B9819EB92DF879" box="[1249,1348,1942,1963]" italics="true" pageId="3" pageNumber="95">S. pyanan</emphasis>
</taxonomicName>
, female; 10, 
<taxonomicName id="58C9AB9C60053A0856E581BCBC97F818" box="[189,254,1972,1994]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="lieni">
<emphasis id="ADBD0C0D60053A0856E581BCBC97F818" box="[189,254,1972,1994]" italics="true" pageId="3" pageNumber="95">S.lieni</emphasis>
</taxonomicName>
, male; 11, 
<taxonomicName id="58C9AB9C60053A08572881BCBDD9F818" box="[368,432,1972,1994]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="lieni">
<emphasis id="ADBD0C0D60053A08572881BCBDD9F818" box="[368,432,1972,1994]" italics="true" pageId="3" pageNumber="95">S.lieni</emphasis>
</taxonomicName>
, female; 12, 
<taxonomicName id="58C9AB9C60053A08546E81BCBED3F81B" box="[566,698,1972,1993]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="tsaopiensis">
<emphasis id="ADBD0C0D60053A08546E81BCBED3F81B" box="[566,698,1972,1993]" italics="true" pageId="3" pageNumber="95">S.tsaopiensis</emphasis>
</taxonomicName>
, male; 13 &amp; 14, 
<taxonomicName id="58C9AB9C60053A08553081BCBF85F81B" box="[872,1004,1972,1993]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="tsaopiensis">
<emphasis id="ADBD0C0D60053A08553081BCBF85F81B" box="[872,1004,1972,1993]" italics="true" pageId="3" pageNumber="95">S.tsaopiensis</emphasis>
</taxonomicName>
, females, 13, normal form, 14, dark form; 15, 
<taxonomicName id="58C9AB9C60053A0856E581DBBD32F83A" box="[189,347,2002,2024]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60053A0856E581DBBD32F83A" box="[189,347,2002,2024]" italics="true" pageId="3" pageNumber="95">S. chiangchinlii</emphasis>
</taxonomicName>
, holotype male; 16 &amp; 17, 
<taxonomicName id="58C9AB9C60053A08543D81DBBF6DF83A" box="[613,772,2002,2024]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60053A08543D81DBBF6DF83A" box="[613,772,2002,2024]" italics="true" pageId="3" pageNumber="95">S. chiangchinlii</emphasis>
</taxonomicName>
, paratype females.
</paragraph>
</caption>
<caption id="CBB6809760023A0F56CF8173BD4BF81D" httpUri="https://zenodo.org/record/227488/files/figure.png" pageId="4" pageNumber="96" targetBox="[248,1345,206,1889]" targetPageId="4">
<paragraph id="9F76D01F60023A0F56CF8173BD4BF81D" blockId="4.[151,1436,1915,1999]" pageId="4" pageNumber="96">
<emphasis id="ADBD0C0D60023A0F56CF8173BD3CF842" bold="true" box="[151,341,1915,1938]" pageId="4" pageNumber="96">FIGURES 18–29.</emphasis>
Different characters of the 
<taxonomicName id="58C9AB9C60023A0F542F8173BF6DF843" box="[631,772,1915,1937]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="ADBD0C0D60023A0F542F8173BF6DF843" box="[631,772,1915,1937]" italics="true" pageId="4" pageNumber="96">Sarasaeschna</emphasis>
</taxonomicName>
of Taiwan. 18–21, male pterothorax, dorsal; 22–25, male cerci, dorsal; 26–29, male cerci, lateral; 18, 22 &amp; 26, 
<taxonomicName id="58C9AB9C60023A0F54DA8193BE8EF862" box="[642,743,1947,1968]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="pyanan">
<emphasis id="ADBD0C0D60023A0F54DA8193BE8EF862" box="[642,743,1947,1968]" italics="true" pageId="4" pageNumber="96">S. pyanan</emphasis>
</taxonomicName>
; 19, 23 &amp; 27, S. 
<taxonomicName id="58C9AB9C60023A0F55C58192BFA3F862" box="[925,970,1946,1968]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="lieni">
<emphasis id="ADBD0C0D60023A0F55C58192BFA3F862" box="[925,970,1946,1968]" italics="true" pageId="4" pageNumber="96">lieni</emphasis>
</taxonomicName>
; 20, 24 &amp; 28, 
<taxonomicName id="58C9AB9C60023A0F523C8193B886F862" box="[1124,1263,1947,1968]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="tsaopiensis">
<emphasis id="ADBD0C0D60023A0F523C8193B886F862" box="[1124,1263,1947,1968]" italics="true" pageId="4" pageNumber="96">S. tsaopiensis</emphasis>
</taxonomicName>
; 21, 25 &amp; 29, 
<taxonomicName id="58C9AB9C60023A0F53D18193BD74F81D" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60023A0F53D18193BD74F81D" italics="true" pageId="4" pageNumber="96">S. chiangchinlii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="9F76D01F60033A0E56CF82E5BE53FA96" blockId="5.[151,1436,1261,1348]" pageId="5" pageNumber="97">
<emphasis id="ADBD0C0D60033A0E56CF82E5BD3DFAD1" bold="true" box="[151,340,1261,1284]" pageId="5" pageNumber="97">FIGURES 30–35.</emphasis>
30, 
<taxonomicName id="58C9AB9C60033A0E572782E7BE60FAD6" box="[383,521,1263,1284]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="tsaopiensis">
<emphasis id="ADBD0C0D60033A0E572782E7BE60FAD6" box="[383,521,1263,1284]" italics="true" pageId="5" pageNumber="97">S. tsaopiensis</emphasis>
</taxonomicName>
, 31, 
<taxonomicName id="58C9AB9C60033A0E546482E7BEF6FAD6" box="[572,671,1263,1284]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="pyanan">
<emphasis id="ADBD0C0D60033A0E546482E7BEF6FAD6" box="[572,671,1263,1284]" italics="true" pageId="5" pageNumber="97">S. pyanan</emphasis>
</taxonomicName>
, 32 &amp; 33, 
<taxonomicName id="58C9AB9C60033A0E555382E7BFC0FAD6" box="[779,937,1262,1284]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60033A0E555382E7BFC0FAD6" box="[779,937,1262,1284]" italics="true" pageId="5" pageNumber="97">S. chiangchinlii</emphasis>
</taxonomicName>
. 30, female cerci, dorsal; 31 &amp; 32, abdominal S2 of male, lateral; 33, male penis, lateral, upside down, 34, the habitat of 
<taxonomicName id="58C9AB9C60033A0E552D8306B867FAF6" box="[885,1038,1294,1316]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60033A0E552D8306B867FAF6" box="[885,1038,1294,1316]" italics="true" pageId="5" pageNumber="97">S.chiangchinlii</emphasis>
</taxonomicName>
in Daxi, Taoyuan County, 
<collectingCountry id="E7DE908F60033A0E537B8306B904FAF6" box="[1315,1389,1294,1316]" name="Taiwan" pageId="5" pageNumber="97">Taiwan</collectingCountry>
, 35, the ovipositing female of 
<taxonomicName id="58C9AB9C60033A0E57C38326BE5AFA96" box="[411,563,1326,1348]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="chiangchinlii">
<emphasis id="ADBD0C0D60033A0E57C38326BE5AFA96" box="[411,563,1326,1348]" italics="true" pageId="5" pageNumber="97">S.chiangchinlii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9F76D01F60033A0E569F837ABC95FA06" blockId="5.[151,1436,1393,2032]" pageId="5" pageNumber="97">
PENILE STRUCTURES (Fig. 33): Configuration of penis similar to those of 
<taxonomicName id="58C9AB9C60033A0E5263837AB816FA5B" box="[1083,1151,1394,1417]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="pryeri">
<emphasis id="ADBD0C0D60033A0E5263837AB816FA5B" box="[1083,1151,1394,1417]" italics="true" pageId="5" pageNumber="97">pryeri</emphasis>
</taxonomicName>
-group 
<taxonomicName id="58C9AB9C60033A0E528A8379B907FA5B" box="[1234,1390,1393,1417]" class="Insecta" family="Aeshnidae" genus="Sarasaeschna" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="ADBD0C0D60033A0E528A8379B907FA5B" box="[1234,1390,1393,1417]" italics="true" pageId="5" pageNumber="97">Sarasaeschna</emphasis>
</taxonomicName>
(cf. Karube &amp; Yeh 2001, 
<bibRefCitation id="FB58ADEE60033A0E57CD839FBE05FA62" author="Yeh" box="[405,620,1431,1456]" pageId="5" pageNumber="97" refString="Yeh, W. C. &amp; Chen, Y. M. (2000) Descriptions of two new species of the genus Oligoaeschna from northern Taiwan, with notes on the status of the pryeri - group (Anisoptera: Aeshnidae). Odonatologica, 29 (2), 137 - 150." type="journal article" year="2000">Yeh &amp; Chen 2000</bibRefCitation>
) with flagella protruding horizontally in relation to the position of 4th segment.
</paragraph>
<paragraph id="9F76D01F60033A0E569F83D7BD85F926" blockId="5.[151,1436,1393,2032]" pageId="5" pageNumber="97">
<emphasis id="ADBD0C0D60033A0E569F83D7BDDDFA2A" bold="true" box="[199,436,1503,1528]" pageId="5" pageNumber="97">
Females (
<typeStatus id="40726EBD60033A0E576383D7BDDDFA2A" box="[315,436,1503,1528]" pageId="5" pageNumber="97" type="paratype">Paratypes</typeStatus>
</emphasis>
, Figs. 2, 4): General body maculations of females similar to those of 
<typeStatus id="40726EBD60033A0E52AA83D7B93DFA2A" box="[1266,1364,1503,1528]" pageId="5" pageNumber="97" type="holotype">holotype</typeStatus>
male, differences as follows: Wings tinged with pale brown entirely with apical half less intensely, color deeper in teneral stage. Abdomen straight and narrowed smoothly and gradually toward apex. On dorsum, S2 with large sagittate AD spot (
<figureCitation id="07F2CC9A60033A0E575C8043BDE3F9B6" box="[260,394,1611,1636]" captionStart="FIGURES 8 – 17" captionStartId="3.[151,264,1940,1963]" captionTargetBox="[268,1317,200,1887]" captionTargetId="figure@3.[256,1333,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 17. Dorsal maculations of abdominal S 2 in the Sarasaeschna of Taiwan. 8, S. pyanan, male; 9, S. pyanan, female; 10, S. lieni, male; 11, S. lieni, female; 12, S. tsaopiensis, male; 13 &amp; 14, S. tsaopiensis, females, 13, normal form, 14, dark form; 15, S. chiangchinlii, holotype male; 16 &amp; 17, S. chiangchinlii, paratype females." httpUri="https://zenodo.org/record/227487/files/figure.png" pageId="5" pageNumber="97">Figs. 16–17</figureCitation>
) and streak-like MD spots, S3 with longitudinal AD spot and streak-like MD spots; on sides, S1 &amp; S2 largely yellow, S3 with large AL spots and tiny PL spots, S4–7 with transverse AL spots, tiny in S7. Ventral side of abdomen black. Female cerci short, as long as S10 (Fig. 7). Ovipositor reddish brown and reaching to posterior margin of S10; dentigerous plate of S10 protruding backward and downward, ventrally armed with numerous closely set spinules.
</paragraph>
<paragraph id="9F76D01F60033A0E569F80F7BDDDF8EE" blockId="5.[151,1436,1393,2032]" pageId="5" pageNumber="97">
<emphasis id="ADBD0C0D60033A0E569F80F7BDAFF8CA" bold="true" box="[199,454,1791,1816]" pageId="5" pageNumber="97">Measurements (mm).</emphasis>
<typeStatus id="40726EBD60033A0E579680F7BE5FF8CA" box="[462,566,1791,1816]" pageId="5" pageNumber="97" type="holotype">Holotype</typeStatus>
male, abd. + cerci 46, hindwing 37, cerci 4.0; 
<typeStatus id="40726EBD60033A0E521F8108B8C0F8CA" box="[1095,1193,1792,1816]" pageId="5" pageNumber="97" type="paratype">Paratype</typeStatus>
females, abd. + cerci 40–42, hindwings 37–40.
</paragraph>
</subSubSection>
<subSubSection id="D7D3839460033A0E569F814FBDDFF822" pageId="5" pageNumber="97" type="materials_examined">
<paragraph id="9F76D01F60033A0E569F814FB89EF875" blockId="5.[151,1436,1393,2032]" pageId="5" pageNumber="97">
<emphasis id="ADBD0C0D60033A0E569F814FBDD4F8B2" bold="true" box="[199,445,1863,1888]" pageId="5" pageNumber="97">Materials examined.</emphasis>
All of the specimens including those listed in additional materials were collected from Daxi, Taoyuan County, N. 
<collectingCountry id="E7DE908F60033A0E57908163BE77F856" box="[456,542,1899,1924]" name="Taiwan" pageId="5" pageNumber="97">Taiwan</collectingCountry>
. 
<typeStatus id="40726EBD60033A0E54728163BEFEF856" box="[554,663,1899,1924]" pageId="5" pageNumber="97" type="holotype">
<emphasis id="ADBD0C0D60033A0E54728163BEFEF856" bold="true" box="[554,663,1899,1924]" pageId="5" pageNumber="97">Holotype</emphasis>
</typeStatus>
: ♂, 
<date id="EB77F6DF60033A0E54938164BF24F856" box="[715,845,1900,1924]" pageId="5" pageNumber="97" value="2013-03-17">17/III/2013</date>
, leg. C.L. Chiang; 
<typeStatus id="40726EBD60033A0E527E8163B8F7F856" box="[1062,1182,1899,1924]" pageId="5" pageNumber="97" type="paratype">
<emphasis id="ADBD0C0D60033A0E527E8163B8F7F856" bold="true" box="[1062,1182,1899,1924]" pageId="5" pageNumber="97">Paratypes</emphasis>
</typeStatus>
: 3♀#, 
<date id="EB77F6DF60033A0E52B68164B9F1F856" box="[1262,1432,1900,1924]" pageId="5" pageNumber="97" value="2010-04-21" valueMax="2010-04-22" valueMin="2010-04-21">21-22/IV/2010</date>
, SLC; 1♀, 
<date id="EB77F6DF60033A0E57528198BDE4F87A" box="[266,397,1936,1960]" pageId="5" pageNumber="97" value="2010-04-30">30/IV/2010</date>
, WCY. All 
<typeStatus id="40726EBD60033A0E54578198BE56F87A" box="[527,575,1936,1960]" pageId="5" pageNumber="97">type</typeStatus>
specimens will be deposited in the Insect Collection of 
<collectionCode id="F9D848DA60033A0E52EB8198B89AF875" box="[1203,1267,1936,1959]" httpUri="http://grbio.org/cool/fitg-90ys" name="Taiwan Fisheries Research Institute" pageId="5" pageNumber="97">TFRI</collectionCode>
.
</paragraph>
<paragraph id="9F76D01F60033A0E569F81BBBDDFF822" blockId="5.[151,1436,1393,2032]" pageId="5" pageNumber="97">
<emphasis id="ADBD0C0D60033A0E569F81BBBDD4F81E" bold="true" box="[199,445,1971,1996]" pageId="5" pageNumber="97">Additional materials</emphasis>
: 
<specimenCount id="89CF1B9660033A0E579481BCBD86F81E" box="[460,495,1972,1996]" pageId="5" pageNumber="97" type="generic">1♂</specimenCount>
(in alcohol), 
<date id="EB77F6DF60033A0E54DF81BCBF62F81E" box="[647,779,1972,1996]" pageId="5" pageNumber="97" value="2010-04-22">22/IV/2010</date>
, C. L. Chiang. (picked up from a spider web); 1♀, teneral, 
<date id="EB77F6DF60033A0E56CF81D0BD70F822" box="[151,281,2008,2032]" pageId="5" pageNumber="97" value="2013-03-17">17/III/2013</date>
, C. L Chiang.
</paragraph>
</subSubSection>
</treatment>
</document>