252 lines
26 KiB
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252 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.953.53977" ID-GBIF-Dataset="77c65687-4a7f-4b63-911d-1d096aaaad76" ID-PMC="PMC7398950" ID-Pensoft-Pub="1313-2970-953-1" ID-Pensoft-UUID="C7001D577310538DBE2A4736802E1BF4" ID-PubMed="32821193" ID-ZooBank="BA81E87988A2495A92FB98D1F2909BA7" ModsDocID="1313-2970-953-1" checkinTime="1595913774766" checkinUser="pensoft" docAuthor="Wesener, Thomas" docDate="2020" docId="15E9BAD2035050778C6E120CB922AB5C" docLanguage="en" docName="ZooKeys 953: 1-29" docOrigin="ZooKeys 953" docSource="http://dx.doi.org/10.3897/zookeys.953.53977" docTitle="Riotintobolus antafoky Wesener 2020, sp. nov." docType="treatment" docUuid="9F3E2163-ACA9-4A1F-B6D0-D24CD3A692D8" docUuidSource="ZooBank" docVersion="4" id="C7001D577310538DBE2A4736802E1BF4" lastPageNumber="1" masterDocId="C7001D577310538DBE2A4736802E1BF4" masterDocTitle="Ecotone shifts in southern Madagascar: first barcoding data and six new species of the endemic millipede genus Riotintobolus (Spirobolida, Pachybolidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="1" updateTime="1668169162946" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Ecotone shifts in southern Madagascar: first barcoding data and six new species of the endemic millipede genus Riotintobolus (Spirobolida, Pachybolidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>953</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>29</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.953.53977</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.953.53977</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-953-1</mods:identifier>
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<mods:identifier type="ZooBank">BA81E87988A2495A92FB98D1F2909BA7</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C7001D577310538DBE2A4736802E1BF4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="166016737" LSID="urn:lsid:zoobank.org:act:9F3E2163-ACA9-4A1F-B6D0-D24CD3A692D8" httpUri="http://treatment.plazi.org/id/15E9BAD2035050778C6E120CB922AB5C" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="15E9BAD2-0350-5077-8C6E-120CB922AB5C" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Riotintobolus antafoky" order="Spirobolida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antafoky" status="sp. nov.">Riotintobolus antafoky</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">Figure 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: ♂,
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<emphasis bold="true" pageId="0" pageNumber="1">CASENT 9032794</emphasis>
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, MGF007, Madagascar, Province Toliara, Antafoky, 80 m, spiny thicket,
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<geoCoordinate degrees="23" direction="south" minutes="29" orientation="latitude" precision="15" seconds="16" value="-23.487778">23°29'16"S</geoCoordinate>
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,
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<geoCoordinate degrees="44" direction="east" minutes="4" orientation="longitude" precision="15" seconds="39" value="44.0775">44°4'39"E</geoCoordinate>
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, coll. Frontier project, millipede dig (3 m x 3 m), 14.xi.2001.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: 1 ♂, 1 ♀,
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<emphasis bold="true" pageId="0" pageNumber="1">CASENT 9032794</emphasis>
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, same data as holotype
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Antafoky, after the type locality, spiny forest of Antafoky (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the newly described Riotintobolus species, as well as of the still undescribed species. Vegetation marks modified from Moat and Smith 2007." figureDoi="10.3897/zookeys.953.53977.figure2" httpUri="https://binary.pensoft.net/fig/436163" pageId="0" pageNumber="1">2</figureCitation>
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). Noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName lsidName="R. antafoky" pageId="0" pageNumber="1" rank="species" species="antafoky">
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<emphasis italics="true" pageId="0" pageNumber="1">R. antafoky</emphasis>
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</taxonomicName>
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sp. nov. shares the flag-like membranous tip of the posterior gonopod with
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<taxonomicName lsidName="R. mandenensis" pageId="0" pageNumber="1" rank="species" species="mandenensis">
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<emphasis italics="true" pageId="0" pageNumber="1">R. mandenensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="R. minutus" pageId="0" pageNumber="1" rank="species" species="minutus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. minutus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="R. aridus" pageId="0" pageNumber="1" rank="species" species="aridus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. aridus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="R. anomalus" pageId="0" pageNumber="1" rank="species" species="anomalus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. anomalus</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="R. makayi" pageId="0" pageNumber="1" rank="species" species="makayi">
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<emphasis italics="true" pageId="0" pageNumber="1">R. makayi</emphasis>
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</taxonomicName>
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sp. nov.
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<taxonomicName lsidName="R. antafoky" pageId="0" pageNumber="1" rank="species" species="antafoky">
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<emphasis italics="true" pageId="0" pageNumber="1">R. antafoky</emphasis>
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</taxonomicName>
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sp. nov. shares the absence of tarsal pads only with
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<taxonomicName lsidName="R. bovinus" pageId="0" pageNumber="1" rank="species" species="bovinus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. bovinus</emphasis>
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</taxonomicName>
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sp. nov., and the relatively simple tip of the posterior gonopod only with
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<taxonomicName lsidName="R. anomalus" pageId="0" pageNumber="1" rank="species" species="anomalus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. anomalus</emphasis>
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</taxonomicName>
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.
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<taxonomicName lsidName="R. antafoky" pageId="0" pageNumber="1" rank="species" species="antafoky">
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<emphasis italics="true" pageId="0" pageNumber="1">R. antafoky</emphasis>
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</taxonomicName>
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sp. nov. differs from the sympatric
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<taxonomicName lsidName="R. anomalus" pageId="0" pageNumber="1" rank="species" species="anomalus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. anomalus</emphasis>
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</taxonomicName>
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in details of the posterior gonopod, the absence of a dorsal red stripe, the much longer antenna (protruding back to body ring 5), and the much smaller size (
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<taxonomicName lsidName="R. anomalus" pageId="0" pageNumber="1" rank="species" species="anomalus">
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<emphasis italics="true" pageId="0" pageNumber="1">R. anomalus</emphasis>
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</taxonomicName>
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males 45 mm long, 4.3 mm wide,
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<taxonomicName lsidName="R. antafoky" pageId="0" pageNumber="1" rank="species" species="antafoky">
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<emphasis italics="true" pageId="0" pageNumber="1">R. antafoky</emphasis>
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</taxonomicName>
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sp. nov. ca. 33 mm long, 3.2 mm wide).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
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(holotype): 51+0 segments, ca. 33 mm long (fragmented) and 3.2 mm wide.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
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: segments grey, with a dark grey posterior margin, ozopore highlighted by a black spot. Head, antennae and legs dark grey.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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: each eye with 34 ommatidia in six rows. Incisura lateralis open. Labrum with standard three irregular teeth and a single row of 10-12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennae long, protruding back to segment 5. Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 and 6 latero-apically with sensilla basiconica.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gnathochilarium</emphasis>
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: lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Endochilarium not dissected.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mandible</emphasis>
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: Stipes without projection, well rounded. Gnathal lobe not dissected.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Collum</emphasis>
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: smooth, laterally not protruding as far as ring 2.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Body rings</emphasis>
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: ozopores starting at segment 6, marked by a black spot. Located slightly before, but touching suture between meso- and metazonite. Rings with smooth, but irregular coriaceous surface, ventrally on metazona with transverse ridges.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Telson</emphasis>
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: paraprocts without lips, abundant micropunctation especially towards edges. Epiproct well-rounded, covering, but not reaching above paraproct. Hypoproct inconspicuous.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
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: leg 1 with a large cylindrical coxa, twice as long as other podomeres. Tarsus with three pairs of ventral spines and an apical spine beyond claw. Leg 2 with an elongated coxa and a strongly swollen prefemur. Tarsus with two pairs of ventral spines and a short apical spine. Midbody legs with a rectangular coxa, as long as other podomeres. Each podomere ventrally with a single or a pair of apical setae, tarsus without a tarsal pad, a single apical and two pairs of ventral spines. Length of midbody legs ca. 0.9 times body diameter in males.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female</emphasis>
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: not investigated.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male sexual characters</emphasis>
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: tarsal pads absent. Coxae 3-7 without coxal processes, but coxae 6 and 7 flattened, rectangular.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Anterior gonopod</emphasis>
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sternite massive (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9A</figureCitation>
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), elongated into a wide, well-rounded triangular lobe (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9A</figureCitation>
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). Sternite in anterior view well-visible, without discernible apodemes, protruding almost as high as coxal processes. Coxite with sharp triangular mesal process (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9A, B</figureCitation>
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), upper rim bending forward in anterior view (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9A</figureCitation>
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). Telopodite with strong process arising mesally (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9B</figureCitation>
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), process apically curved with a short triangular projection (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9A</figureCitation>
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), tip well-rounded, protruding above lateral margin of telopodite (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9B</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
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<taxonomicName authorityName="Wesener" authorityYear="2020" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Riotintobolus antafoky" order="Spirobolida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antafoky">
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<emphasis italics="true" pageId="0" pageNumber="1">Riotintobolus antafoky</emphasis>
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</taxonomicName>
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sp. nov., holotype (CASENT9032794)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior gonopod, anterior view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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anterior gonopod, posterior view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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telopodite of left posterior gonopod, anterior view
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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left posterior gonopod, posterior view.
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<emphasis bold="true" pageId="0" pageNumber="1">Abbreviations</emphasis>
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: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous
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<normalizedToken originalValue="‘flag’">'flag'</normalizedToken>
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. Not to scale.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Posterior gonopods</emphasis>
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consisting of two parts, separated by an articulation: a long coxite and a slightly shorter telopodite, efferent duct discharging apically (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C, D</figureCitation>
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). Process of coxite and telopodite standing in same axis (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9D</figureCitation>
|
||
). Pair of posterior gonopods located parallel to each other, connected by a small, sclerotised and visible sternite (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9D</figureCitation>
|
||
). Basal part of coxite wide, mesally with a triangular sclerite located on lower level than remaining part (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C</figureCitation>
|
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). Coxite elongated. Efferent duct running at mesal margin of coxite (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C, D</figureCitation>
|
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). Telopodite as wide as but slightly shorter than coxite, standing in same axis (Fig.
|
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9D</figureCitation>
|
||
), apically with a membranous
|
||
<normalizedToken originalValue="‘flag’">'flag'</normalizedToken>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C, D</figureCitation>
|
||
). Laterally with sclerotised projection bending laterally and completely surrounded by membranous flag. (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C, D</figureCitation>
|
||
). Efferent duct ending at base of process (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Riotintobolus antafoky sp. nov., holotype (CASENT 9032794) A anterior gonopod, anterior view B anterior gonopod, posterior view C telopodite of left posterior gonopod, anterior view D left posterior gonopod, posterior view. Abbreviations: Cx = coxite; St = sternite; T = telopodite; x = lateral swollen process; y = opening of efferenct duct; z = apical membranous ' flag'. Not to scale." figureDoi="10.3897/zookeys.953.53977.figure9" httpUri="https://binary.pensoft.net/fig/436170" pageId="0" pageNumber="1">9C, D</figureCitation>
|
||
). Apex laterally with weakly developed finger-shaped process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
this species lives in direct sympatry with the much larger
|
||
<taxonomicName lsidName="R. anomalus" pageId="0" pageNumber="1" rank="species" species="anomalus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. anomalus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |