treatments-xml/data/15/F1/26/15F126E6266E58B4AFD6A48F510D1C5B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.98.89413" ID-Pensoft-Pub="1860-0743-2-327" ID-Pensoft-UUID="1C115BD7E10C5139A21E322F336B5A5C" ID-ZooBank="78BD2D176B3446D6816385C87B9652BF" ModsDocID="1860-0743-98-2-327" checkinTime="1662649616165" checkinUser="pensoft" docAuthor="Depra, Gabriel de Carvalho, Aguilera, Gaston, Faustino-Fuster, Dario R., Katz, Axel M. &amp; Azevedo-Santos, Valter M." docDate="2022" docId="15F126E6266E58B4AFD6A48F510D1C5B" docLanguage="en" docName="ZoosystEvol 98(2): 327-343" docOrigin="Zoosystematics and Evolution 98 (2)" docPubDate="2022-09-08" docSource="http://dx.doi.org/10.3897/zse.98.89413" docTitle="Heptapterus Bleeker 1858" docType="treatment" docVersion="1" id="1C115BD7E10C5139A21E322F336B5A5C" lastPageNumber="327" masterDocId="1C115BD7E10C5139A21E322F336B5A5C" masterDocTitle="Redefinition of Heptapterus (Heptapteridae) and description of Heptapterus carmelitanorum, a new species from the upper Parana River basin in Brazil" masterLastPageNumber="343" masterPageNumber="327" pageNumber="327" updateTime="1662649616165" updateUser="pensoft">
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<mods:title>Redefinition of Heptapterus (Heptapteridae) and description of Heptapterus carmelitanorum, a new species from the upper Parana River basin in Brazil</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Depra, Gabriel de Carvalho</mods:namePart>
<mods:affiliation>Universidade Estadual de Maringa. Av. Colombo, 5790, 87020 - 900, Maringa, Parana, Brazil</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Aguilera, Gaston</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo (CONICET) - Fundacion Miguel Lillo. Miguel Lillo 251, San Miguel de Tucuman (CP 4000), Tucuman, Argentina</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Faustino-Fuster, Dario R.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-3495</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Porto Alegre, Brazil &amp; Departamento de Ictiologia, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru &amp; Escuela de Ingenieria y Gestion Ambiental, Facultad de Ingenieria y Gestion, Universidad Nacional Autonoma de Huanta, Huanta, Peru</mods:affiliation>
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<mods:namePart>Katz, Axel M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2933-7163</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Azevedo-Santos, Valter M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8986-6406</mods:nameIdentifier>
<mods:affiliation>Faculdade Eduvale de Avare, Avare, Sao Paulo, Brazil &amp; Nucleo de Ecologia Aquatica e Pesca da Amazonia, Grupo de Ecologia Aquatica, Universidade Federal do Para, Belem, Para, Brazil &amp; Programa de Pos-Graduacao em Biodiversidade, Ecologia e Conservacao, Universidade Federal do Tocantins, Porto Nacional, Tocantins, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">valter.ecologia@gmail.com</mods:nameIdentifier>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-09-08</mods:number>
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<mods:number>98</mods:number>
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<mods:start>327</mods:start>
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<treatment LSID="urn:lsid:plazi:treatment:15F126E6266E58B4AFD6A48F510D1C5B" httpUri="http://treatment.plazi.org/id/15F126E6266E58B4AFD6A48F510D1C5B" lastPageNumber="327" pageId="0" pageNumber="327">
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<paragraph pageId="0" pageNumber="327">
Genus 
<taxonomicName LSID="15F126E6-266E-58B4-AFD6-A48F510D1C5B" authority="Bleeker, 1858" authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Heptapterus Bleeker, 1858</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="327" type="type species">
<paragraph pageId="0" pageNumber="327">Type species.</paragraph>
<paragraph pageId="0" pageNumber="327">
<taxonomicName authorityName="Valenciennes" authorityYear="1835" class="Actinopterygii" family="Pimelodidae" genus="Pimelodus" kingdom="Animalia" lsidName="Pimelodus mustelinus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="mustelinus">
<emphasis italics="true" pageId="0" pageNumber="327">Pimelodus mustelinus</emphasis>
</taxonomicName>
Valenciennes, 1835.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="327" type="diagnosis">
<paragraph pageId="0" pageNumber="327">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="327">
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
</taxonomicName>
differs from all other 
<taxonomicName lsidName="" pageId="0" pageNumber="327" rank="tribe" tribe="Heptapterini">Heptapterini</taxonomicName>
except 
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" class="Actinopterygii" family="Heptapteridae" genus="Acentronichthys" kingdom="Animalia" lsidName="Acentronichthys" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Acentronichthys</emphasis>
</taxonomicName>
Eigenmann &amp; Eigenmann, 1889, 
<taxonomicName authorityName="C.H.Eigenmann &amp; R.S.Eigenmann" authorityYear="1889" class="Actinopterygii" family="Heptapteridae" genus="Nemuroglanis" kingdom="Animalia" lsidName="Nemuroglanis" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Nemuroglanis</emphasis>
</taxonomicName>
Eigenmann &amp; Eigenmann, 1889, 
<taxonomicName class="Pisces" family="Heptapteridae" genus="Chasmocranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmocranus bleekeri" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="bleekeri">
<emphasis italics="true" pageId="0" pageNumber="327">Chasmocranus bleekeri</emphasis>
</taxonomicName>
, ' 
<taxonomicName authorityName="C.H.Eigenmann" authorityYear="1912" class="Pisces" family="Heptapteridae" genus="Chasmocranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmocranus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Chasmocranus</taxonomicName>
' 
<taxonomicName authorityName="Gomes &amp; Schubart" authorityYear="1958" class="Pisces" family="Heptapteridae" genus="Chasmocranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmocranus brachynema" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="brachynema">Chasmocranus brachynema</taxonomicName>
Gomes &amp; Schubart, 1958, ' 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Heptapterus</taxonomicName>
' 
<taxonomicName authorityName="Ihering" authorityYear="1907" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus multiradiatus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="multiradiatus">Heptapterus multiradiatus</taxonomicName>
, ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Haseman" authorityYear="1911" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus stewarti" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="stewarti">Heptapterus stewarti</taxonomicName>
, and ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Buckup" authorityYear="1988" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus sympterygium" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="sympterygium">Heptapterus sympterygium</taxonomicName>
by the presence of an adipose fin extensively fused with the caudal fin (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic representation of the different degrees of proximity and connection between the adipose and caudal fins in Heptapterini. a. Adipose and caudal fins widely separate, as in Imparfinis piperatus, for instance; b. Adipose fin reaching the caudal fin, but not connecting to it (i. e., connective tissue in which dorsal procurrent caudal-fin rays are imbedded is not contiguous with connective tissue forming the adipose fin), as in Chasmocranus longior, for instance; c. Adipose fin connecting with caudal fin (i. e., connective tissue in which dorsal procurrent caudal-fin rays are imbedded is clearly contiguous with connective tissue forming the adipose fin), as in Heptapterus." figureDoi="10.3897/zse.98.89413.figure2" httpUri="https://binary.pensoft.net/fig/739959" pageId="0" pageNumber="327">2</figureCitation>
). 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" class="Actinopterygii" family="Heptapteridae" genus="Acentronichthys" kingdom="Animalia" lsidName="Acentronichthys" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Acentronichthys</emphasis>
</taxonomicName>
by having a non-bifurcate caudal fin (i.e., caudal fin not divided in two lobes; vs. bifurcate, with distinct dorsal and ventral lobes), and from 
<taxonomicName authorityName="C.H.Eigenmann &amp; R.S.Eigenmann" authorityYear="1889" class="Actinopterygii" family="Heptapteridae" genus="Nemuroglanis" kingdom="Animalia" lsidName="Nemuroglanis" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Nemuroglanis</emphasis>
</taxonomicName>
by having dark bars and stripes on back of trunk (vs. absence of dark bars and stripes on back of trunk) and 5-6 pairs of pleural ribs (vs. 8-9 ribs). It is also distinguished from ' 
<taxonomicName genus="C." lsidName="C." pageId="0" pageNumber="327" rank="genus">C.</taxonomicName>
' 
<taxonomicName authorityName="Gomes &amp; Schubart" authorityYear="1958" class="Pisces" family="Heptapteridae" genus="Chasmocranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmocranus brachynema" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="brachynema">Chasmocranus brachynema</taxonomicName>
by having an elongate body, with a head length of 16.1-24.9% (vs. 25.5-27.3%); the posterior extension of mouth rim much shorter, with rictus barely reaching vertical line through posterior nostril (vs. posterior extension of mouth rim much longer, with rictus reaching vertical line between posterior nostril and eye); and the premaxillary tooth plate with no posterolateral extension, or with a small one (vs. with a very long posterolateral extension). It is further distinguished from 
<taxonomicName lsidName="C. bleekeri" pageId="0" pageNumber="327" rank="species" species="bleekeri">
<emphasis italics="true" pageId="0" pageNumber="327">C. bleekeri</emphasis>
</taxonomicName>
by having the pelvic-fin insertion posterior to vertical through insertion of dorsal fin (vs. anterior) and anal-fin insertion posterior to vertical through adipose-fin origin (vs. anterior). Additionally, 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
</taxonomicName>
differs from ' 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">Heptapterus</taxonomicName>
' 
<taxonomicName authorityName="Ihering" authorityYear="1907" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus multiradiatus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="multiradiatus">Heptapterus multiradiatus</taxonomicName>
and ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Haseman" authorityYear="1911" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus stewarti" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="stewarti">Heptapterus stewarti</taxonomicName>
by having fewer anal-fin rays (10-23 in 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
</taxonomicName>
vs. 38-46 and 33-36 in ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Ihering" authorityYear="1907" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus multiradiatus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="multiradiatus">Heptapterus multiradiatus</taxonomicName>
and ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Haseman" authorityYear="1911" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus stewarti" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="stewarti">Heptapterus stewarti</taxonomicName>
, respectively). It is further distinguished from ' 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="H." kingdom="Plantae" lsidName="H." order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="genus">H.</taxonomicName>
' 
<taxonomicName authorityName="Buckup" authorityYear="1988" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus sympterygium" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="sympterygium">Heptapterus sympterygium</taxonomicName>
by having the anal and caudal fins separated (vs. anal fin confluent with the caudal fin), and supraorbital pore 6 (s6) fused or closer to each other (vs. separate and closer to the eye than to each other).
</paragraph>
<caption doi="10.3897/zse.98.89413.figure2" httpUri="https://binary.pensoft.net/fig/739959" pageId="0" pageNumber="327" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="327">
<emphasis bold="true" pageId="0" pageNumber="327">Figure 2.</emphasis>
Schematic representation of the different degrees of proximity and connection between the adipose and caudal fins in 
<taxonomicName lsidName="" pageId="0" pageNumber="327" rank="tribe" tribe="Heptapterini">Heptapterini</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="327">a.</emphasis>
Adipose and caudal fins widely separate, as in 
<taxonomicName authorityName="Eigenmann &amp; Norris" authorityYear="1900" class="Pisces" family="Heptapteridae" genus="Imparfinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imparfinis piperatus" order="Siluriformes" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="piperatus">
<emphasis italics="true" pageId="0" pageNumber="327">Imparfinis piperatus</emphasis>
</taxonomicName>
, for instance; 
<emphasis bold="true" pageId="0" pageNumber="327">b.</emphasis>
Adipose fin reaching the caudal fin, but not connecting to it (i.e., connective tissue in which dorsal procurrent caudal-fin rays are imbedded is not contiguous with connective tissue forming the adipose fin), as in 
<taxonomicName authorityName="Eigenmann" authorityYear="1912" class="Pisces" family="Heptapteridae" genus="Chasmocranus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmocranus longior" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="longior">
<emphasis italics="true" pageId="0" pageNumber="327">Chasmocranus longior</emphasis>
</taxonomicName>
, for instance; 
<emphasis bold="true" pageId="0" pageNumber="327">c.</emphasis>
Adipose fin connecting with caudal fin (i.e., connective tissue in which dorsal procurrent caudal-fin rays are imbedded is clearly contiguous with connective tissue forming the adipose fin), as in 
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="327" type="species included">
<paragraph pageId="0" pageNumber="327">Species included.</paragraph>
<paragraph pageId="0" pageNumber="327">
<taxonomicName class="Pisces" family="Heptapteridae" genus="Heptapterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptapterus borodini" order="Nematognathi" pageId="0" pageNumber="327" phylum="Chordata" rank="species" species="borodini">
<emphasis italics="true" pageId="0" pageNumber="327">Heptapterus borodini</emphasis>
</taxonomicName>
(Mees &amp; Cala, 1989), 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carmelitanorum" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carmelitanorum">
<emphasis italics="true" pageId="0" pageNumber="327">H. carmelitanorum</emphasis>
</taxonomicName>
, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. carnatus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="carnatus">
<emphasis italics="true" pageId="0" pageNumber="327">H. carnatus</emphasis>
</taxonomicName>
Faustino-Fuster, Bockmann &amp; Malabarba, 2019, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. exilis" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="exilis">
<emphasis italics="true" pageId="0" pageNumber="327">H. exilis</emphasis>
</taxonomicName>
Faustino-Fuster, Bockmann &amp; Malabarba, 2019, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. hollandi" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="hollandi">
<emphasis italics="true" pageId="0" pageNumber="327">H. hollandi</emphasis>
</taxonomicName>
(Haseman, 1911), 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mandimbusu" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mandimbusu">
<emphasis italics="true" pageId="0" pageNumber="327">H. mandimbusu</emphasis>
</taxonomicName>
Aguilera, Benitez, 
<normalizedToken originalValue="Terán">Teran</normalizedToken>
, Alonso &amp; Mirande, 2017, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mbya" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mbya">
<emphasis italics="true" pageId="0" pageNumber="327">H. mbya</emphasis>
</taxonomicName>
Azpelicueta, Aguilera &amp; Mirande, 2011, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. mustelinus" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="mustelinus">
<emphasis italics="true" pageId="0" pageNumber="327">H. mustelinus</emphasis>
</taxonomicName>
(Valenciennes, 1835), 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. ornaticeps" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="ornaticeps">
<emphasis italics="true" pageId="0" pageNumber="327">H. ornaticeps</emphasis>
</taxonomicName>
Ahl, 1936, and 
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="H. qenqo" order="Fabales" pageId="0" pageNumber="327" phylum="Tracheophyta" rank="species" species="qenqo">
<emphasis italics="true" pageId="0" pageNumber="327">H. qenqo</emphasis>
</taxonomicName>
Aguilera, Mirande &amp; Azpelicueta, 2011.
</paragraph>
</subSubSection>
</treatment>
</document>