137 lines
17 KiB
XML
137 lines
17 KiB
XML
<document id="10D44FED05EB7187D7A5BB7B31414DE3" ID-CLB-Dataset="28466" ID-DOI="http://dx.doi.org/10.3897/dez.65.27424" ID-GBIF-Dataset="290e099d-cb77-4ad7-975e-32ce4a51926f" ID-Pensoft-Pub="1860-1324-2-177" ID-ZooBank="A047B48DD161424FB8800428DCC5888A" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-177" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M.lophoides (Chaudoir) species complex" checkinTime="1552154842744" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="174B66F1FEBD73DD335A7CF9434D8E64" docLanguage="en" docName="DeutEntomolZeit 65(2): 177-224" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27424" docTitle="Mecyclothorax peryphoides" docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFB33A56FFD86772FFAAFFA9FF882245" masterDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex" masterLastPageNumber="224" masterPageNumber="177" pageNumber="177" updateTime="1701378747773" updateUser="ExternalLinkService">
|
||
<mods:mods id="B00A93F4D14759E5BA4CEA9617F3862F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3465A0B7938B25669C58111996C799FB">
|
||
<mods:title id="53BD5E6C47EF5B46AD979CD7190B4A17">Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E1040692985715FA11878A17D7491EEE" type="personal">
|
||
<mods:role id="BC64D23096120C60AA5DB0D913A35715">
|
||
<mods:roleTerm id="9263CC6ADD6569C5F42FFE30680D404B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2E5ED9DD2A85A15549D9E8FD9736172A">Liebherr, James K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8127CCA5BD18A2CBCD696C5DCB451704">text</mods:typeOfResource>
|
||
<mods:relatedItem id="20A5CC07C56EF77FD626F580AFDC5897" type="host">
|
||
<mods:titleInfo id="F1B66E0B7D08582C3684FDD05EE97BAB">
|
||
<mods:title id="E1BAA09B7A1C3D09E4997D90AA3DDCB9">Deutsche Entomologische Zeitschrift</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1185E6E0441FC30B8FE1FE62E5FF4532">
|
||
<mods:date id="979A7EF32D99026C7763D857FA6599FD">2018</mods:date>
|
||
<mods:detail id="0BF3BB2EFDC89BD263900F792D46094C" type="volume">
|
||
<mods:number id="63B6399A02C9D2B2FF685AF7C6829DE2">65</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3F9EAB4FBB5304C0ECB31A5B6C40EB5A" type="issue">
|
||
<mods:number id="B3EE8CCE6541AF95D76C607BD683BC18">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BD43879D64707ED47CA4C6FF43A346F0" unit="page">
|
||
<mods:start id="099134AEA3B30E72117770BFE99977A9">177</mods:start>
|
||
<mods:end id="791F9E5FC7D62602B6E07C477A3242EC">224</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="BF5F301B4B063309AF822B45B2A0D340">
|
||
<mods:url id="D7D5B8E65D07B1CADCB52EB6D8F29A24">http://dx.doi.org/10.3897/dez.65.27424</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="9792C558B311051AA06ADB07FC6BF577">journal article</mods:classification>
|
||
<mods:identifier id="7FD7DCE14842F714A1B78296C7C926A2" type="DOI">http://dx.doi.org/10.3897/dez.65.27424</mods:identifier>
|
||
<mods:identifier id="1AEC1D4E6DF9BDCF1A22D834B6EECE4A" type="Pensoft-Pub">1860-1324-2-177</mods:identifier>
|
||
<mods:identifier id="985246E3718FE86841F306978B7CFC7B" type="ZooBank">A047B48DD161424FB8800428DCC5888A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="174B66F1FEBD73DD335A7CF9434D8E64" ID-GBIF-Taxon="154365992" LSID="urn:lsid:plazi:treatment:174B66F1FEBD73DD335A7CF9434D8E64" httpUri="http://treatment.plazi.org/id/174B66F1FEBD73DD335A7CF9434D8E64" lastPageNumber="177" pageId="0" pageNumber="177">
|
||
<subSubSection id="D07D7ABE5A4BD497EA780D01A46F7127" pageId="0" pageNumber="177" type="nomenclature">
|
||
<paragraph id="A8F20B3C1D331A1A3CF9396BD8FAB9F6" pageId="0" pageNumber="177">
|
||
<taxonomicName id="D6EE16E92C5AADB3E93F73B6012D9292" ID-CoL="8M7QJ" authority="Blackburn" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax peryphoides" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="peryphoides">Mecyclothorax peryphoides (Blackburn)</taxonomicName>
|
||
Figures 2H, 5B, 9H, 10H, 11D, 12D, 13D, 14
|
||
<normalizedToken id="769B667B59710715937035C45F9657D3" originalValue="E–F">E-F</normalizedToken>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="034D5BFF751412595327DFC59D71CE60" pageId="0" pageNumber="177" type="reference_group">
|
||
<paragraph id="2C8879A432507805A3233430ADF1FBB5" pageId="0" pageNumber="177">
|
||
<taxonomicName id="265E6C44558794038E718F2C7752A506" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax peryphoides" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="peryphoides">Cyclothorax peryphoides</taxonomicName>
|
||
<bibRefCitation id="93FEAF88DEB200075984F7196C471120" author="Blackburn, T" journalOrPublisher=")" pageId="0" pageNumber="177" pagination="1387 - 1506" title="Further notes on Australian Coleoptera, with descriptions of new genera and species. Proceedings of the Linnean Society of New South Wales (2 nd ser." volume="3" year="1889">Blackburn 1889</bibRefCitation>
|
||
: 1392.
|
||
</paragraph>
|
||
<paragraph id="18861B8D7EB09FB7870899EDB68F70F0" pageId="0" pageNumber="177">
|
||
<taxonomicName id="DC65541F2F1E795B33B43A16C728D03F" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax peryphoides" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="peryphoides">Cyclothorax peryphoides</taxonomicName>
|
||
Sloane, 1895: 446.
|
||
</paragraph>
|
||
<paragraph id="FDAB8277A46A272F67CDDD8803784DC6" pageId="0" pageNumber="177">
|
||
<taxonomicName id="74D0DB1C95810714F467921D15BEED1B" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax peryphoides" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="peryphoides">Mecyclothorax peryphoides</taxonomicName>
|
||
Csiki, 1929: 489.
|
||
</paragraph>
|
||
<paragraph id="B903B47188BB33AC634B45B3AF0D8FB4" pageId="0" pageNumber="177">
|
||
<taxonomicName id="B423BD64C5D507A62430FD90A2EC2AD2" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax cordicollis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="cordicollis">Mecyclothorax cordicollis</taxonomicName>
|
||
Jeannel, 1940: 100 (misidentification).
|
||
</paragraph>
|
||
<paragraph id="C5589BFDE50F3B71AC88AC6311165998" pageId="0" pageNumber="177">
|
||
<taxonomicName id="818D5A8F63FFEF090AF7A98327151E9F" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax lophoides" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="lophoides">Mecyclothorax lophoides</taxonomicName>
|
||
Liebherr, 2011a: 292, table 2 (misidentification).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="48445FF071769B0DB243A705376DFE3E" pageId="0" pageNumber="177" type="diagnosis">
|
||
<paragraph id="C5DE09D9065B37CE8D2853D3AE7367FD" pageId="0" pageNumber="177">Diagnosis</paragraph>
|
||
<paragraph id="929496CCB2D0023651844DF9D4A1F634" pageId="0" pageNumber="177">
|
||
(n = 5). Among the species of the
|
||
<taxonomicName id="706D190DE80E7287822ED7AFB0192134" lsidName="M. lophoides" pageId="0" pageNumber="177" rank="species" species="lophoides">M. lophoides</taxonomicName>
|
||
complex characterized by darker bodies and contrastingly pale legs (Figs 4B, 5
|
||
<normalizedToken id="F208140BB0F3EE00B8149A89473E8BA7" originalValue="B–C">B-C</normalizedToken>
|
||
), this species has an ellipsoid elytra with large strial punctures, a pronotum with distinct obtuse hind angles, and concolorous elytral disc and apex, i.e. without an apical flavous marginal band (Fig. 5B). The eyes are broadly convex, MHW/mFW = 1.60-166. The pronotal margins are sinuate anterad the well-defined hind angles, but the posterior margin behind the angle is at most slightly sinuate, and usually convex, smoothly meeting the curved median basal margin (Fig. 2H). Body coloration is dark, with the head, pronotum and elytra piceous. The pronotum is transverse and constricted
|
||
<normalizedToken id="14C8F8D05A93A2AFF0CED071D53919EA" originalValue="basally–MEW">basally-MEW</normalizedToken>
|
||
/EL = 1.20-1.28, MPW/BPW = 1.63
|
||
<normalizedToken id="FA15A1223E6A51E4A23508A4381A8AEB" originalValue="–1.71–">-1.71-</normalizedToken>
|
||
though the basal constriction less than that observed in beetles of
|
||
<taxonomicName id="392B8B8A5D94AF7ADE55942D45234CAC" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
; MEW/BPW = 1.76-1.82. The pronotal median base is covered with distinct punctures, about 20 punctulae each side from midline to the very shallow, poorly defined laterobasal depression (Fig. 2H). The pronotal lateral marginal depression is narrow, but broad enough for sculpticells to line the deepest part, and the margin is beaded. Like
|
||
<taxonomicName id="721DD16838E7703607B7DE82569A940D" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
: 1, the prosternum is flat to depressed medially, the medial area bearing a longitudinal series of 4-7 distinct punctures; 2, the anteapical groove is deep and distinctly punctate laterally, continuous and more shallowly punctate medially; 3, the marginal bead of the procoxal cavity is bordered anteriorly by 5-6 strigose punctulae; and 4, the concave mesepisternal surface is lined with 13 deep punctures arrayed in 3-4 irregular rows. The elytral striae are punctate on the disc, though the punctures are far enough apart that the cuticle is coplanar with the intervals between the punctures. Like
|
||
<taxonomicName id="A71331A5EBB4D6D3DDEFB759D9B111FF" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
, the punctures of striae 1-6 are progressively more isolated laterally on the elytra, but the sutural stria is more shallowly and irregularly punctured mesad the posterior dorsal elytral seta in this species (Fig. 12
|
||
<normalizedToken id="069C6ADC7CF7FF235192BD252EAB2E65" originalValue="C–D">C-D</normalizedToken>
|
||
). Cuticular microsculpture is well developed in this species, with: 1, frons covered with indistinct transverse lines, vertex with more well-developed sculpticells, transforming from transverse just behind posterior supraorbital setae to nearly isodiametric near pronotum; 2, pronotal disc with indistinct transverse lines visible outside areas of direct reflection, irregular surface of pronotal base covered with irregular transverse mesh; 3, elytral disc with transverse lines visible outside areas of direct reflection, apex covered with well-developed elongate transverse mesh. Standardized body length 4.3-5.0 mm. Setal formula ++/++/+2++.
|
||
</paragraph>
|
||
<paragraph id="5700D92E36ACFA435E184A1DA6D0918B" pageId="0" pageNumber="177">
|
||
Male genitalia (n = 16). As stated under
|
||
<taxonomicName id="767479C74BB7C499ED498D6ECA72681A" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
, the aedeagal median lobe and flagellar complex of that species and
|
||
<taxonomicName id="AF885D1F88A13FFF04FCFB43A7BF6EF0" lsidName="M. peryphoides" pageId="0" pageNumber="177" rank="species" species="peryphoides">M. peryphoides</taxonomicName>
|
||
show no differences (Fig. 14
|
||
<normalizedToken id="042168538D6DD1A7E2FF1222B946005F" originalValue="D–E">D-E</normalizedToken>
|
||
). However the right paramere of this species bears significantly more setae (9-17) along its ventral margin.
|
||
</paragraph>
|
||
<paragraph id="45FF8E3EFE5145D4A1FCB3616C9B8B6C" pageId="0" pageNumber="177">
|
||
Female reproductive tract (n = 3). Bursa copulatrix short, slightly longer than broad (Fig. 9H); helminthoid sclerite broad basally, with narrow mediodistal projection; spermathecal duct elongate, sclerotized enough to hold coiled configuration, length ~2
|
||
<normalizedToken id="DD44449EB40CB28923B98C6688D821F8" originalValue="×">x</normalizedToken>
|
||
length of spermathecal reservoir; basal gonocoxite with 3-6 apical setae, 2-3 large and the balance smaller, plus a similarly sized seta at the apicomedial angle (Fig. 10H); medial surface of basal gonocoxite with larger setae apically; apical gonocoxite subtriangular, narrowly rounded apically; lateral ensiform setae small, narrow; apical nematiform setae in apical sensory furrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B4FE09B862EB7E407FF25DD9C429E8B" pageId="0" pageNumber="177" type="type information">
|
||
<paragraph id="D3C866A7E96B7E28BCC62B5CDC78BB85" pageId="0" pageNumber="177">Type information.</paragraph>
|
||
<paragraph id="0EA2848AC9809290F0EF4734D5610DD8" pageId="0" pageNumber="177">
|
||
Holotype male (BMNH): platen mounted with
|
||
<normalizedToken id="06C9E372A94EB723254DC3AE74A5C265" originalValue="“1614”">"1614"</normalizedToken>
|
||
in red ink and
|
||
<normalizedToken id="A9C0B8A1A153050BC01C2E4326A79908" originalValue="“T”">"T"</normalizedToken>
|
||
in black ink on the obverse, with basal marginal black and red lines, the red crossed by an arrowhead // Type (round red-margined label) // Blackburn coll. 1910-236. //
|
||
<taxonomicName id="D68AE4AC7D2A037A3D17437E101FEDB8" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Cyclothorax</taxonomicName>
|
||
periphoides [sic.], Blackb. Blackburn states the type locality as "Woodville, near Adelaide; a single specimen (
|
||
<bibRefCitation id="08CECA89E8E3E9E9BA0EEC93E35A6AD5" author="Blackburn, T" journalOrPublisher=")" pageId="0" pageNumber="177" pagination="1387 - 1506" title="Further notes on Australian Coleoptera, with descriptions of new genera and species. Proceedings of the Linnean Society of New South Wales (2 nd ser." volume="3" year="1889">Blackburn 1889</bibRefCitation>
|
||
: 1393)."
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7D73AD489C45ACF8C6B9ABA7310362C3" pageId="0" pageNumber="177" type="distribution">
|
||
<paragraph id="BF078109C4B169620B01052BC7731757" pageId="0" pageNumber="177">Distribution and habitat.</paragraph>
|
||
<paragraph id="F91B2097611E4DA1FBD48B9B42922238" pageId="0" pageNumber="177">
|
||
This species is distributed (Fig. 11D) to the south of its sister species,
|
||
<taxonomicName id="6CA7B3308E2E49EC95D500805D245EEE" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
, in South Australia. The type locality of Woodville is near recent collections made at Belair Recreation Park, 10 miles S of Adelaide (FMNH), Blackwood, Sturt Gorge Reserve (ZMUC), and Hale Conservation Park SE Williamstown (FMNH). To the east the species is recorded from the following localities: A.C.T.: Black Mountain (CUIC, 8; EMEC, 1); NSW: Belmore (AMS, 1); Bogan R. (AMS, 1); "
|
||
<taxonomicName id="09A55D14825C465F9AE94080647C0F83" class="Insecta" family="Carabidae" genus="Calosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calosoma" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Calosoma</taxonomicName>
|
||
" via Gundaroo (ANIC, 4); Federal Hwy. (ANIC, 1); Jerrawa (AMCS, 2); Mt. Wilson (CUIC, 1; FMNH, 4); Narrabeen (AMS, 1); Queanbeyan (ANIC, 1); Tuglo Wldlf. Ref. 48 km N Singleton (FMNH, 1); Weddin Mtns. N. P. (FMNH, 1); VIC: Lake Eildon N. P., Sebastopol Ck. (ZMUC, 1). Collections have been made via litter sifting along streams or in
|
||
<taxonomicName id="FF12E5D59538A3702A5DAD8B083DF95E" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="0" pageNumber="177" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
|
||
(
|
||
<taxonomicName id="60F58392056A3991891F1CDCD8B863F4" family="Myrtaceae" lsidName="Moriomorphina" pageId="0" pageNumber="177" rank="family">Myrtaceae</taxonomicName>
|
||
) or mixed deciduous forest, or in rainforest on Mt. Wilson. As for
|
||
<taxonomicName id="0777123A3BD1CEDBC44432D1CABCB9B4" lsidName="M. eyrensis" pageId="0" pageNumber="177" rank="species" species="eyrensis">M. eyrensis</taxonomicName>
|
||
, even though this species occupies predominantly riparian situations, it appears to be vestigially winged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |