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55 KiB
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447 lines
55 KiB
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<mods:title id="3BE85ACDC4FA639B8A98922FF921E1CE">Redefinition And Generic Revision Of The North American Vaejovid Scorpion Subfamily Syntropinae Kraepelin, 1905, With Descriptions Of Six New Genera</mods:title>
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<taxonomicName id="58E52ED0FFE62E450B399C019699FA97" ID-CoL="37QJ" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[844,961,1368,1391]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="AD918941FFE62E450B399C019699FA97" bold="true" box="[844,961,1368,1391]" italics="true" pageId="21" pageNumber="21">Balsateres</emphasis>
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,
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<taxonomicNameLabel id="B6A2343AFFE62E450BA49C009166FA96" box="[977,1086,1369,1390]" pageId="21" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="9F5A5553FFE62E450A989C2E91C4FA74" blockId="21.[749,1180,1368,1421]" box="[749,1180,1399,1421]" pageId="21" pageNumber="21">
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<heading id="C412E23FFFE62E450A989C2E91C4FA74" box="[749,1180,1399,1421]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="21" reason="2">
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<figureCitation id="07DE49D6FFE62E450A989C2E9605FA74" box="[749,861,1399,1421]" captionStart="Fig" captionStartId="7.[150,188,1091,1110]" captionTargetBox="[128,1229,211,1059]" captionTargetId="figure-215@7.[128,1229,211,1060]" captionTargetPageId="7" captionText="Fig. 3. Map of southwestern North America, plotting known locality records for two genera in the vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, based on data collected in the present study: Balsateres, gen. nov. (circles); Paravaejovis Williams, 1980 (squares)." figureDoi="http://doi.org/10.5281/zenodo.4611875" httpUri="https://zenodo.org/record/4611875/files/figure.png" pageId="21" pageNumber="21">Figures 3</figureCitation>
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,
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<figureCitation id="07DE49D6FFE62E450B199C219624FA75" box="[876,892,1400,1421]" captionStart="Fig" captionStartId="13.[139,178,1308,1328]" captionTargetBox="[341,1397,490,1278]" captionTargetId="figure-0@13.[341,1397,490,1278]" captionTargetPageId="13" captionText="Fig. 7. Most parsimonious tree (length: 10894.593, CI: 0.190, RI: 0.628, fit: 244.08), obtained by simultaneous phylogenetic analysis of 250 qualitative and quantitative morphological characters and 4221 aligned DNA nucleotides from three mitochondrial and two nuclear gene markers, for 18 genera and 61 species in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905. Analyses were conducted with an implied weighting regime (k 518) that maximized average support (González-Santillán and Prendini, in press). Synapomorphies are optimized with accelerated transformation. Black bars indicate uniquely derived apomorphic states, white bars indicate parallel derivations of apomorphic states, numbers above indicate characters, and numbers below indicate states (appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.4611887" httpUri="https://zenodo.org/record/4611887/files/figure.png" pageId="21" pageNumber="21">7</figureCitation>
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,
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<figureCitation id="07DE49D6FFE62E450BFF9C2E96E5FA75" box="[906,957,1399,1421]" captionStart="Fig" captionStartId="28.[116,155,1616,1635]" captionTargetBox="[95,1191,211,1586]" captionTargetId="figure-1@28.[95,1191,211,1586]" captionTargetPageId="28" captionText="Fig. 16. Syntropinae Kraepelin, 1905, carapace, dorsal aspect, illustrating surface ornamentation. A. Balsateres cisnerosi (Ponce-Saavedra and Sissom, 2004), comb. nov., 8 (IBUNAM). B. Chihuahuanus crassimanus (Pocock, 1898), comb. nov., 8 (AMNH). C. Kochius bruneus (Williams, 1970), 8 (AMNH). D. Konetontli acapulco (Armas and Martín-Frías, 2001), comb. nov., holotype 8 (IBUNAM). Scale bars 5 1 mm (A–C), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.4611915" httpUri="https://zenodo.org/record/4611915/files/figure.png" pageId="21" pageNumber="21">16A</figureCitation>
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,
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<figureCitation id="07DE49D6FFE62E450BBF9C2E96A6FA74" box="[970,1022,1399,1420]" captionStart="Fig" captionStartId="37.[150,188,1460,1479]" captionTargetBox="[139,1213,218,1430]" captionTargetId="figure-75@37.[128,1228,211,1430]" captionTargetPageId="37" captionText="Fig. 20. Syntropinae Kraepelin, 1905, dextral pedipalp chela movable finger, dorsal aspect, illustrating dentition. A. Balsateres cisnerosi (Ponce-Saavedra and Sissom, 2004), comb. nov., 8 (IBUNAM). B. Thorellius intrepidus (Thorell, 1876), 8 (AMNH). C. Vizcaino viscainensis (Williams, 1970), comb. nov., 8 (AMNH). Scale bars 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4611927" httpUri="https://zenodo.org/record/4611927/files/figure.png" pageId="21" pageNumber="21">20A</figureCitation>
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,
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<figureCitation id="07DE49D6FFE62E450C7E9C2E9166FA74" box="[1035,1086,1399,1420]" captionStart="Fig" captionStartId="57.[150,188,1645,1664]" captionTargetBox="[227,1132,211,1594]" captionTargetId="figure-0@57.[218,1137,211,1615]" captionTargetPageId="57" captionText="Fig. 28. Syntropinae Kraepelin, 1905, metasomal segments I–V and telson, lateral aspect, illustrating carinae. A. Balsateres cisnerosi (Ponce-Saavedra and Sissom, 2004), comb. nov., 8 (IBUNAM). B. Konetontli acapulco (Armas and Martín-Frías, 2004), comb. nov., holotype 8 (IBUNAM). C. Paravaejovis pumilis (Williams, 1980), 8 (AMNH). Scale bars 5 2.5 mm (A), 1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.4611953" httpUri="https://zenodo.org/record/4611953/files/figure.png" pageId="21" pageNumber="21">28A</figureCitation>
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;
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<tableCitation id="D26760E8FFE62E450C399C2E91C4FA74" box="[1100,1180,1399,1421]" captionStart="TABLE 1" captionStartId="6.[590,672,214,233]" captionTargetBox="[93,1183,531,993]" captionTargetPageId="6" captionText="TABLE 1 Currently recognized genera in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with countries, states and geographical regions from which they have been recorded Abbreviations: Mex. (Mexico): Ags (Aguascalientes), B.C. (Baja California), B.C.S. (Baja California Sur), Chis (Chiapas), Chih. (Chihuahua), Coah. (Coahuila), Col. (Colima), Dgo (Durango), Gto (Guanajuato), Gro (Guerrero), Hgo (Hidalgo), Jal. (Jalisco), Edomex. (Estado de México), Mich. (Michoacán), Mor. (Morelos), Nay. (Nayarit), N.L. (Nuevo León), Oax. (Oaxaca), Pue. (Puebla), Qro (Querétaro), Sin. (Sinaloa), Son. (Sonora), Tamps (Tamaulipas), Tlax. (Tlaxcala), Ver. (Veracruz), Zac. (Zacatecas); U.S.A.: AZ (Arizona), CA (California), NM (New Mexico), NV (Nevada), OR (Oregon), TX (Texas), UT (Utah); Balsas Depression (BD), Baja California Peninsula (BCP), Chihuahuan Desert (CD), Great Basin Desert (GBD), Mojave Desert (MD), Pacific Coast (PC), Sonoran Desert (SD), Trans-Mexican Volcanic Belt (TMVB)." httpUri="http://table.plazi.org/id/CB9A05DBFFF52E560A3B998F95B5FE08" pageId="21" pageNumber="21" tableUuid="CB9A05DBFFF52E560A3B998F95B5FE08">table 1</tableCitation>
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<emphasis id="AD918941FFE62E450AC89CFC9623FA4F" box="[701,891,1445,1463]" italics="true" pageId="21" pageNumber="21">Vaejovis cisnerosi</emphasis>
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<bibRefCitation id="FB7428A2FFE62E450BF99CFD9651FA2A" author="Ponce-Saavedra, J. & W. D. Sissom" pageId="21" pageNumber="21" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004</bibRefCitation>
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[
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<emphasis id="AD918941FFE62E450B6B9CE7966AFA2B" box="[798,818,1470,1491]" italics="true" pageId="21" pageNumber="21">=</emphasis>
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<taxonomicName id="58E52ED0FFE62E450B349CE696CFFA15" ID-CoL="KJ2S" authority="(Ponce-Saavedra and Sissom, 2004)" authorityName="González-Santillán & Prendini" authorityYear="2013" baseAuthorityName="Ponce-Saavedra and Sissom" baseAuthorityYear="2004" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="cisnerosi" status="comb. nov.">
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<emphasis id="AD918941FFE62E450B349CE69157FA2A" box="[833,1039,1471,1491]" italics="true" pageId="21" pageNumber="21">Balsateres cisnerosi</emphasis>
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(
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<bibRefCitation id="FB7428A2FFE62E450C519CE696C9FA15" author="Ponce-Saavedra, J. & W. D. Sissom" pageId="21" pageNumber="21" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004</bibRefCitation>
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)
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,
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<taxonomicNameLabel id="B6A2343AFFE62E450BD79C839141FA15" box="[930,1049,1498,1517]" pageId="21" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
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],
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species, by monotypy.
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<taxonomicName id="58E52ED0FFE62E450AC89F49967BF9DC" ID-CoL="7W89" authorityName="Soleglad and Fet" authorityYear="2008" box="[701,803,1552,1572]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="AD918941FFE62E450AC89F49967BF9DC" box="[701,803,1552,1572]" italics="true" pageId="21" pageNumber="21">Thorellius</emphasis>
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(part):
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<bibRefCitation id="FB7428A2FFE62E450B089F4991C3F9DC" author="Soleglad, M. E. & V. Fet" box="[893,1179,1552,1572]" pageId="21" pageNumber="21" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet, 2008: 1</bibRefCitation>
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, 95, 102;
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<treatmentCitation id="1E447342FFE62E450B7E9F75916EF9C6" author="Ayrey, R. F. & M. E. Soleglad" box="[779,1078,1579,1599]" page="1" pageId="21" pageNumber="21" year="2011">
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<bibRefCitation id="FB7428A2FFE62E450B7E9F75916EF9C6" author="Ayrey, R. F. & M. E. Soleglad" box="[779,1078,1579,1599]" pageId="21" pageNumber="21" pagination="1 - 15" refId="ref36088" refString="Ayrey, R. F., and M. E. Soleglad. 2011. A new species of Vaejovis from Prescott, Arizona (Scorpiones: Vaejovidae). Euscorpius 114: 1 - 15." type="journal article" year="2011">Ayrey and Soleglad, 2011: 1</bibRefCitation>
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.
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</paragraph>
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<paragraph id="9F5A5553FFE62E450AC89F1E96E5F98D" blockId="21.[701,1228,1444,1653]" pageId="21" pageNumber="21">
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<taxonomicName id="58E52ED0FFE62E450AC89F1E9625F9A2" authorityName="Thorell" authorityYear="1876" box="[701,893,1606,1626]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="intrepidus">
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<emphasis id="AD918941FFE62E450AC89F1E9625F9A2" box="[701,893,1606,1626]" italics="true" pageId="21" pageNumber="21">Vaejovis intrepidus</emphasis>
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group (part):
|
||
<treatmentCitation id="1E447342FFE62E450C649F1E96E2F98D" author="Santibanez-Lopez, C. & W. D. Sissom" page="52" pageId="21" pageNumber="21" year="2010">
|
||
<bibRefCitation id="FB7428A2FFE62E450C649F1E96E2F98D" author="Santibanez-Lopez, C. & W. D. Sissom" pageId="21" pageNumber="21" pagination="49 - 58" refId="ref38931" refString="Santibanez-Lopez, C., and W. D. Sissom. 2010. A new species of the Vaejovis eusthenura group in Oaxaca, Mexico (Scorpiones: Vaejovidae). Zootaxa 2493: 49 - 58." type="journal article" year="2010">Santibáñez-López and Sissom, 2010: 52</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFE62E480AAF9FD5940AFEBA" lastPageId="24" lastPageNumber="24" pageId="21" pageNumber="21" type="etymology">
|
||
<paragraph id="9F5A5553FFE62E480AAF9FD5940AFEBA" blockId="21.[701,1228,1676,1759]" lastBlockId="24.[93,620,213,1759]" lastPageId="24" lastPageNumber="24" pageId="21" pageNumber="21">
|
||
ETYMOLOGY: The name
|
||
<taxonomicName id="58E52ED0FFE62E450C649FD491DEF95C" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[1041,1158,1677,1700]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFE62E450C649FD491DEF95C" box="[1041,1158,1677,1700]" italics="true" pageId="21" pageNumber="21">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFE62E450CEE9FD697ABF939" pageId="21" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, a compound word, masculine in gender, is derived from the Balsas Depression, to which the genus is endemic, and the Latin adjective
|
||
<emphasis id="AD918941FFEB2E4808AD99AA9456FEF0" box="[216,270,243,264]" italics="true" pageId="24" pageNumber="24">teres</emphasis>
|
||
(‘‘smooth’’ or ‘‘polished’’) which refers to the glabrous integument typical of this genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="CB9A05DBFFE52E4608019F3295E6F936" ID-DOI="http://doi.org/10.5281/zenodo.4611907" ID-Zenodo-Dep="4611907" httpUri="https://zenodo.org/record/4611907/files/figure.png" pageId="22" pageNumber="22" startId="22.[116,155,1643,1662]" targetBox="[229,1058,211,1613]" targetPageId="22">
|
||
<paragraph id="9F5A5553FFE52E4608019F3295E6F936" blockId="22.[93,1193,1640,1743]" pageId="22" pageNumber="22">
|
||
Fig. 13.
|
||
<taxonomicName id="58E52ED0FFE52E4608AE9F32972FF986" authority="Soleglad et al., 2007" authorityName="Soleglad" authorityYear="2007" box="[219,631,1642,1663]" class="Arachnida" family="Vaejovidae" genus="Syntropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="22" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="AD918941FFE52E4608AE9F3294C4F986" box="[219,412,1642,1662]" italics="true" pageId="22" pageNumber="22">Syntropis williamsi</emphasis>
|
||
<bibRefCitation id="FB7428A2FFE52E4609D69F32972FF986" author="Soleglad, M. E. & G. Lowe & V. Fet" box="[419,631,1643,1663]" pageId="22" pageNumber="22" pagination="119 - 136" refId="ref39688" refString="Soleglad, M. E., G. Lowe, and V. Fet. 2007. Systematic observations on the scorpion genus Syntropis, with descriptions of two new species (Scorpiones: Vaejovidae). Boletin de la Sociedad Entomologica Aragonesa 40: 119 - 136." type="journal article" year="2007">Soleglad et al., 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♂ (AMNH), dextral leg II, basitarsus and telotarsus, dorsal (
|
||
<emphasis id="AD918941FFE52E4608D89FDF95E7F961" bold="true" box="[173,191,1670,1689]" pageId="22" pageNumber="22">A</emphasis>
|
||
) and ventral (
|
||
<emphasis id="AD918941FFE52E4609229FDF943FF961" bold="true" box="[343,359,1670,1689]" pageId="22" pageNumber="22">B</emphasis>
|
||
) aspects, illustrating rows of spinules. Abbreviations:
|
||
<emphasis id="AD918941FFE52E460BD69FDF96E2F961" bold="true" box="[931,954,1670,1689]" pageId="22" pageNumber="22">pv</emphasis>
|
||
, prolateral ventral;
|
||
<emphasis id="AD918941FFE52E460CF89FDF91FAF961" bold="true" box="[1165,1186,1670,1689]" pageId="22" pageNumber="22">rd</emphasis>
|
||
, retrolateral dorsal;
|
||
<emphasis id="AD918941FFE52E4609529FF8946FF94C" bold="true" box="[295,311,1697,1716]" pageId="22" pageNumber="22">rl</emphasis>
|
||
, retrolateral;
|
||
<emphasis id="AD918941FFE52E4609B39FF89482F94C" bold="true" box="[454,474,1697,1716]" pageId="22" pageNumber="22">rv</emphasis>
|
||
, retrolateral ventral;
|
||
<emphasis id="AD918941FFE52E460ACF9FF89789F94C" bold="true" box="[698,721,1697,1716]" pageId="22" pageNumber="22">vd</emphasis>
|
||
, ventral distal;
|
||
<emphasis id="AD918941FFE52E460B019FF896C9F94C" bold="true" box="[884,913,1697,1716]" pageId="22" pageNumber="22">vm</emphasis>
|
||
, ventral median. Scale bar
|
||
<emphasis id="AD918941FFE52E4608289FE39529F937" box="[93,113,1722,1743]" italics="true" pageId="22" pageNumber="22">=</emphasis>
|
||
1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CB9A05DBFFE42E4708E39F3497F1F928" ID-DOI="http://doi.org/10.5281/zenodo.4611909" ID-Zenodo-Dep="4611909" httpUri="https://zenodo.org/record/4611909/files/figure.png" pageId="23" pageNumber="23" startId="23.[150,188,1645,1664]" targetBox="[263,1092,211,1615]" targetPageId="23">
|
||
<paragraph id="9F5A5553FFE42E4708E39F3497F1F928" blockId="23.[128,1228,1642,1745]" pageId="23" pageNumber="23">
|
||
Fig. 14.
|
||
<taxonomicName id="58E52ED0FFE42E47088B9F3497C2F978" authority="Soleglad et al., 2007" authorityName="Soleglad" authorityYear="2007" box="[254,666,1644,1665]" class="Arachnida" family="Vaejovidae" genus="Syntropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="23" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="AD918941FFE42E47088B9F3494E7F978" box="[254,447,1644,1664]" italics="true" pageId="23" pageNumber="23">Syntropis williamsi</emphasis>
|
||
<bibRefCitation id="FB7428A2FFE42E4709B39F3497C2F978" author="Soleglad, M. E. & G. Lowe & V. Fet" box="[454,666,1645,1665]" pageId="23" pageNumber="23" pagination="119 - 136" refId="ref39688" refString="Soleglad, M. E., G. Lowe, and V. Fet. 2007. Systematic observations on the scorpion genus Syntropis, with descriptions of two new species (Scorpiones: Vaejovidae). Boletin de la Sociedad Entomologica Aragonesa 40: 119 - 136." type="journal article" year="2007">Soleglad et al., 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♂ (AMNH), dextral leg II, basitarsus and telotarsus, dorsal (
|
||
<emphasis id="AD918941FFE42E4708A59FD195BAF963" bold="true" box="[208,226,1672,1691]" pageId="23" pageNumber="23">A</emphasis>
|
||
) and ventral (
|
||
<emphasis id="AD918941FFE42E47090F9FD194D2F963" bold="true" box="[378,394,1672,1691]" pageId="23" pageNumber="23">B</emphasis>
|
||
) aspects, illustrating distribution of macrosetae. Abbreviations:
|
||
<emphasis id="AD918941FFE42E470C449FD1911CF963" bold="true" box="[1073,1092,1672,1691]" pageId="23" pageNumber="23">D</emphasis>
|
||
, dorsal;
|
||
<emphasis id="AD918941FFE42E470CD79FD1919DF963" bold="true" box="[1186,1221,1672,1691]" pageId="23" pageNumber="23">PD</emphasis>
|
||
, prolateral dorsal;
|
||
<emphasis id="AD918941FFE42E47094C9FFB943AF94D" bold="true" box="[313,354,1698,1717]" pageId="23" pageNumber="23">PM</emphasis>
|
||
, prolateral median;
|
||
<emphasis id="AD918941FFE42E470A409FFB970FF94D" bold="true" box="[565,599,1698,1717]" pageId="23" pageNumber="23">PV</emphasis>
|
||
, prolateral ventral;
|
||
<emphasis id="AD918941FFE42E470B539FFB9612F94D" bold="true" box="[806,842,1698,1717]" pageId="23" pageNumber="23">RD</emphasis>
|
||
, retrolateral dorsal;
|
||
<emphasis id="AD918941FFE42E470C559FFB9112F94D" bold="true" box="[1056,1098,1698,1717]" pageId="23" pageNumber="23">RM</emphasis>
|
||
, retrolateral median;
|
||
<emphasis id="AD918941FFE42E4708AE9FE495A6F928" bold="true" box="[219,254,1725,1744]" pageId="23" pageNumber="23">RV</emphasis>
|
||
, retrolateral ventral. Scale bar
|
||
<emphasis id="AD918941FFE42E470A3D9FE59704F929" box="[584,604,1724,1745]" italics="true" pageId="23" pageNumber="23">=</emphasis>
|
||
1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D7FF06D8FFEB2E48080F981095E1FA63" pageId="24" pageNumber="24" type="diagnosis">
|
||
<paragraph id="9F5A5553FFEB2E48080F98109703FCA9" blockId="24.[93,620,213,1759]" pageId="24" pageNumber="24">
|
||
DIAGNOSIS:
|
||
<taxonomicName id="58E52ED0FFEB2E480967981394DFFE99" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[274,391,330,353]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E480967981394DFFE99" box="[274,391,330,353]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809E998129748FE98" box="[412,528,331,352]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, is distinguished from other genera of
|
||
<taxonomicName id="58E52ED0FFEB2E4809AB98319733FE86" authorityName="KRAEPELIN" authorityYear="1905" box="[478,619,360,382]" class="Arachnida" family="Vaejovidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="64" phylum="Arthropoda" rank="subFamily" subFamily="Syntropinae">Syntropinae</taxonomicName>
|
||
by the smooth dl carinae on metasomal segments I–IV. The carinae and intercarinal surfaces of the carapace (fig. 16A), pedipalp chela, patella, tergites, metasoma, and telson (fig. 28A) are smooth in this genus. Counts of macrosetae on the carinae of metasomal segments I–IV are also greatly reduced, as follows: dl, 0/0:0/0:0/0:1/1:4/4; lm, 0/0:0/0:0/ 0:0/0:2/2; vl and vsm, 1/1:1/1:1/1:1/1:3/3. Two or more macrosetae are present on these carinae in all other
|
||
<taxonomicName id="58E52ED0FFEB2E4809299BF094B5FD47" authorityName="KRAEPELIN" authorityYear="1905" box="[348,493,681,703]" class="Arachnida" family="Vaejovidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="64" phylum="Arthropoda" rank="subFamily" subFamily="Syntropinae">Syntropinae</taxonomicName>
|
||
, including
|
||
<taxonomicName id="58E52ED0FFEB2E4808289B9F9597FD25" authorityName="Soleglad and Fet" authorityYear="2008" box="[93,207,710,733]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E4808289B9F9597FD25" box="[93,207,710,733]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
, which also presents low setal counts, e.g.,
|
||
<taxonomicName id="58E52ED0FFEB2E48088C9BBA95C5FCEF" authority="(Pocock, 1989)" baseAuthorityName="Pocock" baseAuthorityYear="1989" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="cristimanus">
|
||
<emphasis id="AD918941FFEB2E48088C9BBA94A1FD01" box="[249,505,739,762]" italics="true" pageId="24" pageNumber="24">Thorellius cristimanus</emphasis>
|
||
(Pocock, 1989)
|
||
</taxonomicName>
|
||
with the following counts on segments I– V: dl, 2/2:3/3:3/3:3/3:7/6; lm, 1/1:3/3:3/3:4/4:4/4; vl, 2/2:3/3:3/3:3/3:7/7; vsm, 3/3:3/3:3/3:3/3:5/5.
|
||
</paragraph>
|
||
<paragraph id="9F5A5553FFEB2E48080F9A0095E1FA63" blockId="24.[93,620,213,1759]" pageId="24" pageNumber="24">
|
||
<taxonomicName id="58E52ED0FFEB2E48080F9A0095B7FC88" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[122,239,857,880]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E48080F9A0095B7FC88" box="[122,239,857,880]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809789A0394D2FC97" box="[269,394,858,879]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, is most closely related to
|
||
<taxonomicName id="58E52ED0FFEB2E4808949A2F940BFC75" authorityName="Soleglad and Fet" authorityYear="2008" box="[225,339,886,909]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E4808949A2F940BFC75" box="[225,339,886,909]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
(fig. 7), with which it shares broad pedipalpal and metasomal carinae and, with
|
||
<taxonomicName id="58E52ED0FFEB2E48095B9AE99597FC1C" authority="(Thorell, 1867)" baseAuthorityName="Thorell" baseAuthorityYear="1867" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="intrepidus">
|
||
<emphasis id="AD918941FFEB2E48095B9AE9974EFC3F" box="[302,534,944,967]" italics="true" pageId="24" pageNumber="24">Thorellius intrepidus</emphasis>
|
||
(Thorell, 1867)
|
||
</taxonomicName>
|
||
, double basal prolateral denticles on the movable finger of the pedipalp chela (fig. 20A, B). However, all carinae are smooth in
|
||
<taxonomicName id="58E52ED0FFEB2E48080C9D7C95B6FBC4" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[121,238,1061,1084]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E48080C9D7C95B6FBC4" box="[121,238,1061,1084]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E48088E9D7E943FFBC4" box="[251,359,1063,1084]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, instead of moderately to densely granular, as in
|
||
<taxonomicName id="58E52ED0FFEB2E4809C19D1A977EFBA2" authorityName="Soleglad and Fet" authorityYear="2008" box="[436,550,1091,1114]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E4809C19D1A977EFBA2" box="[436,550,1091,1114]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
. The distinct proximal gap, evident when the pedipalp chela fingers are closed, in both sexes of
|
||
<taxonomicName id="58E52ED0FFEB2E48080E9DC395B5FB49" authorityName="Soleglad and Fet" authorityYear="2008" box="[123,237,1178,1201]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E48080E9DC395B5FB49" box="[123,237,1178,1201]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
, is absent in
|
||
<taxonomicName id="58E52ED0FFEB2E4809F59DC394ADFB49" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[384,501,1178,1201]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E4809F59DC394ADFB49" box="[384,501,1178,1201]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E480A779DC29733FB48" box="[514,619,1179,1200]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
The two genera also differ in base coloration and infuscation.
|
||
<taxonomicName id="58E52ED0FFEB2E48094D9D8C94F5FB14" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[312,429,1237,1260]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E48094D9D8C94F5FB14" box="[312,429,1237,1260]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809BD9D8F9719FB13" box="[456,577,1238,1259]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, is yellowish and almost immaculate, except for the ocular tubercle, which is outlined by dark infuscation, whereas
|
||
<taxonomicName id="58E52ED0FFEB2E4809169C75948DFABB" authorityName="Soleglad and Fet" authorityYear="2008" box="[355,469,1324,1347]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E4809169C75948DFABB" box="[355,469,1324,1347]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
tend to be darker and reddish in color, with various degrees of infuscation on the carapace and tergites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFEB2E48080F9CFB9734F991" pageId="24" pageNumber="24" type="discussion">
|
||
<paragraph id="9F5A5553FFEB2E48080F9CFB9484F9D6" blockId="24.[93,620,213,1759]" pageId="24" pageNumber="24">
|
||
<taxonomicName id="58E52ED0FFEB2E48080F9CFB95B7FA41" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[122,239,1442,1465]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E48080F9CFB95B7FA41" box="[122,239,1442,1465]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809769CFA942DFA40" box="[259,373,1443,1464]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, resembles
|
||
<taxonomicName id="58E52ED0FFEB2E480A779CFD95CFFA2E" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" class="Arachnida" family="Vaejovidae" genus="Mesomexovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E480A779CFD95CFFA2E" italics="true" pageId="24" pageNumber="24">Mesomexovis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4808C39C98946BFA2E" box="[182,307,1473,1494]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, in possessing obsolete ventral carinae on metasomal segments I– IV, but species of the latter genus differ in being more setose and infuscate.
|
||
</paragraph>
|
||
<paragraph id="9F5A5553FFEB2E48080F9F6D9734F991" blockId="24.[93,620,213,1759]" pageId="24" pageNumber="24">
|
||
INCLUDED SPECIES:
|
||
<taxonomicName id="58E52ED0FFEB2E4809F59F6C94B1F991" authority="(Ponce-Saavedra and Sissom, 2004)" authorityName="González-Santillán & Prendini" authorityYear="2013" baseAuthorityName="Ponce-Saavedra and Sissom" baseAuthorityYear="2004" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="cisnerosi" status="comb. nov.">
|
||
<emphasis id="AD918941FFEB2E4809F59F6C9733F9B4" box="[384,619,1589,1612]" italics="true" pageId="24" pageNumber="24">Balsateres cisnerosi</emphasis>
|
||
(
|
||
<bibRefCitation id="FB7428A2FFEB2E4808109F0A94BAF991" author="Ponce-Saavedra, J. & W. D. Sissom" box="[101,482,1619,1641]" pageId="24" pageNumber="24" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809869F0A9734F991" box="[499,620,1619,1641]" pageId="24" pageNumber="24" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFEB2E48080F9F36942AF927" pageId="24" pageNumber="24" type="distribution">
|
||
<paragraph id="9F5A5553FFEB2E48080F9F36942AF927" blockId="24.[93,620,213,1759]" pageId="24" pageNumber="24">
|
||
DISTRIBUTION:
|
||
<taxonomicName id="58E52ED0FFEB2E4809489F2894EAF970" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[317,434,1649,1672]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E4809489F2894EAF970" box="[317,434,1649,1672]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E4809BE9F2B971BF97F" box="[459,579,1650,1671]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, is endemic to the Balsas Depression of
|
||
<collectingRegion id="5D219BB1FFEB2E480A619FD69733F95D" box="[532,619,1679,1701]" country="Mexico" name="Mexico" pageId="24" pageNumber="24">Mexico</collectingRegion>
|
||
and recorded only from
|
||
<collectingRegion id="5D219BB1FFEB2E4809FB9FF59733F93A" box="[398,619,1708,1730]" country="Mexico" name="Mexico" pageId="24" pageNumber="24">Estado de México</collectingRegion>
|
||
and
|
||
<collectingRegion id="5D219BB1FFEB2E4808E79F90944DF927" box="[146,277,1737,1759]" country="Mexico" name="Michoacan de Ocampo" pageId="24" pageNumber="24">Michoacán</collectingRegion>
|
||
(fig. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFEB2E480ACD998A966AFCCC" pageId="24" pageNumber="24" type="biology_ecology">
|
||
<paragraph id="9F5A5553FFEB2E480ACD998A966AFCCC" blockId="24.[667,1193,211,1438]" pageId="24" pageNumber="24">
|
||
NATURAL HISTORY:
|
||
<taxonomicName id="58E52ED0FFEB2E480BC7998D917FFF13" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[946,1063,212,235]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E480BC7998D917FFF13" box="[946,1063,212,235]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E480C43998C91FCFF12" box="[1078,1188,213,234]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, inhabits tropical deciduous forest at altitudes between
|
||
<quantity id="581DF8B6FFEB2E480B79985696C0FEDD" box="[780,920,271,293]" metricMagnitude="2" metricUnit="m" metricValue="4.335" metricValueMax="6.72" metricValueMin="1.95" pageId="24" pageNumber="24" unit="m" value="433.5" valueMax="672.0" valueMin="195.0">
|
||
195–
|
||
<specimenCount id="89E39EDAFFEB2E480B34985696C0FEDD" box="[833,920,271,293]" pageId="24" pageNumber="24" type="generic">672 m</specimenCount>
|
||
</quantity>
|
||
. Specimens were observed at night, doorkeeping at the entrances of their burrows in open, sandy areas, and excavated from the burrows (
|
||
<bibRefCitation id="FB7428A2FFEB2E480B98983E9636FE63" author="Ponce-Saavedra, J. & W. D. Sissom" pageId="24" pageNumber="24" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004</bibRefCitation>
|
||
). The glabrous integument and burrowing biology in sandy habitats suggest that this species is psammophilous although the leg setation is atypical for the psammophilous ecomorphotype, because the dorsal and retrolateral dorsal macrosetae are arranged into two separate rows, not forming a setal comb. This monotypic genus was collected at the same locality as another monotypic syntropine genus,
|
||
<taxonomicName id="58E52ED0FFEB2E480C7F9BD49134FD5A" authorityName="Francke and Ponce-Saavedra" authorityYear="2010" box="[1034,1132,653,674]" class="Arachnida" family="Vaejovidae" genus="Kuarapu" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E480C7F9BD49134FD5A" box="[1034,1132,653,674]" italics="true" pageId="24" pageNumber="24">Kuarapu</emphasis>
|
||
</taxonomicName>
|
||
. Although sympatric, these genera are allotopic:
|
||
<taxonomicName id="58E52ED0FFEB2E480AEE9B9F9648FD25" authority="González-Santillán & Prendini, 2013" authorityName="González-Santillán & Prendini" authorityYear="2013" box="[667,784,710,733]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="AD918941FFEB2E480AEE9B9F9648FD25" box="[667,784,710,733]" italics="true" pageId="24" pageNumber="24">Balsateres</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E480B579B9E96CAFD24" box="[802,914,711,732]" pageId="24" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, was collected in open, sandy areas, whereas
|
||
<taxonomicName id="58E52ED0FFEB2E480BD59BBD915BFD01" authorityName="Francke and Ponce-Saavedra" authorityYear="2010" box="[928,1027,740,761]" class="Arachnida" family="Vaejovidae" genus="Kuarapu" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E480BD59BBD915BFD01" box="[928,1027,740,761]" italics="true" pageId="24" pageNumber="24">Kuarapu</emphasis>
|
||
</taxonomicName>
|
||
was collected on road-cuts, suggesting a lithophilous ecomorphotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFEB2E480ACD9A62910CFB37" pageId="24" pageNumber="24" type="discussion">
|
||
<paragraph id="9F5A5553FFEB2E480ACD9A62910CFB37" blockId="24.[667,1193,211,1438]" pageId="24" pageNumber="24">
|
||
REMARKS: Due to the unique morphology of this species,
|
||
<bibRefCitation id="FB7428A2FFEB2E480B289A0397B9FC75" author="Ponce-Saavedra, J. & W. D. Sissom" pageId="24" pageNumber="24" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom (2004)</bibRefCitation>
|
||
refrained from assigning it to one of the five species groups of
|
||
<taxonomicName id="58E52ED0FFEB2E480BEA9ACC96A7FC52" authorityName="C. L. Koch" authorityYear="1836" box="[927,1023,917,938]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E480BEA9ACC96A7FC52" box="[927,1023,917,938]" italics="true" pageId="24" pageNumber="24">Vaejovis</emphasis>
|
||
</taxonomicName>
|
||
recognized at the time.
|
||
<bibRefCitation id="FB7428A2FFEB2E480B7F9AEB917AFC30" author="Soleglad, M. E. & V. Fet" box="[778,1058,946,968]" pageId="24" pageNumber="24" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet (2008)</bibRefCitation>
|
||
transferred it to
|
||
<taxonomicName id="58E52ED0FFEB2E480A949A97960BFC1D" authorityName="Soleglad and Fet" authorityYear="2008" box="[737,851,974,997]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E480A949A97960BFC1D" box="[737,851,974,997]" italics="true" pageId="24" pageNumber="24">Thorellius</emphasis>
|
||
</taxonomicName>
|
||
, a name devised for
|
||
<bibRefCitation id="FB7428A2FFEB2E480C1D9A96961AFBFA" author="Hoffmann, C. C." pageId="24" pageNumber="24" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann’s (1931)</bibRefCitation>
|
||
‘‘second section’’ of
|
||
<taxonomicName id="58E52ED0FFEB2E480C379AB491FAFBFA" authorityName="C. L. Koch" authorityYear="1836" box="[1090,1186,1005,1026]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AD918941FFEB2E480C379AB491FAFBFA" box="[1090,1186,1005,1026]" italics="true" pageId="24" pageNumber="24">Vaejovis</emphasis>
|
||
</taxonomicName>
|
||
, later termed the
|
||
<taxonomicName id="58E52ED0FFEB2E480B139D50968DFBD8" baseAuthorityName="Thorell" baseAuthorityYear="1867" box="[870,981,1033,1056]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="intrepidus">
|
||
<emphasis id="AD918941FFEB2E480B139D50968DFBD8" box="[870,981,1033,1056]" italics="true" pageId="24" pageNumber="24">intrepidus</emphasis>
|
||
</taxonomicName>
|
||
group by
|
||
<bibRefCitation id="FB7428A2FFEB2E480C2C9D5397BDFBC5" author="Sissom, W. D." pageId="24" pageNumber="24" pagination="179 - 187" refId="ref39026" refString="Sissom, W. D. 1989. Redescription of Vaejovis occidentalis Hoffmann with a revised diagnosis for Vaejovis subcristatus Pocock (Scorpiones: Vaejovidae). Revue Arachnologique 8: 179 - 187." type="journal article" year="1989">Sissom (1989)</bibRefCitation>
|
||
, without quantitatively testing its monophyly or composition.
|
||
<bibRefCitation id="FB7428A2FFEB2E480BDD9D1D9663FB80" author="Santibanez-Lopez, C. & W. D. Sissom" pageId="24" pageNumber="24" pagination="49 - 58" refId="ref38931" refString="Santibanez-Lopez, C., and W. D. Sissom. 2010. A new species of the Vaejovis eusthenura group in Oaxaca, Mexico (Scorpiones: Vaejovidae). Zootaxa 2493: 49 - 58." type="journal article" year="2010">Santibáñez-López and Sissom (2010)</bibRefCitation>
|
||
also referred this species to the
|
||
<taxonomicName id="58E52ED0FFEB2E480AEE9D279652FB6D" baseAuthorityName="Thorell" baseAuthorityYear="1867" box="[667,778,1150,1173]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="intrepidus">
|
||
<emphasis id="AD918941FFEB2E480AEE9D279652FB6D" box="[667,778,1150,1173]" italics="true" pageId="24" pageNumber="24">intrepidus</emphasis>
|
||
</taxonomicName>
|
||
group. The creation of a monotypic genus is merited by the unique diagnostic character combination of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7FF06D8FFEB2E480ACD9D8F96A7FA65" pageId="24" pageNumber="24" type="materials_examined">
|
||
<paragraph id="9F5A5553FFEB2E480ACD9D8F96A7FA65" blockId="24.[667,1193,211,1438]" pageId="24" pageNumber="24">
|
||
MATERIAL EXAMINED:
|
||
<materialsCitation id="2F8D5F0EFFEB2E480BA99D8E96A7FA65" ID-GBIF-Occurrence="3757437392" collectingDate="2007-05-20" collectionCode="IBUNAM" collectorName="O. F. Francke & J. Ponce & M. Villasenor & A. Quijano" country="Mexico" elevation="198" location="El Vado" municipality="Municipio de La Huacana" pageId="24" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Michoacan">
|
||
<taxonomicName id="58E52ED0FFEB2E480BA99D8E91FDFAF3" authority="(Ponce-Saavedra and Sissom, 2004)" authorityName="González-Santillán & Prendini" authorityYear="2013" baseAuthorityName="Ponce-Saavedra and Sissom" baseAuthorityYear="2004" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="cisnerosi" status="comb. nov.">
|
||
<emphasis id="AD918941FFEB2E480BA99D8E97E3FAF2" italics="true" pageId="24" pageNumber="24">Balsateres cisnerosi</emphasis>
|
||
(
|
||
<bibRefCitation id="FB7428A2FFEB2E480AAF9DAC91C5FAF3" author="Ponce-Saavedra, J. & W. D. Sissom" box="[730,1181,1269,1291]" pageId="24" pageNumber="24" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B6A2343AFFEB2E480AEE9C4B967DFAD0" box="[667,805,1298,1320]" pageId="24" pageNumber="24" rank="species">comb. nov.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="AD918941FFEB2E480B409C4B911CFAD0" bold="true" box="[821,1092,1297,1320]" pageId="24" pageNumber="24">
|
||
<collectingCountry id="E7F215C3FFEB2E480B409C4B96F5FAD0" box="[821,941,1298,1320]" name="Mexico" pageId="24" pageNumber="24">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="5D219BB1FFEB2E480BCB9C489167FAD0" box="[958,1087,1297,1320]" country="Mexico" name="Michoacan de Ocampo" pageId="24" pageNumber="24">Michoacán</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="AD918941FFEB2E480C3A9C4A96DFFABD" italics="true" pageId="24" pageNumber="24">
|
||
<collectingMunicipality id="7F3ECF29FFEB2E480C3A9C4A96DFFABD" pageId="24" pageNumber="24">Municipio de La Huacana</collectingMunicipality>
|
||
</emphasis>
|
||
:
|
||
<location id="9A3A0388FFEB2E480BE99C76915EFABD" LSID="urn:lsid:plazi:treatment:174CE445FFE62E480B399C0196A7FA65:9A3A0388FFEB2E480BE99C76915EFABD" box="[924,1030,1327,1349]" country="Mexico" municipality="Municipio de La Huacana" name="El Vado" pageId="24" pageNumber="24" stateProvince="Michoacan">El Vado</location>
|
||
, 18
|
||
<emphasis id="AD918941FFEB2E480C469C769166FABC" bold="true" box="[1075,1086,1327,1348]" pageId="24" pageNumber="24">°</emphasis>
|
||
48.908
|
||
<emphasis id="AD918941FFEB2E480CFC9C7791CAFABD" box="[1161,1170,1326,1349]" italics="true" pageId="24" pageNumber="24">′</emphasis>
|
||
N 101
|
||
<emphasis id="AD918941FFEB2E480AB19C149797FA9A" bold="true" box="[708,719,1357,1378]" pageId="24" pageNumber="24">°</emphasis>
|
||
54.976
|
||
<emphasis id="AD918941FFEB2E480B6F9C12967BFA9A" box="[794,803,1355,1378]" italics="true" pageId="24" pageNumber="24">′</emphasis>
|
||
W,
|
||
<quantity id="581DF8B6FFEB2E480B149C149698FA9A" box="[865,960,1357,1378]" metricMagnitude="2" metricUnit="m" metricValue="1.98" pageId="24" pageNumber="24" unit="m" value="198.0">
|
||
<elevation id="14C8B260FFEB2E480B149C149698FA9A" box="[865,960,1357,1378]" metricMagnitude="2" metricUnit="m" metricValue="1.98" pageId="24" pageNumber="24" unit="m" value="198.0">198 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="EB5B7393FFEB2E480B949C14910AFA9A" box="[993,1106,1357,1378]" pageId="24" pageNumber="24" value="2007-05-20">
|
||
<collectingDate id="FB1F8A7BFFEB2E480B949C14910AFA9A" box="[993,1106,1357,1378]" pageId="24" pageNumber="24" value="2007-05-20">20.v.2007</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="32103085FFEB2E480C069C1497A6FA78" pageId="24" pageNumber="24">O.F. Francke</collectorName>
|
||
,
|
||
<collectorName id="32103085FFEB2E480B679C339626FA87" box="[786,894,1386,1407]" pageId="24" pageNumber="24">J. Ponce</collectorName>
|
||
,
|
||
<collectorName id="32103085FFEB2E480BE79C339165FA78" box="[914,1085,1386,1408]" pageId="24" pageNumber="24">M. Villaseñor</collectorName>
|
||
, and
|
||
<collectorName id="32103085FFEB2E480CF99C3397A5FA65" pageId="24" pageNumber="24">A. Quijano</collectorName>
|
||
,
|
||
<specimenCount id="89E39EDAFFEB2E480B7F9CDE9677FA66" box="[778,815,1412,1438]" pageId="24" pageNumber="24" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="89E39EDAFFEB2E480B4A9CDE963AFA66" box="[831,866,1412,1438]" pageId="24" pageNumber="24" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="F9F4CD96FFEB2E480B069CDE96ABFA65" box="[883,1011,1415,1437]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="24" pageNumber="24" type="University or college">IBUNAM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |