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<document id="77E2BD61993C2873CEF447FC686D0E43" ID-DOI="10.11646/zootaxa.3752.1.11" ID-GBIF-Dataset="aa45f110-cd42-4c88-84c8-b53031a5914f" ID-ISSN="1175-5326" ID-Zenodo-Dep="219785" ID-ZooBank="F51BDBEB-E635-46A8-B103-ED785C986F0B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460711933348" checkinUser="plazi" docAuthor="Last, Peter R., Ho, Hsuan-Ching &amp; Chen, Rou-Rong" docDate="2013" docId="045587EE5D72FF82FF44FCC2FD685ABA" docLanguage="en" docName="zt03752p198.pdf" docOrigin="Zootaxa 3752 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhynchobatus immaculatus Last, Ho &amp; Chen, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="196" masterDocId="F86CFF965D70FF89FFD3FF80FF915C5A" masterDocTitle="A new species of wedgefish, Rhynchobatus immaculatus (Chondrichthyes, Rhynchobatidae), from Taiwan" masterLastPageNumber="198" masterPageNumber="185" pageNumber="187" updateTime="1698358133627" updateUser="plazi">
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<mods:title id="E9E49989E06881090F8278086EC4A2B3">A new species of wedgefish, Rhynchobatus immaculatus (Chondrichthyes, Rhynchobatidae), from Taiwan</mods:title>
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<mods:namePart id="84C820772193D4D99AD8A5BD449BC24B">Last, Peter R.</mods:namePart>
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<mods:namePart id="609CE3401D90EC6A8C0A30DD2A4B63E8">Ho, Hsuan-Ching</mods:namePart>
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<mods:namePart id="67F259BF2A8BD050FB5D46920A1E68F5">Chen, Rou-Rong</mods:namePart>
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<mods:title id="70B1A0E1BE0F9682B7F6FE1A14424270">Zootaxa</mods:title>
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<mods:date id="2FB114DFF970E91E43FF76415FA461CC">2013</mods:date>
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<treatment id="045587EE5D72FF82FF44FCC2FD685ABA" ID-DOI="http://doi.org/10.5281/zenodo.5623785" ID-GBIF-Taxon="119585179" ID-Zenodo-Dep="5623785" LSID="urn:lsid:plazi:treatment:045587EE5D72FF82FF44FCC2FD685ABA" httpUri="http://treatment.plazi.org/id/045587EE5D72FF82FF44FCC2FD685ABA" lastPageId="11" lastPageNumber="196" pageId="2" pageNumber="187">
<subSubSection id="C4E665735D72FF8BFF44FCC2FE1E5F24" pageId="2" pageNumber="187" type="nomenclature">
<paragraph id="8C4336F85D72FF8BFF44FCC2FDDC5F06" blockId="2.[151,589,834,894]" box="[151,589,834,860]" pageId="2" pageNumber="187">
<heading id="D70B81945D72FF8BFF44FCC2FDDC5F06" bold="true" box="[151,589,834,860]" fontSize="11" level="1" pageId="2" pageNumber="187" reason="1">
<emphasis id="BE88EAEA5D72FF8BFF44FCC2FDDC5F06" bold="true" box="[151,589,834,860]" pageId="2" pageNumber="187">
<taxonomicName id="4BFC4D7B5D72FF8BFF44FCC2FE795F06" box="[151,488,834,860]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="187" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D72FF8BFF44FCC2FE795F06" bold="true" box="[151,488,834,860]" italics="true" pageId="2" pageNumber="187">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D72FF8BFE23FCC2FDDC5F06" box="[496,589,834,860]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8C4336F85D72FF8BFF44FCE5FE1E5F24" blockId="2.[151,589,834,894]" box="[151,399,869,894]" pageId="2" pageNumber="187">
(
<figureCitation id="14C72A7D5D72FF8BFF4DFCE5FEB95F24" box="[158,296,869,894]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStartId-0="3.[151,244,1363,1384]" captionStartId-1="4.[151,244,1256,1277]" captionStartId-2="5.[151,250,1899,1921]" captionStartId-3="6.[151,244,1191,1212]" captionStartId-4="6.[151,244,1798,1819]" captionStartId-5="7.[151,244,1931,1952]" captionStartId-6="10.[151,244,1954,1975]" captionTargetBox-0="[153,1432,398,1343]" captionTargetBox-1="[226,1360,395,1241]" captionTargetBox-2="[767,1436,475,1874]" captionTargetBox-3="[415,1171,154,1169]" captionTargetBox-4="[491,1095,1293,1774]" captionTargetBox-5="[183,1404,683,1906]" captionTargetBox-6="[339,1246,940,1926]" captionTargetId-0="figure@3.[149,1434,850,1347]" captionTargetId-1="figure@4.[226,1360,391,1242]" captionTargetId-2="figure@5.[767,1372,964,1460]" captionTargetId-3="figure@6.[415,1171,149,1169]" captionTargetId-4="figure@6.[491,1096,1293,1774]" captionTargetId-5="figure@7.[183,1404,668,1908]" captionTargetId-6="figure@10.[339,1246,940,1926]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="10" captionText-0="FIGURE 2. Rhynchobatus immaculatus sp. nov. immature male holotype (NMMB P 16274, 730 mm TL, recently preserved). A. dorsal view; B. ventral view." captionText-1="FIGURE 3. Rhynchobatus immaculatus sp. nov. View of central dorsal disc of immature male holotype (NMMB P 16274, 730 mm TL, preserved) showing markings and the spine rows." captionText-2="FIGURE 4. Rhynchobatus immaculatus sp. nov. fin shape and development. Immature male holotype (NMMB P 16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMB P 15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin." captionText-3="FIGURE 5. Rhynchobatus immaculatus sp. nov. Snout and orbito-spiracular region of immature male non-type (NMMB P 15684, 992 mm TL, preserved)." captionText-4="FIGURE 6. Rhynchobatus immaculatus sp. nov. View of oronasal region and snout immature male holotype (NMMB P 16274, 730 mm TL, recently preserved)." captionText-5="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." captionText-6="FIGURE 8. Distribution of Rhynchobatus immaculatus sp. nov. Yellow star—holotype; yellow dots—market sites for other material. Map extracted from Google Earth." httpUri-0="https://zenodo.org/record/219787/files/figure.png" httpUri-1="https://zenodo.org/record/219788/files/figure.png" httpUri-2="https://zenodo.org/record/219789/files/figure.png" httpUri-3="https://zenodo.org/record/219790/files/figure.png" httpUri-4="https://zenodo.org/record/219791/files/figure.png" httpUri-5="https://zenodo.org/record/219792/files/figure.png" httpUri-6="https://zenodo.org/record/219793/files/figure.png" pageId="2" pageNumber="187">Figures 28</figureCitation>
,
<tableCitation id="C17E03435D72FF8BFEE0FCE5FE155F24" box="[307,388,869,894]" captionStart="TABLE 1" captionStartId="8.[151,232,151,172]" captionTargetBox="[165,1436,268,2004]" captionTargetPageId="8" captionText="TABLE 1. Morphometric data for the immature male holotype (NMMB P 16274) and paratypes (CSIRO H 7418 01, NMMB-P 16275 &amp; P 16135) of Rhynchobatus immaculatus sp. nov. Measurements are expressed as percentages of total length. Data for paratypes are expressed as minima, maxima and means." httpUri="http://table.plazi.org/id/D88366705D78FF81FF44FF17FDBA5CB5" pageId="2" pageNumber="187" tableUuid="D88366705D78FF81FF44FF17FDBA5CB5">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D72FF8BFF44FC2DFBA25820" pageId="2" pageNumber="187" type="materials_examined">
<paragraph id="8C4336F85D72FF8BFF44FC2DFDE05FB0" blockId="2.[151,1437,941,2010]" pageId="2" pageNumber="187">
<emphasis id="BE88EAEA5D72FF8BFF44FC2DFE9A5F9F" bold="true" box="[151,267,941,965]" pageId="2" pageNumber="187">
<typeStatus id="5347885A5D72FF8BFF44FC2DFE975F9F" box="[151,262,941,965]" pageId="2" pageNumber="187" type="holotype">Holotype</typeStatus>
.
</emphasis>
NMMBP16274, immature male
<quantity id="4B049B1D5D72FF8BFD42FC2EFD615F9F" box="[657,752,942,966]" metricMagnitude="-1" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="187" unit="mm" value="730.0">730 mm</quantity>
TL, Keelung fish market,
<collectingCountry id="F4EB76685D72FF8BFBF2FC2DFBE75F9C" box="[1057,1142,941,966]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
,
<date id="F84210385D72FF8BFB51FC2DFA8F5F9C" box="[1154,1310,941,966]" pageId="2" pageNumber="187" value="2012-03-16">16 Mar. 2012</date>
, purchased from market by P. Last, field no. HO179.
</paragraph>
<paragraph id="8C4336F85D72FF8BFF14FC75FBA25820" blockId="2.[151,1437,941,2010]" pageId="2" pageNumber="187">
<emphasis id="BE88EAEA5D72FF8BFF14FC75FED75857" bold="true" box="[199,326,1013,1037]" pageId="2" pageNumber="187">
<typeStatus id="5347885A5D72FF8BFF14FC75FED05857" box="[199,321,1013,1037]" pageId="2" pageNumber="187" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9AFAFD715D72FF8BFE82FC76FE755854" box="[337,484,1013,1038]" pageId="2" pageNumber="187" type="generic">3 specimens</specimenCount>
: CSIRO H 741801, immature male
<quantity id="4B049B1D5D72FF8BFC4DFC76FB915857" box="[926,1024,1014,1038]" metricMagnitude="-1" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="187" unit="mm" value="890.0">890 mm</quantity>
TL, Wu-chi fish market, Taichung County,
<collectingCountry id="F4EB76685D72FF8BFF24FB99FED85868" box="[247,329,1049,1074]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
Strait, western
<collectingCountry id="F4EB76685D72FF8BFE2EFB99FDC35868" box="[509,594,1049,1074]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
, ca.
<geoCoordinate id="E9C8503F5D72FF8BFD54FB9AFD7B5868" box="[647,746,1050,1074]" direction="north" orientation="latitude" pageId="2" pageNumber="187" precision="925" value="24.3">24°18N</geoCoordinate>
,
<geoCoordinate id="E9C8503F5D72FF8BFD24FB9AFCF55868" box="[759,868,1050,1074]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="925" value="120.48333">120°29E</geoCoordinate>
,
<date id="F84210385D72FF8BFCA1FB9AFB9F5868" box="[882,1038,1050,1074]" pageId="2" pageNumber="187" value="2010-05-21">21 May 2010</date>
, purchased from market by H. Ho; NMMBP16135, female
<quantity id="4B049B1D5D72FF8BFE60FBBEFD81580F" box="[435,528,1086,1110]" metricMagnitude="-1" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="187" unit="mm" value="780.0">780 mm</quantity>
TL, Keelung fish market,
<collectingCountry id="F4EB76685D72FF8BFCE5FBBDFC1A580C" box="[822,907,1085,1110]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
,
<date id="F84210385D72FF8BFC45FBBEFB8B580C" box="[918,1050,1086,1110]" pageId="2" pageNumber="187" value="2012-03-09">9 Mar 2012</date>
, purchased from market by H. Ho; NMMBP16275 [HO180], female
<quantity id="4B049B1D5D72FF8BFDFDFBE2FD1A5823" box="[558,651,1122,1146]" metricMagnitude="-1" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="187" unit="mm" value="740.0">740 mm</quantity>
TL, collected together with
<typeStatus id="5347885A5D72FF8BFC19FBE1FBA15820" box="[970,1072,1121,1146]" pageId="2" pageNumber="187" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D72FF8BFF14FB05FB0E59C3" pageId="2" pageNumber="187" type="description">
<paragraph id="8C4336F85D72FF8BFF14FB05FB0E59C3" blockId="2.[151,1437,941,2010]" pageId="2" pageNumber="187">
<emphasis id="BE88EAEA5D72FF8BFF14FB05FED658C7" bold="true" box="[199,327,1157,1181]" pageId="2" pageNumber="187">Non-types.</emphasis>
<specimenCount id="9AFAFD715D72FF8BFE87FB05FE7658C4" box="[340,487,1157,1182]" pageId="2" pageNumber="187" type="generic">6 specimens</specimenCount>
: ASIZP 63513, immature male
<quantity id="4B049B1D5D72FF8BFCB4FB06FC5B58C7" box="[871,970,1158,1182]" metricMagnitude="-1" metricUnit="m" metricValue="4.08" pageId="2" pageNumber="187" unit="mm" value="408.0">408 mm</quantity>
TL, Ao-di, Taipei county, northeastern
<collectingCountry id="F4EB76685D72FF8BFF44FB29FF7D5898" box="[151,236,1193,1218]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
, ca.
<geoCoordinate id="E9C8503F5D72FF8BFEFDFB29FE005898" box="[302,401,1193,1218]" direction="north" orientation="latitude" pageId="2" pageNumber="187" precision="925" value="25.05">25°03N</geoCoordinate>
,
<geoCoordinate id="E9C8503F5D72FF8BFE76FB29FD825898" box="[421,531,1193,1218]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="925" value="121.933334">121°56E</geoCoordinate>
,
<date id="F84210385D72FF8BFDC4FB29FD245898" box="[535,693,1193,1218]" pageId="2" pageNumber="187" value="2001-07-17">17 Jul. 2001</date>
, coll. Z.-H. Wu; ASIZP 66972, female
<quantity id="4B049B1D5D72FF8BFB7BFB29FA9F589B" box="[1192,1294,1193,1218]" metricMagnitude="-1" metricUnit="m" metricValue="8.35" pageId="2" pageNumber="187" unit="mm" value="835.0">835 mm</quantity>
TL, Da-xi, northeastern
<collectingCountry id="F4EB76685D72FF8BFEFBFB4DFEEC58BC" box="[296,381,1229,1254]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
,
<date id="F84210385D72FF8BFE5BFB4DFD9858BC" box="[392,521,1229,1254]" pageId="2" pageNumber="187" value="2007-02-05">5 Feb 2007</date>
; NMMBP16225, immature male
<quantity id="4B049B1D5D72FF8BFC5DFB4EFC7A58BF" box="[910,1003,1230,1254]" metricMagnitude="-1" metricUnit="m" metricValue="7.44" pageId="2" pageNumber="187" unit="mm" value="744.0">744 mm</quantity>
TL, Da-xi, Yilan County, northeastern
<collectingCountry id="F4EB76685D72FF8BFF44FB71FF7D5950" box="[151,236,1265,1290]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
, ca.
<geoCoordinate id="E9C8503F5D72FF8BFEFBFB71FE1A5950" box="[296,395,1265,1290]" direction="north" orientation="latitude" pageId="2" pageNumber="187" precision="925" value="24.933332">24°56N</geoCoordinate>
,
<geoCoordinate id="E9C8503F5D72FF8BFE48FB71FD9B5950" box="[411,522,1265,1290]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="925" value="121.9">121°54E</geoCoordinate>
,
<date id="F84210385D72FF8BFDDDFB71FD3D5950" box="[526,684,1265,1290]" pageId="2" pageNumber="187" value="2012-03-13">13 Mar 2012</date>
, purchased from market by P. Last; NMMBP16224, immature female
<quantity id="4B049B1D5D72FF8BFF39FA95FED65977" box="[234,327,1301,1326]" metricMagnitude="-1" metricUnit="m" metricValue="9.15" pageId="2" pageNumber="187" unit="mm" value="915.0">915 mm</quantity>
TL, Wu-chi fish market, Taichung,
<collectingCountry id="F4EB76685D72FF8BFD08FA95FCBC5974" box="[731,813,1301,1326]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
Strait, western
<collectingCountry id="F4EB76685D72FF8BFC0EFA95FBA35974" box="[989,1074,1301,1326]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
, ca.
<geoCoordinate id="E9C8503F5D72FF8BFBB7FA96FB595974" box="[1124,1224,1302,1326]" direction="north" orientation="latitude" pageId="2" pageNumber="187" precision="925" value="24.3">24°18N</geoCoordinate>
,
<geoCoordinate id="E9C8503F5D72FF8BFB00FA96FAD05974" box="[1235,1345,1302,1326]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="925" value="120.48333">120°29E</geoCoordinate>
,
<date id="F84210385D72FF8BFA9EFA96FF435908" pageId="2" pageNumber="187" value="2012-04-10">10 Apr 2012</date>
, purchased from the market by H. Ho; NMMBP15684, immature male
<quantity id="4B049B1D5D72FF8BFBD6FABAFBF2590B" box="[1029,1123,1338,1362]" metricMagnitude="-1" metricUnit="m" metricValue="9.92" pageId="2" pageNumber="187" unit="mm" value="992.0">992 mm</quantity>
TL, collected together with CSIRO H 741801; NMMBP15661, immature male
<quantity id="4B049B1D5D72FF8BFCD2FADDFCF1592F" box="[769,864,1373,1398]" metricMagnitude="-1" metricUnit="m" metricValue="5.91" pageId="2" pageNumber="187" unit="mm" value="591.0">591 mm</quantity>
TL, Wu-chi fish market, Taichung,
<collectingCountry id="F4EB76685D72FF8BFB2DFADDFAC1592C" box="[1278,1360,1373,1398]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
Strait, western
<collectingCountry id="F4EB76685D72FF8BFF26FA01FEDB59C0" box="[245,330,1409,1434]" name="Taiwan" pageId="2" pageNumber="187">Taiwan</collectingCountry>
, ca.
<geoCoordinate id="E9C8503F5D72FF8BFEAFFA02FE4E59C0" box="[380,479,1410,1434]" direction="north" orientation="latitude" pageId="2" pageNumber="187" precision="925" value="24.3">24°18N</geoCoordinate>
,
<geoCoordinate id="E9C8503F5D72FF8BFE39FA02FDC959C0" box="[490,600,1410,1434]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="925" value="120.48333">120°29E</geoCoordinate>
,
<date id="F84210385D72FF8BFDB7FA02FD6159C0" box="[612,752,1410,1434]" pageId="2" pageNumber="187" value="2011-09-11">11 Sep 2011</date>
, purchased from the market by H. Ho.
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D72FF8BFF14FA24FA055B7C" pageId="2" pageNumber="187" type="diagnosis">
<paragraph id="8C4336F85D72FF8BFF14FA24FA055B7C" blockId="2.[151,1437,941,2010]" pageId="2" pageNumber="187">
<emphasis id="BE88EAEA5D72FF8BFF14FA24FED059E7" bold="true" box="[199,321,1444,1469]" pageId="2" pageNumber="187">Diagnosis.</emphasis>
A medium-sized species of the genus
<taxonomicName id="4BFC4D7B5D72FF8BFD20FA25FC0359E7" box="[755,914,1445,1469]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="BE88EAEA5D72FF8BFD20FA25FC0359E7" box="[755,914,1445,1469]" italics="true" pageId="2" pageNumber="187">Rhynchobatus</emphasis>
</taxonomicName>
with the following combination of characters: a broad, wedge-shaped snout; preoral snout 1921% TL; eye small, length
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preorbital snout; interorbital space
<quantity id="4B049B1D5D72FF8BFF0DFA6DFEC05A5C" box="[222,337,1517,1542]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.366" metricValueMin="6.604" pageId="2" pageNumber="187" unit="in" value="2.75" valueMax="2.9" valueMin="2.6">2.62.9 in</quantity>
preorbital snout; tooth rows about 48; spines absent from snout; supraorbital spines small, well differentiated, extending from preorbit to beyond end of spiracle; predorsal spines relatively well developed; two disjunct rows of 69 small scapular spines on each side of disc; predorsal space 4849% of total length; origin of first dorsal fin well behind origin of pelvic-fin bases; dark greenish brown dorsally with a few white spots scattered on pectoral disc; black pectoral marking absent; prominent row of white spots commencing just forward of a single white spot (above pelvic-fin origin) and coalescing posteriorly to form a white mid-lateral line on tail; orbital membrane white with dark bar over eye; no alternating light and dark markings on interorbital space; total pectoralfin radials 6568; vertebrae with 2830 monospondylous precaudal centra, 124131 precaudal free centra, 3842 diplospondylous caudal (free) centra, 165170 total free centra, 179184 total centra (including synarcual centra).
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D72FF82FF14F8B1FC925D47" lastPageId="11" lastPageNumber="196" pageId="2" pageNumber="187" type="description">
<paragraph id="8C4336F85D72FF8AFF14F8B1FB075D3F" blockId="2.[151,1437,941,2010]" lastBlockId="3.[151,1436,151,357]" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187">
<emphasis id="BE88EAEA5D72FF8BFF14F8B1FEC85B10" bold="true" box="[199,345,1841,1866]" pageId="2" pageNumber="187">Description.</emphasis>
Body relatively robust; snout in front of eyes bluntly angular to obtusely wedge-shaped, angle about 50° in
<typeStatus id="5347885A5D72FF8BFEF5F8D5FE195B34" box="[294,392,1877,1902]" pageId="2" pageNumber="187" type="holotype">holotype</typeStatus>
(5253° in
<typeStatus id="5347885A5D72FF8BFDDFF8D6FDED5B34" box="[524,636,1878,1902]" pageId="2" pageNumber="187" type="paratype">paratypes</typeStatus>
). Lateral margin of snout slightly convex beside orbit, otherwise almost straight. Preorbital length
<quantity id="4B049B1D5D72FF8BFE13F8FAFD905BC8" box="[448,513,1913,1938]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" pageId="2" pageNumber="187" unit="in" value="3.3">3.3 in</quantity>
<typeStatus id="5347885A5D72FF8BFDDAF8F9FDFA5BC8" box="[521,619,1913,1938]" pageId="2" pageNumber="187" type="holotype">holotype</typeStatus>
(
<quantity id="4B049B1D5D72FF8BFDAFF8FAFD615BC8" box="[636,752,1913,1938]" metricMagnitude="-2" metricUnit="m" metricValue="8.508999999999999" metricValueMax="8.636" metricValueMin="8.382" pageId="2" pageNumber="187" unit="in" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.33.4 in</quantity>
<typeStatus id="5347885A5D72FF8BFD2BF8FAFCF45BC8" box="[760,869,1914,1938]" pageId="2" pageNumber="187" type="paratype">paratypes</typeStatus>
) times interorbital width. Preoral length 3.4 (3.0 3.3) times mouth width. Disc width across pectoral-fin apices 75% (7577%) of disc length from snout tip to pectoral-fin free rear tips. Head strongly depressed, trowel-shaped, disc thickness 1.5 (1.61.7) times in interorbital space; ventral head length 3.2 (3.23.4) times in total length; surface between eyes and spiracles almost flat. Precloacal length 85% (8688%) of length of tail from anterior vent to caudal-fin tip. Tail moderately depressed; in cross section, arched dorsally, almost flat ventrally, angular ventrolaterally at lateral keels, tapering evenly from pelvic-fin insertions. Width of tail at first dorsal-fin insertions of
<typeStatus id="5347885A5D73FF8AFC71FE84FB955D47" box="[930,1028,260,285]" pageId="3" pageNumber="188" type="holotype">holotype</typeStatus>
1.8 (1.61.9) times interspiracular distance. Lateral keels extended forward as a thickened angular edge along precaudal tail, reaching forward to below posterior half of first-dorsal fin; strongly differentiated on anterior part of caudal fin.
</paragraph>
<caption id="D88366705D73FF8AFF44FAD3FE3A59DC" httpUri="https://zenodo.org/record/219787/files/figure.png" pageId="3" pageNumber="188" targetBox="[153,1432,398,1343]" targetPageId="3">
<paragraph id="8C4336F85D73FF8AFF44FAD3FE3A59DC" blockId="3.[151,1437,1363,1414]" pageId="3" pageNumber="188">
<emphasis id="BE88EAEA5D73FF8AFF44FAD3FE9D5932" bold="true" box="[151,268,1363,1384]" pageId="3" pageNumber="188">FIGURE 2.</emphasis>
<taxonomicName id="4BFC4D7B5D73FF8AFEC1FAD3FD8F5932" box="[274,542,1363,1384]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="188" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D73FF8AFEC1FAD3FD8F5932" box="[274,542,1363,1384]" italics="true" pageId="3" pageNumber="188">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D73FF8AFDF4FAD3FDE45932" box="[551,629,1363,1384]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
immature male holotype (NMMBP16274, 730 mm TL, recently preserved). A. dorsal view; B. ventral view.
</paragraph>
</caption>
<paragraph id="8C4336F85D73FF8AFF14FA32FDC75A25" blockId="3.[151,1436,1458,2023]" pageId="3" pageNumber="188">Horizontal eye (eyeball) diameter about 63% (6467%) of interspiracular width, distance from anterior margin of orbit to posterior margin of spiracle slightly smaller than interspiracular width; greatest dimension of spiracles 58% (4750%) of horizontal eye diameter; distance between spiracle and eye less than a fifth horizontal eye diameter, membrane of orbit almost continuous with spiracular opening. Spiracle dorsolateral, anterior margin with a partly concealed thickened valve, posterior margin with two short anteriorly directed spiracular folds; outer fold slightly taller and larger than inner fold.</paragraph>
<paragraph id="8C4336F85D73FF8AFF14F90AFC815BC5" blockId="3.[151,1436,1458,2023]" pageId="3" pageNumber="188">Nostrils diagonal, forming about a 45° angle with body axis, incurrent aperture directed more laterally. Nasal cavity fully exposed, without dividing flaps; aperture straight anterolaterally, recurved posteromedially. Anterior nasal flap narrow, low, anteromedial on nasal aperture, inserted near midlength of nasal aperture; anterior process short, bilobed, its base length about twice as long as its width. Posterolateral nasal flap low, narrow and elongated, weakly lobate; originating just behind anterior lateral edge of incurrent aperture, extending posteriorly to about midlength of nasal aperture. Posterior nasal flap low, short based; joined to undersurface of posterolateral flap near anterior third of its length, junction concealed beneath posterolateral nasal flap; inserted near posterior quarter of nostril. Nostril width 1.3 (1.2) times in internarial width.</paragraph>
<paragraph id="8C4336F85D73FF8DFF14F82AFEB45D3F" blockId="3.[151,1436,1458,2023]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="189" pageId="3" pageNumber="188">
Mouth opening somewhat arcuate, weakly undulating to nearly straight laterally; strong medial depression on upper jaw corresponding to a very prominent anterior protrusion at symphysis of lower jaw; much weaker corresponding depressions and convexities laterally. Labial folds and furrows short, but well developed at corners of mouth; shallow pockets, circumoral grooves, and low folds and depressions, surround jaws laterally to labial folds; grooves most prominent on lower jaw. Teeth in quincunx, about 48 rows in both upper and lower jaws of
<typeStatus id="5347885A5D74FF8DFF44FE85FF665D47" box="[151,247,261,285]" pageId="4" pageNumber="189" type="paratype">paratype</typeStatus>
CSIRO H 741801. First four gill openings subequal in length, the fifth slightly shorter. Third gill opening 2.6 (2.32.6) in internarial width, 3.3 (2.83.1) times in nostril length, 1.2 (1.21.7) times length of fifth gill opening.
</paragraph>
<caption id="D88366705D74FF8DFF44FB68FD0D5940" httpUri="https://zenodo.org/record/219788/files/figure.png" pageId="4" pageNumber="189" targetBox="[226,1360,395,1241]" targetPageId="4">
<paragraph id="8C4336F85D74FF8DFF44FB68FD0D5940" blockId="4.[151,1435,1256,1306]" pageId="4" pageNumber="189">
<emphasis id="BE88EAEA5D74FF8DFF44FB68FE9D58A7" bold="true" box="[151,268,1256,1277]" pageId="4" pageNumber="189">FIGURE 3.</emphasis>
<taxonomicName id="4BFC4D7B5D74FF8DFEC1FB68FD8D58A7" box="[274,540,1256,1277]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="189" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D74FF8DFEC1FB68FD8D58A7" box="[274,540,1256,1277]" italics="true" pageId="4" pageNumber="189">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D74FF8DFDF0FB68FDFE58A7" box="[547,623,1256,1277]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
View of central dorsal disc of immature male holotype (NMMBP16274, 730 mm TL, preserved) showing markings and the spine rows.
</paragraph>
</caption>
<paragraph id="8C4336F85D74FF8DFF14FAC7FAD75AB6" blockId="4.[151,1436,1351,2024]" pageId="4" pageNumber="189">
Dermal denticles minute, covering all of body surface, varying in shape and size over body (
<figureCitation id="14C72A7D5D74FF8DFAF5FAC7FA1C5905" box="[1318,1421,1351,1376]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
). Denticles with slender pedicels and elevated crowns; crowns rhomboidal, subcircular to broadly ovoid, margins irregularly rounded to crenulate anteriorly; crowns weakly unicuspidate, rounded or tricuspidate posteriorly, usually with prominent medial and lateral ridges. On snout, broadly ovoid, textured with 35 pronounced longitudinal ridges; median ridge best developed; posterior margin weakly tricuspidate; scales in two main sizes, weakly imbricated with some gaps (
<figureCitation id="14C72A7D5D74FF8DFDE8FA7BFD0E5A4E" box="[571,671,1531,1556]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
a). On interorbit, somewhat similar to snout (
<figureCitation id="14C72A7D5D74FF8DFB7DFA7BFA835A4E" box="[1198,1298,1531,1556]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
b); those on orbital membrane slightly smaller than those on interorbit; an indistinct patch of slightly enlarged (often more elongate) denticles present in front of eyes. On mid-dorsal surface, more regular in size, dense, often weakly imbricate; narrow patches of skin exposed where scales are missing; posterior margins irregular to rounded (
<figureCitation id="14C72A7D5D74FF8DFA82F9E7FF365AFE" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
c). On outer pectoral fin, much smaller, similar to those of snout (
<figureCitation id="14C72A7D5D74FF8DFC74F90BFB9B5AFE" box="[935,1034,1675,1700]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
d). On flank below first dorsal fin, similar to outer pectoral fin but relatively much broader (
<figureCitation id="14C72A7D5D74FF8DFCFBF92FFC1A5A9D" box="[808,907,1711,1736]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
e). On ventral surface, subrhomboidal, usually acuspid, strongly imbricate; slightly larger than on dorsal flank but mostly smaller than on disc (
<figureCitation id="14C72A7D5D74FF8DFB1FF953FABC5AB6" box="[1228,1325,1747,1772]" captionStart="FIGURE 7" captionStartId="7.[151,244,1931,1952]" captionTargetBox="[183,1404,683,1906]" captionTargetId="figure@7.[183,1404,668,1908]" captionTargetPageId="7" captionText="FIGURE 7. Rhynchobatus immaculatus sp. nov. SEM images (x 100) showing squamation on the left side of a immature male nontype (NMMB P 15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1." httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="4" pageNumber="189">Figure 7</figureCitation>
f).
</paragraph>
<paragraph id="8C4336F85D74FF8CFF14F977FC365DF6" blockId="4.[151,1436,1351,2024]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="190" pageId="4" pageNumber="189">
Small, variable-sized spines present on dorsal surface of body and tail; present in series on orbital margin, along dorsal midline of disc and tail, and in scapular region; those of nuchal and mid-scapular regions largest; rostral spines absent. Spines along midline of disc and tail narrow and long based or globular and weakly oblique; distal surfaces of globular spines smooth, with corrugated bases; keel-like spines with corrugated bases and sometimes lateral edges. Orbit of
<typeStatus id="5347885A5D74FF8DFDCDF808FDEF5BFA" box="[542,638,1928,1952]" pageId="4" pageNumber="189" type="paratype">paratype</typeStatus>
CSIRO H 741801 with a single, continuous series of ca. 20, small, variable-sized spines; series extending along inner margin of orbit from anterior mid-eye to beyond posterior margin of spiracle; similar in form to those along midline; row partly interrupted above mid spiracle in some
<typeStatus id="5347885A5D74FF8DFAB2F850FA0D5BB2" box="[1377,1436,2000,2024]" pageId="4" pageNumber="189">types</typeStatus>
(partially subdivided into orbital and spiracular groups of spines). Predorsal series with ca. 27 spines of varying size in CSIRO H 741801, well developed, spines partly embedded in a low, cutaneous ridge; extending from anterior nuchal region to over pelvic-fin origin; spine bases often within a shallow groove; closely and more or less evenly spaced anteriorly, more widely spaced posteriorly. Interdorsal series barely emergent; absent and not forming a ridge behind second dorsal fin. Scapular spines of CSIRO H 741801in two short, disjunct, linear series of on each side of disc; commencing just behind origin of predorsal series, terminating at level of pectoral-fin apex; anterior series with ca 89 spines, its length subequal to eye diameter; posterior series with 69 spines, short, slightly longer than anterior series; no obvious lateral patches present.
</paragraph>
<paragraph id="8C4336F85D75FF8FFF7FFD87FA0D5BBC" blockId="5.[172,208,519,562]" lastBlockId="6.[151,1436,1889,2022]" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190">
<emphasis id="BE88EAEA5D75FF8CFF7FFD87FF415E68" bold="true" box="[172,208,519,562]" pageId="5" pageNumber="190">A</emphasis>
Dorsal fins similar in shape in young, raked, shark-like, with convex anterior margins (shallowly concave at base), bluntly pointed apices, deeply concave posterior margins, sharply acute free rear tips, and straight inner margins. Inner margin of first dorsal fin about twice (1.92.0 times) in its base length. First dorsal fin larger than second; origin just forward of mid-base of pelvic fins; free rear tip slightly behind free rear tips of pelvic fin.
</paragraph>
<caption id="D88366705D75FF8CFF44F8EBFDE75BBD" httpUri="https://zenodo.org/record/219789/files/figure.png" pageId="5" pageNumber="190" targetBox="[767,1436,475,1874]" targetPageId="5">
<paragraph id="8C4336F85D75FF8CFF44F8EBFDE75BBD" blockId="5.[151,1436,1899,2023]" pageId="5" pageNumber="190">
<emphasis id="BE88EAEA5D75FF8CFF44F8EBFE855BDA" bold="true" box="[151,276,1899,1921]" pageId="5" pageNumber="190">FIGURE 4.</emphasis>
<taxonomicName id="4BFC4D7B5D75FF8CFEC8F8EBFDA25BDB" box="[283,563,1899,1921]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="190" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D75FF8CFEC8F8EBFDA25BDB" box="[283,563,1899,1921]" italics="true" pageId="5" pageNumber="190">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D75FF8CFDE8F8ECFD1A5BDB" box="[571,651,1900,1921]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
fin shape and development. Immature male holotype (NMMBP16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMBP15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin.
</paragraph>
</caption>
<caption id="D88366705D76FF8FFF44FB27FE135883" httpUri="https://zenodo.org/record/219790/files/figure.png" pageId="6" pageNumber="191" targetBox="[415,1171,154,1169]" targetPageId="6">
<paragraph id="8C4336F85D76FF8FFF44FB27FE135883" blockId="6.[151,1436,1191,1241]" pageId="6" pageNumber="191">
<emphasis id="BE88EAEA5D76FF8FFF44FB27FE9D58E6" bold="true" box="[151,268,1191,1212]" pageId="6" pageNumber="191">FIGURE 5.</emphasis>
<taxonomicName id="4BFC4D7B5D76FF8FFEC1FB27FD8F58E6" box="[274,542,1191,1212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="191" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D76FF8FFEC1FB27FD8F58E6" box="[274,542,1191,1212]" italics="true" pageId="6" pageNumber="191">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D76FF8FFDF5FB27FDE258E6" box="[550,627,1191,1212]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
Snout and orbito-spiracular region of immature male non-type (NMMBP15684, 992 mm TL, preserved).
</paragraph>
</caption>
<caption id="D88366705D76FF8FFF44F886FE3A5B63" httpUri="https://zenodo.org/record/219791/files/figure.png" pageId="6" pageNumber="191" targetBox="[491,1095,1293,1774]" targetPageId="6">
<paragraph id="8C4336F85D76FF8FFF44F886FE3A5B63" blockId="6.[151,1436,1798,1849]" pageId="6" pageNumber="191">
<emphasis id="BE88EAEA5D76FF8FFF44F886FE9D5B41" bold="true" box="[151,268,1798,1819]" pageId="6" pageNumber="191">FIGURE 6.</emphasis>
<taxonomicName id="4BFC4D7B5D76FF8FFEC1F886FD8A5B41" box="[274,539,1798,1819]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="191" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D76FF8FFEC1F886FD8A5B41" box="[274,539,1798,1819]" italics="true" pageId="6" pageNumber="191">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D76FF8FFDF0F886FDFE5B41" box="[547,623,1798,1819]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
View of oronasal region and snout immature male holotype (NMMBP16274, 730 mm TL, recently preserved).
</paragraph>
</caption>
<paragraph id="8C4336F85D77FF8EFF44FF17FD585EDF" blockId="7.[151,1436,151,645]" pageId="7" pageNumber="192">Interdorsal space 2.8 (2.32.7) times length of first dorsal base, 3.5 (2.93.3) of length of second dorsal-fin base. Caudal fin rather short, strongly forked; dorsal caudal margin 6.5 (5.96.5) in total length, subequal to interdorsal space. Dorsal caudal margin strongly convex, weakly concave near its origin; tip bluntly pointed. Preventral caudal margin strongly convex; ventral lobe well developed, tip strongly angular. Lower postventral caudal margin short, weakly concave, 2.9 (2.23.2) in length of upper margin. Upper postventral margin weakly concave. Caudal axis elevated slightly, forming a narrow angle to main body axis. Pectoral fin originating at about level of spiracle, with initially concave then convex anterior margins; apices broadly pointed, posterior margins moderately convex; free rear tips narrowly rounded, extending 101% (8589%) of distance between pectoral and pelvic-fin bases (pectoral pelvic space); inner margins straight to weakly concave. Pelvic fins small, with weakly convex to straight anterior margins, broadly rounded apices, weakly concave posterior margins, elongate and very narrowly rounded free rear tips, and concave inner margins; inner margin very long, 1.5 (1.31.5) times length of pelvic base; fin base 1.0 (0.9) in pectoralpelvic space; height of pelvic fin about 2.1 (2.02.3) in their length. Distance between pelvic-fin insertions much longer than pelvic-fin base length. Vent with well-developed folds laterally; well separated from pelvic-fin inner margins. No data on adult clasper.</paragraph>
<caption id="D88366705D77FF8EFF44F80BFCA95B81" httpUri="https://zenodo.org/record/219792/files/figure.png" pageId="7" pageNumber="192" targetBox="[183,1404,683,1906]" targetPageId="7">
<paragraph id="8C4336F85D77FF8EFF44F80BFCA95B81" blockId="7.[151,1436,1931,2011]" pageId="7" pageNumber="192">
<emphasis id="BE88EAEA5D77FF8EFF44F80BFE9D5BFA" bold="true" box="[151,268,1931,1952]" pageId="7" pageNumber="192">FIGURE 7.</emphasis>
<taxonomicName id="4BFC4D7B5D77FF8EFEC1F80BFD8A5BFA" box="[274,539,1931,1952]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="192" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D77FF8EFEC1F80BFD8A5BFA" box="[274,539,1931,1952]" italics="true" pageId="7" pageNumber="192">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D77FF8EFDF0F80BFDFE5BFA" box="[547,623,1931,1952]" pageId="7" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
SEM images (x100) showing squamation on the left side of a immature male nontype (NMMBP15684, 992 mm TL, preserved). A. Snout; B. Supraorbit; C. Mid-disc; D. Pectoral fin; E. Flank beneath first dorsal fin; F. Ventral anterior tail. Letters correspond to those in the Figure 1.
</paragraph>
</caption>
<caption id="D88366705D78FF81FF44FF17FDBA5CB5" ID-Table-UUID="D88366705D78FF81FF44FF17FDBA5CB5" httpUri="http://table.plazi.org/id/D88366705D78FF81FF44FF17FDBA5CB5" pageId="8" pageNumber="193" targetBox="[165,1436,268,2004]" targetIsTable="true" targetPageId="8">
<paragraph id="8C4336F85D78FF81FF44FF17FDBA5CB5" blockId="8.[151,1436,151,239]" pageId="8" pageNumber="193">
<emphasis id="BE88EAEA5D78FF81FF44FF17FE915CF6" bold="true" box="[151,256,151,172]" pageId="8" pageNumber="193">TABLE 1.</emphasis>
Morphometric data for the immature male holotype (NMMBP16274) and paratypes (CSIRO H 741801, NMMB-P16275 &amp; P16135) of
<taxonomicName id="4BFC4D7B5D78FF81FEF1FF37FDBA5C96" box="[290,555,183,204]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="193" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D78FF81FEF1FF37FDBA5C96" box="[290,555,183,204]" italics="true" pageId="8" pageNumber="193">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D78FF81FDE2FF37FDEE5C96" box="[561,639,183,204]" pageId="8" pageNumber="193" rank="species">
<emphasis id="BE88EAEA5D78FF81FDE2FF37FDEE5C96" bold="true" box="[561,639,183,204]" pageId="8" pageNumber="193">sp. nov.</emphasis>
</taxonomicNameLabel>
Measurements are expressed as percentages of total length. Data for paratypes are expressed as minima, maxima and means.
</paragraph>
</caption>
<paragraph id="8C4336F85D78FF81FC84FE8CFA0D5B8E" pageId="8" pageNumber="193">
<table id="FEFCC4585D780076FF76FE8CFA0D5B8E" box="[165,1436,268,2004]" gridcols="3" gridrows="45" pageId="8" pageNumber="193">
<tr id="32CC34BA5D780076FF76FE8CFA0D5D7B" box="[165,1436,268,289]" gridrow="0" pageId="8" pageNumber="193" rowspan-0="1">
<th id="711D5DC65D780076FC84FE8CFC215D7B" box="[855,944,268,289]" gridcol="1" gridrow="0" pageId="8" pageNumber="193">Holotype</th>
<th id="711D5DC65D780076FBF5FE8CFA0D5D7B" box="[1062,1436,268,289]" gridcol="2" gridrow="0" pageId="8" pageNumber="193">Paratypes (n=3)</th>
</tr>
<tr id="32CC34BA5D780076FF76FEB2FA0D5D1D" box="[165,1436,306,327]" gridrow="1" pageId="8" pageNumber="193" rowspan-0="1" rowspan-1="1">
<td id="711D5DC65D780076FBF5FEB2FA0D5D1D" box="[1062,1436,306,327]" gridcol="2" gridrow="1" pageId="8" pageNumber="193">Min Max Mean</td>
</tr>
<tr id="32CC34BA5D780076FF76FED8FA0D5D37" box="[165,1436,344,365]" gridrow="2" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FED8FE725D37" box="[165,483,344,365]" gridcol="0" gridrow="2" pageId="8" pageNumber="193">TOT—Total length</th>
<td id="711D5DC65D780076FC84FED8FC215D37" box="[855,944,344,365]" gridcol="1" gridrow="2" pageId="8" pageNumber="193">730</td>
<td id="711D5DC65D780076FBF5FED8FA0D5D37" box="[1062,1436,344,365]" gridcol="2" gridrow="2" pageId="8" pageNumber="193">740 889</td>
</tr>
<tr id="32CC34BA5D780076FF76FEFEFA0D5DC9" box="[165,1436,382,403]" gridrow="3" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FEFEFE725DC9" box="[165,483,382,403]" gridcol="0" gridrow="3" pageId="8" pageNumber="193">FOR—Fork length</th>
<td id="711D5DC65D780076FC84FEFEFC215DC9" box="[855,944,382,403]" gridcol="1" gridrow="3" pageId="8" pageNumber="193">92.7</td>
<td id="711D5DC65D780076FBF5FEFEFA0D5DC9" box="[1062,1436,382,403]" gridcol="2" gridrow="3" pageId="8" pageNumber="193">91.5 92.8 92.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FE24FA0D5DE3" box="[165,1436,420,441]" gridrow="4" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FE24FE725DE3" box="[165,483,420,441]" gridcol="0" gridrow="4" pageId="8" pageNumber="193">PCL—Precaudal length</th>
<td id="711D5DC65D780076FC84FE24FC215DE3" box="[855,944,420,441]" gridcol="1" gridrow="4" pageId="8" pageNumber="193">83.7</td>
<td id="711D5DC65D780076FBF5FE24FA0D5DE3" box="[1062,1436,420,441]" gridcol="2" gridrow="4" pageId="8" pageNumber="193">83.2 84.6 84.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FE4EFA0D5DB9" box="[165,1436,462,483]" gridrow="5" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FE4EFE725DB9" box="[165,483,462,483]" gridcol="0" gridrow="5" pageId="8" pageNumber="193">PD2—Pre-second dorsal length</th>
<td id="711D5DC65D780076FC84FE4EFC215DB9" box="[855,944,462,483]" gridcol="1" gridrow="5" pageId="8" pageNumber="193">70.7</td>
<td id="711D5DC65D780076FBF5FE4EFA0D5DB9" box="[1062,1436,462,483]" gridcol="2" gridrow="5" pageId="8" pageNumber="193">69.2 70.4 69.8</td>
</tr>
<tr id="32CC34BA5D780076FF76FE74FA0D5E53" box="[165,1436,500,521]" gridrow="6" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FE74FE725E53" box="[165,483,500,521]" gridcol="0" gridrow="6" pageId="8" pageNumber="193">PD1—Pre-first dorsal length</th>
<td id="711D5DC65D780076FC84FE74FC215E53" box="[855,944,500,521]" gridcol="1" gridrow="6" pageId="8" pageNumber="193">48.8</td>
<td id="711D5DC65D780076FBF5FE74FA0D5E53" box="[1062,1436,500,521]" gridcol="2" gridrow="6" pageId="8" pageNumber="193">47.8 48.4 48.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FD9AFA0D5E75" box="[165,1436,538,559]" gridrow="7" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FD9AFE725E75" box="[165,483,538,559]" gridcol="0" gridrow="7" pageId="8" pageNumber="193">PP2—Prepelvic length</th>
<td id="711D5DC65D780076FC84FD9AFC215E75" box="[855,944,538,559]" gridcol="1" gridrow="7" pageId="8" pageNumber="193">45.8</td>
<td id="711D5DC65D780076FBF5FD9AFA0D5E75" box="[1062,1436,538,559]" gridcol="2" gridrow="7" pageId="8" pageNumber="193">44.1 45.6 44.7</td>
</tr>
<tr id="32CC34BA5D780076FF76FDC0FA0D5E0F" box="[165,1436,576,597]" gridrow="8" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FDC0FE725E0F" box="[165,483,576,597]" gridcol="0" gridrow="8" pageId="8" pageNumber="193">SVL—Snoutvent length</th>
<td id="711D5DC65D780076FC84FDC0FC215E0F" box="[855,944,576,597]" gridcol="1" gridrow="8" pageId="8" pageNumber="193">46.0</td>
<td id="711D5DC65D780076FBF5FDC0FA0D5E0F" box="[1062,1436,576,597]" gridcol="2" gridrow="8" pageId="8" pageNumber="193">44.5 46.8 45.8</td>
</tr>
<tr id="32CC34BA5D780076FF76FDE6FA0D5E21" box="[165,1436,614,635]" gridrow="9" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FDE6FE725E21" box="[165,483,614,635]" gridcol="0" gridrow="9" pageId="8" pageNumber="193">PSP—Prespiracular length</th>
<td id="711D5DC65D780076FC84FDE6FC215E21" box="[855,944,614,635]" gridcol="1" gridrow="9" pageId="8" pageNumber="193">21.7</td>
<td id="711D5DC65D780076FBF5FDE6FA0D5E21" box="[1062,1436,614,635]" gridcol="2" gridrow="9" pageId="8" pageNumber="193">21.1 21.4 21.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FD0CFA0D5EFB" box="[165,1436,652,673]" gridrow="10" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FD0CFE725EFB" box="[165,483,652,673]" gridcol="0" gridrow="10" pageId="8" pageNumber="193">PG1—Prebranchial length</th>
<td id="711D5DC65D780076FC84FD0CFC215EFB" box="[855,944,652,673]" gridcol="1" gridrow="10" pageId="8" pageNumber="193">27.5</td>
<td id="711D5DC65D780076FBF5FD0CFA0D5EFB" box="[1062,1436,652,673]" gridcol="2" gridrow="10" pageId="8" pageNumber="193">25.5 27.3 26.5</td>
</tr>
<tr id="32CC34BA5D780076FF76FD32FA0D5E9D" box="[165,1436,690,711]" gridrow="11" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FD32FE725E9D" box="[165,483,690,711]" gridcol="0" gridrow="11" pageId="8" pageNumber="193">HDL—Head length</th>
<td id="711D5DC65D780076FC84FD32FC215E9D" box="[855,944,690,711]" gridcol="1" gridrow="11" pageId="8" pageNumber="193">31.4</td>
<td id="711D5DC65D780076FBF5FD32FA0D5E9D" box="[1062,1436,690,711]" gridcol="2" gridrow="11" pageId="8" pageNumber="193">29.6 31.0 30.5</td>
</tr>
<tr id="32CC34BA5D780076FF76FD58FA0D5EB7" box="[165,1436,728,749]" gridrow="12" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FD58FE725EB7" box="[165,483,728,749]" gridcol="0" gridrow="12" pageId="8" pageNumber="193">POB—Preorbital length (direct)</th>
<td id="711D5DC65D780076FC84FD58FC215EB7" box="[855,944,728,749]" gridcol="1" gridrow="12" pageId="8" pageNumber="193">17.7</td>
<td id="711D5DC65D780076FBF5FD58FA0D5EB7" box="[1062,1436,728,749]" gridcol="2" gridrow="12" pageId="8" pageNumber="193">16.0 17.0 16.7</td>
</tr>
<tr id="32CC34BA5D780076FF76FD7EFA0D5F49" box="[165,1436,766,787]" gridrow="13" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FD7EFE725F49" box="[165,483,766,787]" gridcol="0" gridrow="13" pageId="8" pageNumber="193">POR—Preoral length</th>
<td id="711D5DC65D780076FC84FD7EFC215F49" box="[855,944,766,787]" gridcol="1" gridrow="13" pageId="8" pageNumber="193">21.1</td>
<td id="711D5DC65D780076FBF5FD7EFA0D5F49" box="[1062,1436,766,787]" gridcol="2" gridrow="13" pageId="8" pageNumber="193">19.3 20.5 20.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FCA4FA0D5F63" box="[165,1436,804,825]" gridrow="14" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FCA4FE725F63" box="[165,483,804,825]" gridcol="0" gridrow="14" pageId="8" pageNumber="193">PRN—Prenarial length</th>
<td id="711D5DC65D780076FC84FCA4FC215F63" box="[855,944,804,825]" gridcol="1" gridrow="14" pageId="8" pageNumber="193">16.7</td>
<td id="711D5DC65D780076FBF5FCA4FA0D5F63" box="[1062,1436,804,825]" gridcol="2" gridrow="14" pageId="8" pageNumber="193">15.2 16.3 15.9</td>
</tr>
<tr id="32CC34BA5D780076FF76FCCAFA0D5F05" box="[165,1436,842,863]" gridrow="15" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FCCAFE725F05" box="[165,483,842,863]" gridcol="0" gridrow="15" pageId="8" pageNumber="193">IDS—Interdorsal space</th>
<td id="711D5DC65D780076FC84FCCAFC215F05" box="[855,944,842,863]" gridcol="1" gridrow="15" pageId="8" pageNumber="193">16.2</td>
<td id="711D5DC65D780076FBF5FCCAFA0D5F05" box="[1062,1436,842,863]" gridcol="2" gridrow="15" pageId="8" pageNumber="193">15.5 16.2 15.9</td>
</tr>
<tr id="32CC34BA5D780076FF76FCF0FA0D5FDF" box="[165,1436,880,901]" gridrow="16" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FCF0FE725FDF" box="[165,483,880,901]" gridcol="0" gridrow="16" pageId="8" pageNumber="193">DCS—Dorsalcaudal space</th>
<td id="711D5DC65D780076FC84FCF0FC215FDF" box="[855,944,880,901]" gridcol="1" gridrow="16" pageId="8" pageNumber="193">9.2</td>
<td id="711D5DC65D780076FBF5FCF0FA0D5FDF" box="[1062,1436,880,901]" gridcol="2" gridrow="16" pageId="8" pageNumber="193">8.8 9.6 9.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FC16FA0D5FF1" box="[165,1436,918,939]" gridrow="17" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FC16FE725FF1" box="[165,483,918,939]" gridcol="0" gridrow="17" pageId="8" pageNumber="193">PPS—Pectoralpelvic space</th>
<td id="711D5DC65D780076FC84FC16FC215FF1" box="[855,944,918,939]" gridcol="1" gridrow="17" pageId="8" pageNumber="193">4.8</td>
<td id="711D5DC65D780076FBF5FC16FA0D5FF1" box="[1062,1436,918,939]" gridcol="2" gridrow="17" pageId="8" pageNumber="193">5.2 5.3 5.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FC3CFA0D5F8B" box="[165,1436,956,977]" gridrow="18" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FC3CFE725F8B" box="[165,483,956,977]" gridcol="0" gridrow="18" pageId="8" pageNumber="193">PCS—Pelviccaudal space</th>
<td id="711D5DC65D780076FC84FC3CFC215F8B" box="[855,944,956,977]" gridcol="1" gridrow="18" pageId="8" pageNumber="193">34.7</td>
<td id="711D5DC65D780076FBF5FC3CFA0D5F8B" box="[1062,1436,956,977]" gridcol="2" gridrow="18" pageId="8" pageNumber="193">33.8 34.4 34.1</td>
</tr>
<tr id="32CC34BA5D780076FF76FC62FA0D5FAD" box="[165,1436,994,1015]" gridrow="19" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FC62FE725FAD" box="[165,483,994,1015]" gridcol="0" gridrow="19" pageId="8" pageNumber="193">PDS—Pelvicdorsal space</th>
<td id="711D5DC65D780076FC84FC62FC215FAD" box="[855,944,994,1015]" gridcol="1" gridrow="19" pageId="8" pageNumber="193">2.9</td>
<td id="711D5DC65D780076FBF5FC62FA0D5FAD" box="[1062,1436,994,1015]" gridcol="2" gridrow="19" pageId="8" pageNumber="193">2.4 3.0 2.7</td>
</tr>
<tr id="32CC34BA5D780076FF76FB88FA0D5847" box="[165,1436,1032,1053]" gridrow="20" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FB88FE725847" box="[165,483,1032,1053]" gridcol="0" gridrow="20" pageId="8" pageNumber="193">DW—Disc width</th>
<td id="711D5DC65D780076FC84FB88FC215847" box="[855,944,1032,1053]" gridcol="1" gridrow="20" pageId="8" pageNumber="193">33.8</td>
<td id="711D5DC65D780076FBF5FB88FA0D5847" box="[1062,1436,1032,1053]" gridcol="2" gridrow="20" pageId="8" pageNumber="193">33.1 34.2 33.6</td>
</tr>
<tr id="32CC34BA5D780076FF76FBAEFA0D5819" box="[165,1436,1070,1091]" gridrow="21" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FBAEFE725819" box="[165,483,1070,1091]" gridcol="0" gridrow="21" pageId="8" pageNumber="193">DL—Disc length</th>
<td id="711D5DC65D780076FC84FBAEFC215819" box="[855,944,1070,1091]" gridcol="1" gridrow="21" pageId="8" pageNumber="193">45.1</td>
<td id="711D5DC65D780076FBF5FBAEFA0D5819" box="[1062,1436,1070,1091]" gridcol="2" gridrow="21" pageId="8" pageNumber="193">43.4 44.9 44.1</td>
</tr>
<tr id="32CC34BA5D780076FF76FBD4FA0D5833" box="[165,1436,1108,1129]" gridrow="22" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FBD4FE725833" box="[165,483,1108,1129]" gridcol="0" gridrow="22" pageId="8" pageNumber="193">DT—Disc thickness</th>
<td id="711D5DC65D780076FC84FBD4FC215833" box="[855,944,1108,1129]" gridcol="1" gridrow="22" pageId="8" pageNumber="193">8.3</td>
<td id="711D5DC65D780076FBF5FBD4FA0D5833" box="[1062,1436,1108,1129]" gridcol="2" gridrow="22" pageId="8" pageNumber="193">8.1 8.3 8.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FBFAFA0D58D5" box="[165,1436,1146,1167]" gridrow="23" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FBFAFE7258D5" box="[165,483,1146,1167]" gridcol="0" gridrow="23" pageId="8" pageNumber="193">Snout—Greatest width</th>
<td id="711D5DC65D780076FC84FBFAFC2158D5" box="[855,944,1146,1167]" gridcol="1" gridrow="23" pageId="8" pageNumber="193">34.7</td>
<td id="711D5DC65D780076FBF5FBFAFA0D58D5" box="[1062,1436,1146,1167]" gridcol="2" gridrow="23" pageId="8" pageNumber="193">32.8 33.1 33.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FB20FA0D58EF" box="[165,1436,1184,1205]" gridrow="24" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FB20FE7258EF" box="[165,483,1184,1205]" gridcol="0" gridrow="24" pageId="8" pageNumber="193">SWB—Snout width at base</th>
<td id="711D5DC65D780076FC84FB20FC2158EF" box="[855,944,1184,1205]" gridcol="1" gridrow="24" pageId="8" pageNumber="193">16.3</td>
<td id="711D5DC65D780076FBF5FB20FA0D58EF" box="[1062,1436,1184,1205]" gridcol="2" gridrow="24" pageId="8" pageNumber="193">15.3 16.6 16.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FB46FA0D5881" box="[165,1436,1222,1243]" gridrow="25" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FB46FE725881" box="[165,483,1222,1243]" gridcol="0" gridrow="25" pageId="8" pageNumber="193">COL—Corneal/eye length</th>
<td id="711D5DC65D780076FC84FB46FC215881" box="[855,944,1222,1243]" gridcol="1" gridrow="25" pageId="8" pageNumber="193">2.4</td>
<td id="711D5DC65D780076FBF5FB46FA0D5881" box="[1062,1436,1222,1243]" gridcol="2" gridrow="25" pageId="8" pageNumber="193">1.7 2.5 2.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FB6CFA0D595B" box="[165,1436,1260,1281]" gridrow="26" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FB6CFE72595B" box="[165,483,1260,1281]" gridcol="0" gridrow="26" pageId="8" pageNumber="193">COH—Corneal/eye height</th>
<td id="711D5DC65D780076FC84FB6CFC21595B" box="[855,944,1260,1281]" gridcol="1" gridrow="26" pageId="8" pageNumber="193">1.2</td>
<td id="711D5DC65D780076FBF5FB6CFA0D595B" box="[1062,1436,1260,1281]" gridcol="2" gridrow="26" pageId="8" pageNumber="193">0.9 1.4 1.2</td>
</tr>
<tr id="32CC34BA5D780076FF76FA92FA0D597D" box="[165,1436,1298,1319]" gridrow="27" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FA92FE72597D" box="[165,483,1298,1319]" gridcol="0" gridrow="27" pageId="8" pageNumber="193">EYL—Eye [eyeball] length</th>
<td id="711D5DC65D780076FC84FA92FC21597D" box="[855,944,1298,1319]" gridcol="1" gridrow="27" pageId="8" pageNumber="193">3.9</td>
<td id="711D5DC65D780076FBF5FA92FA0D597D" box="[1062,1436,1298,1319]" gridcol="2" gridrow="27" pageId="8" pageNumber="193">3.9 4.1 4.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FAB8FA0D5917" box="[165,1436,1336,1357]" gridrow="28" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FAB8FE725917" box="[165,483,1336,1357]" gridcol="0" gridrow="28" pageId="8" pageNumber="193">EYH—Eye (eyeball) height</th>
<td id="711D5DC65D780076FC84FAB8FC215917" box="[855,944,1336,1357]" gridcol="1" gridrow="28" pageId="8" pageNumber="193">2.6</td>
<td id="711D5DC65D780076FBF5FAB8FA0D5917" box="[1062,1436,1336,1357]" gridcol="2" gridrow="28" pageId="8" pageNumber="193">2.4 2.6 2.5</td>
</tr>
<tr id="32CC34BA5D780076FF76FADEFA0D5929" box="[165,1436,1374,1395]" gridrow="29" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FADEFE725929" box="[165,483,1374,1395]" gridcol="0" gridrow="29" pageId="8" pageNumber="193">INO—Interorbital space</th>
<td id="711D5DC65D780076FC84FADEFC215929" box="[855,944,1374,1395]" gridcol="1" gridrow="29" pageId="8" pageNumber="193">5.4</td>
<td id="711D5DC65D780076FBF5FADEFA0D5929" box="[1062,1436,1374,1395]" gridcol="2" gridrow="29" pageId="8" pageNumber="193">4.9 5.1 5.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FA04FA0D59C3" box="[165,1436,1412,1433]" gridrow="30" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FA04FE7259C3" box="[165,483,1412,1433]" gridcol="0" gridrow="30" pageId="8" pageNumber="193">SPL—Spiracle length</th>
<td id="711D5DC65D780076FC84FA04FC2159C3" box="[855,944,1412,1433]" gridcol="1" gridrow="30" pageId="8" pageNumber="193">1.5</td>
<td id="711D5DC65D780076FBF5FA04FA0D59C3" box="[1062,1436,1412,1433]" gridcol="2" gridrow="30" pageId="8" pageNumber="193">1.3 2.2 1.6</td>
</tr>
<tr id="32CC34BA5D780076FF76FA2AFA0D59E5" box="[165,1436,1450,1471]" gridrow="31" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FA2AFE7259E5" box="[165,483,1450,1471]" gridcol="0" gridrow="31" pageId="8" pageNumber="193">SPH—Spiracle height</th>
<td id="711D5DC65D780076FC84FA2AFC2159E5" box="[855,944,1450,1471]" gridcol="1" gridrow="31" pageId="8" pageNumber="193">2.2</td>
<td id="711D5DC65D780076FBF5FA2AFA0D59E5" box="[1062,1436,1450,1471]" gridcol="2" gridrow="31" pageId="8" pageNumber="193">1.9 2.0 2.0</td>
</tr>
<tr id="32CC34BA5D780076FF76FA50FA0D59BF" box="[165,1436,1488,1509]" gridrow="32" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FA50FE7259BF" box="[165,483,1488,1509]" gridcol="0" gridrow="32" pageId="8" pageNumber="193">ESL—Eyespiracle space</th>
<td id="711D5DC65D780076FC84FA50FC2159BF" box="[855,944,1488,1509]" gridcol="1" gridrow="32" pageId="8" pageNumber="193">5.6</td>
<td id="711D5DC65D780076FBF5FA50FA0D59BF" box="[1062,1436,1488,1509]" gridcol="2" gridrow="32" pageId="8" pageNumber="193">5.3 5.6 5.4</td>
</tr>
<tr id="32CC34BA5D780076FF76FA76FA0D5A51" box="[165,1436,1526,1547]" gridrow="33" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76FA76FE725A51" box="[165,483,1526,1547]" gridcol="0" gridrow="33" pageId="8" pageNumber="193">INS—Interspiracular space</th>
<td id="711D5DC65D780076FC84FA76FC215A51" box="[855,944,1526,1547]" gridcol="1" gridrow="33" pageId="8" pageNumber="193">6.1</td>
<td id="711D5DC65D780076FBF5FA76FA0D5A51" box="[1062,1436,1526,1547]" gridcol="2" gridrow="33" pageId="8" pageNumber="193">5.9 6.4 6.1</td>
</tr>
<tr id="32CC34BA5D780076FF76F99CFA0D5A6B" box="[165,1436,1564,1585]" gridrow="34" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F99CFE725A6B" box="[165,483,1564,1585]" gridcol="0" gridrow="34" pageId="8" pageNumber="193">NOW—Nostril width</th>
<td id="711D5DC65D780076FC84F99CFC215A6B" box="[855,944,1564,1585]" gridcol="1" gridrow="34" pageId="8" pageNumber="193">5.1</td>
<td id="711D5DC65D780076FBF5F99CFA0D5A6B" box="[1062,1436,1564,1585]" gridcol="2" gridrow="34" pageId="8" pageNumber="193">4.8 4.9 4.8</td>
</tr>
<tr id="32CC34BA5D780076FF76F9C2FA0D5A0D" box="[165,1436,1602,1623]" gridrow="35" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F9C2FE725A0D" box="[165,483,1602,1623]" gridcol="0" gridrow="35" pageId="8" pageNumber="193">INW—Internarial space</th>
<td id="711D5DC65D780076FC84F9C2FC215A0D" box="[855,944,1602,1623]" gridcol="1" gridrow="35" pageId="8" pageNumber="193">4.0</td>
<td id="711D5DC65D780076FBF5F9C2FA0D5A0D" box="[1062,1436,1602,1623]" gridcol="2" gridrow="35" pageId="8" pageNumber="193">3.9 4.2 4.0</td>
</tr>
<tr id="32CC34BA5D780076FF76F9ECFA0D5ADB" box="[165,1436,1644,1665]" gridrow="36" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F9ECFE725ADB" box="[165,483,1644,1665]" gridcol="0" gridrow="36" pageId="8" pageNumber="193">ANF—Anterior nasal flap length</th>
<td id="711D5DC65D780076FC84F9ECFC215ADB" box="[855,944,1644,1665]" gridcol="1" gridrow="36" pageId="8" pageNumber="193">2.0</td>
<td id="711D5DC65D780076FBF5F9ECFA0D5ADB" box="[1062,1436,1644,1665]" gridcol="2" gridrow="36" pageId="8" pageNumber="193">1.7 1.8 1.8</td>
</tr>
<tr id="32CC34BA5D780076FF76F912FA0D5AFD" box="[165,1436,1682,1703]" gridrow="37" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F912FE725AFD" box="[165,483,1682,1703]" gridcol="0" gridrow="37" pageId="8" pageNumber="193">NSE—Nostril to snout edge</th>
<td id="711D5DC65D780076FC84F912FC215AFD" box="[855,944,1682,1703]" gridcol="1" gridrow="37" pageId="8" pageNumber="193">1.7</td>
<td id="711D5DC65D780076FBF5F912FA0D5AFD" box="[1062,1436,1682,1703]" gridcol="2" gridrow="37" pageId="8" pageNumber="193">1.5 1.6 1.5</td>
</tr>
<tr id="32CC34BA5D780076FF76F938FA0D5A97" box="[165,1436,1720,1741]" gridrow="38" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F938FE725A97" box="[165,483,1720,1741]" gridcol="0" gridrow="38" pageId="8" pageNumber="193">MOL—Mouth length</th>
<td id="711D5DC65D780076FC84F938FC215A97" box="[855,944,1720,1741]" gridcol="1" gridrow="38" pageId="8" pageNumber="193">0.5</td>
<td id="711D5DC65D780076FBF5F938FA0D5A97" box="[1062,1436,1720,1741]" gridcol="2" gridrow="38" pageId="8" pageNumber="193">0.4 0.5 0.4</td>
</tr>
<tr id="32CC34BA5D780076FF76F95EFA0D5AA9" box="[165,1436,1758,1779]" gridrow="39" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F95EFE725AA9" box="[165,483,1758,1779]" gridcol="0" gridrow="39" pageId="8" pageNumber="193">MOW—Mouth width</th>
<td id="711D5DC65D780076FC84F95EFC215AA9" box="[855,944,1758,1779]" gridcol="1" gridrow="39" pageId="8" pageNumber="193">6.1</td>
<td id="711D5DC65D780076FBF5F95EFA0D5AA9" box="[1062,1436,1758,1779]" gridcol="2" gridrow="39" pageId="8" pageNumber="193">6.2 6.5 6.3</td>
</tr>
<tr id="32CC34BA5D780076FF76F884FA0D5B43" box="[165,1436,1796,1817]" gridrow="40" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F884FE725B43" box="[165,483,1796,1817]" gridcol="0" gridrow="40" pageId="8" pageNumber="193">GS1—First gill slit height</th>
<td id="711D5DC65D780076FC84F884FC215B43" box="[855,944,1796,1817]" gridcol="1" gridrow="40" pageId="8" pageNumber="193">1.7</td>
<td id="711D5DC65D780076FBF5F884FA0D5B43" box="[1062,1436,1796,1817]" gridcol="2" gridrow="40" pageId="8" pageNumber="193">1.4 1.8 1.6</td>
</tr>
<tr id="32CC34BA5D780076FF76F8AAFA0D5B65" box="[165,1436,1834,1855]" gridrow="41" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F8AAFE725B65" box="[165,483,1834,1855]" gridcol="0" gridrow="41" pageId="8" pageNumber="193">GS2—Second gill slit height</th>
<td id="711D5DC65D780076FC84F8AAFC215B65" box="[855,944,1834,1855]" gridcol="1" gridrow="41" pageId="8" pageNumber="193">1.7</td>
<td id="711D5DC65D780076FBF5F8AAFA0D5B65" box="[1062,1436,1834,1855]" gridcol="2" gridrow="41" pageId="8" pageNumber="193">1.6 1.8 1.7</td>
</tr>
<tr id="32CC34BA5D780076FF76F8D0FA0D5B3F" box="[165,1436,1872,1893]" gridrow="42" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F8D0FE725B3F" box="[165,483,1872,1893]" gridcol="0" gridrow="42" pageId="8" pageNumber="193">GS3—Third gill slit height</th>
<td id="711D5DC65D780076FC84F8D0FC215B3F" box="[855,944,1872,1893]" gridcol="1" gridrow="42" pageId="8" pageNumber="193">1.5</td>
<td id="711D5DC65D780076FBF5F8D0FA0D5B3F" box="[1062,1436,1872,1893]" gridcol="2" gridrow="42" pageId="8" pageNumber="193">1.5 1.7 1.6</td>
</tr>
<tr id="32CC34BA5D780076FF76F8F6FA0D5BD1" box="[165,1436,1910,1931]" gridrow="43" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F8F6FE725BD1" box="[165,483,1910,1931]" gridcol="0" gridrow="43" pageId="8" pageNumber="193">GS4—Fourth gill slit height</th>
<td id="711D5DC65D780076FC84F8F6FC215BD1" box="[855,944,1910,1931]" gridcol="1" gridrow="43" pageId="8" pageNumber="193">1.5</td>
<td id="711D5DC65D780076FBF5F8F6FA0D5BD1" box="[1062,1436,1910,1931]" gridcol="2" gridrow="43" pageId="8" pageNumber="193">1.4 1.7 1.5</td>
</tr>
<tr id="32CC34BA5D780076FF76F81CFA0D5B8E" box="[165,1436,1948,2004]" gridrow="44" pageId="8" pageNumber="193">
<th id="711D5DC65D780076FF76F81CFE725B8E" box="[165,483,1948,2004]" gridcol="0" gridrow="44" pageId="8" pageNumber="193">GS5—Fifth gill slit height</th>
<td id="711D5DC65D780076FC84F81CFC215B8E" box="[855,944,1948,2004]" gridcol="1" gridrow="44" pageId="8" pageNumber="193">1.2</td>
<td id="711D5DC65D780076FBF5F81CFA0D5B8E" box="[1062,1436,1948,2004]" gridcol="2" gridrow="44" pageId="8" pageNumber="193">1.0 1.3 1.2 ...... continued on the next page</td>
</tr>
</table>
</paragraph>
<caption id="D88366705D79FF80FF44FF1BFE1A5CE8" ID-Table-UUID="D88366705D79FF80FF44FF1BFE1A5CE8" box="[151,395,155,178]" httpUri="http://table.plazi.org/id/D88366705D79FF80FF44FF1BFE1A5CE8" pageId="9" pageNumber="194" targetBox="[151,1385,274,1951]" targetIsTable="true" targetPageId="9">
<paragraph id="8C4336F85D79FF80FF44FF1BFE1A5CE8" blockId="9.[151,1385,155,257]" box="[151,395,155,178]" pageId="9" pageNumber="194">
<emphasis id="BE88EAEA5D79FF80FF44FF1BFE985CE8" bold="true" box="[151,265,155,178]" pageId="9" pageNumber="194">TABLE 1.</emphasis>
(continued)
</paragraph>
</caption>
<paragraph id="8C4336F85D79FF80FC84FF46FA875C81" blockId="9.[151,1385,155,257]" box="[855,1302,198,219]" pageId="9" pageNumber="194">
<typeStatus id="5347885A5D79FF80FC84FF46FC215C81" box="[855,944,198,219]" pageId="9" pageNumber="194" type="holotype">Holotype</typeStatus>
<typeStatus id="5347885A5D79FF80FBADFF46FB4A5C81" box="[1150,1243,198,219]" pageId="9" pageNumber="194" type="paratype">Paratypes</typeStatus>
(n=3)
</paragraph>
<paragraph id="8C4336F85D79FF83FBF4FF6CFE325DD2" blockId="9.[151,1385,155,257]" lastBlockId="10.[151,1436,151,896]" lastPageId="10" lastPageNumber="195" pageId="9" pageNumber="194">Min Max Mean Vertebral column with 184 (179181) total centra (excluding segments); 14 (1415) synarcual centra, 28 (29 30) monospondylous centra, 131 (124127) precaudal free centra, 39 (3842) diplospondylous caudal (free) centra, 170 (165166) total free centra. Total synarcual segments 7.6% (7.88.3)%, monospondylous 15.2% (16.1 16.8)%, diplospondylous precaudal centra 56.0% (51.954.2)%, and precaudal free centra 71.2% (68.570.9)% of total centra count. Total pectoral radials 65 (6668): 4 (57) free radials before propterygium, 22 (22) propterygials, 5 (5) mesopterygials, 5 (6) neopterygials, 29 (28) metapterygials, 61 (5961) total basal radials (excluding free radials).</paragraph>
<paragraph id="8C4336F85D79FF80FF76FE92FACC5BC5" pageId="9" pageNumber="194">
<table id="FEFCC4585D790076FF44FE92FAF85BC5" box="[151,1385,274,1951]" gridcols="5" gridrows="43" pageId="9" pageNumber="194">
<tr id="32CC34BA5D790076FF44FE92FAF85D7D" box="[151,1385,274,295]" gridrow="0" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FE92FDA65D7D" box="[151,567,274,295]" gridcol="0" gridrow="0" pageId="9" pageNumber="194">ING1—Inter 1st gill</th>
<th id="711D5DC65D790076FCBDFE92FC095D7D" box="[878,920,274,295]" gridcol="1" gridrow="0" pageId="9" pageNumber="194">14.4</th>
<th id="711D5DC65D790076FBF5FE92FBC15D7D" box="[1062,1104,274,295]" gridcol="2" gridrow="0" pageId="9" pageNumber="194">13.7</th>
<th id="711D5DC65D790076FB7AFE92FB425D7D" box="[1193,1235,274,295]" gridcol="3" gridrow="0" pageId="9" pageNumber="194">14.1</th>
<th id="711D5DC65D790076FAEAFE92FAF85D7D" box="[1337,1385,274,295]" gridcol="4" gridrow="0" pageId="9" pageNumber="194">13.9</th>
</tr>
<tr id="32CC34BA5D790076FF44FEB8FAF85D17" box="[151,1385,312,333]" gridrow="1" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FEB8FDA65D17" box="[151,567,312,333]" gridcol="0" gridrow="1" pageId="9" pageNumber="194">ING5—Inter 5th gill</th>
<td id="711D5DC65D790076FCBDFEB8FC095D17" box="[878,920,312,333]" gridcol="1" gridrow="1" pageId="9" pageNumber="194">10.8</td>
<td id="711D5DC65D790076FBF5FEB8FBC15D17" box="[1062,1104,312,333]" gridcol="2" gridrow="1" pageId="9" pageNumber="194">10.6</td>
<td id="711D5DC65D790076FB7AFEB8FB425D17" box="[1193,1235,312,333]" gridcol="3" gridrow="1" pageId="9" pageNumber="194">10.7</td>
<td id="711D5DC65D790076FAEAFEB8FAF85D17" box="[1337,1385,312,333]" gridcol="4" gridrow="1" pageId="9" pageNumber="194">10.6</td>
</tr>
<tr id="32CC34BA5D790076FF44FEDEFAF85D29" box="[151,1385,350,371]" gridrow="2" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FEDEFDA65D29" box="[151,567,350,371]" gridcol="0" gridrow="2" pageId="9" pageNumber="194">HDH—Head height</th>
<td id="711D5DC65D790076FCBDFEDEFC095D29" box="[878,920,350,371]" gridcol="1" gridrow="2" pageId="9" pageNumber="194">6.7</td>
<td id="711D5DC65D790076FBF5FEDEFBC15D29" box="[1062,1104,350,371]" gridcol="2" gridrow="2" pageId="9" pageNumber="194">6.9</td>
<td id="711D5DC65D790076FB7AFEDEFB425D29" box="[1193,1235,350,371]" gridcol="3" gridrow="2" pageId="9" pageNumber="194">7.5</td>
<td id="711D5DC65D790076FAEAFEDEFAF85D29" box="[1337,1385,350,371]" gridcol="4" gridrow="2" pageId="9" pageNumber="194">7.1</td>
</tr>
<tr id="32CC34BA5D790076FF44FE04FAF85DC3" box="[151,1385,388,409]" gridrow="3" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FE04FDA65DC3" box="[151,567,388,409]" gridcol="0" gridrow="3" pageId="9" pageNumber="194">TRH—Trunk height</th>
<td id="711D5DC65D790076FCBDFE04FC095DC3" box="[878,920,388,409]" gridcol="1" gridrow="3" pageId="9" pageNumber="194">7.7</td>
<td id="711D5DC65D790076FBF5FE04FBC15DC3" box="[1062,1104,388,409]" gridcol="2" gridrow="3" pageId="9" pageNumber="194">7.9</td>
<td id="711D5DC65D790076FB7AFE04FB425DC3" box="[1193,1235,388,409]" gridcol="3" gridrow="3" pageId="9" pageNumber="194">8.3</td>
<td id="711D5DC65D790076FAEAFE04FAF85DC3" box="[1337,1385,388,409]" gridcol="4" gridrow="3" pageId="9" pageNumber="194">8.1</td>
</tr>
<tr id="32CC34BA5D790076FF44FE2AFAF85DE5" box="[151,1385,426,447]" gridrow="4" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FE2AFDA65DE5" box="[151,567,426,447]" gridcol="0" gridrow="4" pageId="9" pageNumber="194">TRW—Trunk width</th>
<td id="711D5DC65D790076FCBDFE2AFC095DE5" box="[878,920,426,447]" gridcol="1" gridrow="4" pageId="9" pageNumber="194">13.2</td>
<td id="711D5DC65D790076FBF5FE2AFBC15DE5" box="[1062,1104,426,447]" gridcol="2" gridrow="4" pageId="9" pageNumber="194">13.3</td>
<td id="711D5DC65D790076FB7AFE2AFB425DE5" box="[1193,1235,426,447]" gridcol="3" gridrow="4" pageId="9" pageNumber="194">13.9</td>
<td id="711D5DC65D790076FAEAFE2AFAF85DE5" box="[1337,1385,426,447]" gridcol="4" gridrow="4" pageId="9" pageNumber="194">13.6</td>
</tr>
<tr id="32CC34BA5D790076FF44FE50FAF85DBF" box="[151,1385,464,485]" gridrow="5" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FE50FDA65DBF" box="[151,567,464,485]" gridcol="0" gridrow="5" pageId="9" pageNumber="194">ABH—Abdomen height</th>
<td id="711D5DC65D790076FCBDFE50FC095DBF" box="[878,920,464,485]" gridcol="1" gridrow="5" pageId="9" pageNumber="194">5.9</td>
<td id="711D5DC65D790076FBF5FE50FBC15DBF" box="[1062,1104,464,485]" gridcol="2" gridrow="5" pageId="9" pageNumber="194">6.2</td>
<td id="711D5DC65D790076FB7AFE50FB425DBF" box="[1193,1235,464,485]" gridcol="3" gridrow="5" pageId="9" pageNumber="194">6.4</td>
<td id="711D5DC65D790076FAEAFE50FAF85DBF" box="[1337,1385,464,485]" gridcol="4" gridrow="5" pageId="9" pageNumber="194">6.3</td>
</tr>
<tr id="32CC34BA5D790076FF44FE76FAF85E51" box="[151,1385,502,523]" gridrow="6" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FE76FDA65E51" box="[151,567,502,523]" gridcol="0" gridrow="6" pageId="9" pageNumber="194">ABW—Abdomen width</th>
<td id="711D5DC65D790076FCBDFE76FC095E51" box="[878,920,502,523]" gridcol="1" gridrow="6" pageId="9" pageNumber="194">11.1</td>
<td id="711D5DC65D790076FBF5FE76FBC15E51" box="[1062,1104,502,523]" gridcol="2" gridrow="6" pageId="9" pageNumber="194">0.0</td>
<td id="711D5DC65D790076FB7AFE76FB425E51" box="[1193,1235,502,523]" gridcol="3" gridrow="6" pageId="9" pageNumber="194">11.6</td>
<td id="711D5DC65D790076FAEAFE76FAF85E51" box="[1337,1385,502,523]" gridcol="4" gridrow="6" pageId="9" pageNumber="194">7.5</td>
</tr>
<tr id="32CC34BA5D790076FF44FDA0FAF85E6F" box="[151,1385,544,565]" gridrow="7" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FDA0FDA65E6F" box="[151,567,544,565]" gridcol="0" gridrow="7" pageId="9" pageNumber="194">CPH—Caudal peduncle height</th>
<td id="711D5DC65D790076FCBDFDA0FC095E6F" box="[878,920,544,565]" gridcol="1" gridrow="7" pageId="9" pageNumber="194">1.7</td>
<td id="711D5DC65D790076FBF5FDA0FBC15E6F" box="[1062,1104,544,565]" gridcol="2" gridrow="7" pageId="9" pageNumber="194">1.6</td>
<td id="711D5DC65D790076FB7AFDA0FB425E6F" box="[1193,1235,544,565]" gridcol="3" gridrow="7" pageId="9" pageNumber="194">1.7</td>
<td id="711D5DC65D790076FAEAFDA0FAF85E6F" box="[1337,1385,544,565]" gridcol="4" gridrow="7" pageId="9" pageNumber="194">1.7</td>
</tr>
<tr id="32CC34BA5D790076FF44FDCAFAF85E05" box="[151,1385,586,607]" gridrow="8" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FDCAFDA65E05" box="[151,567,586,607]" gridcol="0" gridrow="8" pageId="9" pageNumber="194">CPW—Caudal peduncle width</th>
<td id="711D5DC65D790076FCBDFDCAFC095E05" box="[878,920,586,607]" gridcol="1" gridrow="8" pageId="9" pageNumber="194">3.4</td>
<td id="711D5DC65D790076FBF5FDCAFBC15E05" box="[1062,1104,586,607]" gridcol="2" gridrow="8" pageId="9" pageNumber="194">2.9</td>
<td id="711D5DC65D790076FB7AFDCAFB425E05" box="[1193,1235,586,607]" gridcol="3" gridrow="8" pageId="9" pageNumber="194">3.7</td>
<td id="711D5DC65D790076FAEAFDCAFAF85E05" box="[1337,1385,586,607]" gridcol="4" gridrow="8" pageId="9" pageNumber="194">3.3</td>
</tr>
<tr id="32CC34BA5D790076FF44FDF0FAF85EDF" box="[151,1385,624,645]" gridrow="9" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FDF0FDA65EDF" box="[151,567,624,645]" gridcol="0" gridrow="9" pageId="9" pageNumber="194">VNL—Vent length</th>
<td id="711D5DC65D790076FCBDFDF0FC095EDF" box="[878,920,624,645]" gridcol="1" gridrow="9" pageId="9" pageNumber="194">2.7</td>
<td id="711D5DC65D790076FBF5FDF0FBC15EDF" box="[1062,1104,624,645]" gridcol="2" gridrow="9" pageId="9" pageNumber="194">2.3</td>
<td id="711D5DC65D790076FB7AFDF0FB425EDF" box="[1193,1235,624,645]" gridcol="3" gridrow="9" pageId="9" pageNumber="194">2.8</td>
<td id="711D5DC65D790076FAEAFDF0FAF85EDF" box="[1337,1385,624,645]" gridcol="4" gridrow="9" pageId="9" pageNumber="194">2.6</td>
</tr>
<tr id="32CC34BA5D790076FF44FD16FAF85EF1" box="[151,1385,662,683]" gridrow="10" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FD16FDA65EF1" box="[151,567,662,683]" gridcol="0" gridrow="10" pageId="9" pageNumber="194">TFL—Tail fold length</th>
<td id="711D5DC65D790076FCBDFD16FC095EF1" box="[878,920,662,683]" gridcol="1" gridrow="10" pageId="9" pageNumber="194">34.0</td>
<td id="711D5DC65D790076FBF5FD16FBC15EF1" box="[1062,1104,662,683]" gridcol="2" gridrow="10" pageId="9" pageNumber="194">32.7</td>
<td id="711D5DC65D790076FB7AFD16FB425EF1" box="[1193,1235,662,683]" gridcol="3" gridrow="10" pageId="9" pageNumber="194">35.5</td>
<td id="711D5DC65D790076FAEAFD16FAF85EF1" box="[1337,1385,662,683]" gridcol="4" gridrow="10" pageId="9" pageNumber="194">33.9</td>
</tr>
<tr id="32CC34BA5D790076FF44FD3CFAF85E8B" box="[151,1385,700,721]" gridrow="11" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FD3CFDA65E8B" box="[151,567,700,721]" gridcol="0" gridrow="11" pageId="9" pageNumber="194">P1L—Pectoral-fin length</th>
<td id="711D5DC65D790076FCBDFD3CFC095E8B" box="[878,920,700,721]" gridcol="1" gridrow="11" pageId="9" pageNumber="194">21.2</td>
<td id="711D5DC65D790076FBF5FD3CFBC15E8B" box="[1062,1104,700,721]" gridcol="2" gridrow="11" pageId="9" pageNumber="194">19.8</td>
<td id="711D5DC65D790076FB7AFD3CFB425E8B" box="[1193,1235,700,721]" gridcol="3" gridrow="11" pageId="9" pageNumber="194">21.4</td>
<td id="711D5DC65D790076FAEAFD3CFAF85E8B" box="[1337,1385,700,721]" gridcol="4" gridrow="11" pageId="9" pageNumber="194">20.8</td>
</tr>
<tr id="32CC34BA5D790076FF44FD66FAF85EA1" box="[151,1385,742,763]" gridrow="12" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FD66FDA65EA1" box="[151,567,742,763]" gridcol="0" gridrow="12" pageId="9" pageNumber="194">P1A—Pectoral-fin anterior margin</th>
<td id="711D5DC65D790076FCBDFD66FC095EA1" box="[878,920,742,763]" gridcol="1" gridrow="12" pageId="9" pageNumber="194">11.0</td>
<td id="711D5DC65D790076FBF5FD66FBC15EA1" box="[1062,1104,742,763]" gridcol="2" gridrow="12" pageId="9" pageNumber="194">10.9</td>
<td id="711D5DC65D790076FB7AFD66FB425EA1" box="[1193,1235,742,763]" gridcol="3" gridrow="12" pageId="9" pageNumber="194">11.9</td>
<td id="711D5DC65D790076FAEAFD66FAF85EA1" box="[1337,1385,742,763]" gridcol="4" gridrow="12" pageId="9" pageNumber="194">11.4</td>
</tr>
<tr id="32CC34BA5D790076FF44FC8CFAF85F7B" box="[151,1385,780,801]" gridrow="13" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FC8CFDA65F7B" box="[151,567,780,801]" gridcol="0" gridrow="13" pageId="9" pageNumber="194">P1B—Pectoral-fin base</th>
<td id="711D5DC65D790076FCBDFC8CFC095F7B" box="[878,920,780,801]" gridcol="1" gridrow="13" pageId="9" pageNumber="194">15.2</td>
<td id="711D5DC65D790076FBF5FC8CFBC15F7B" box="[1062,1104,780,801]" gridcol="2" gridrow="13" pageId="9" pageNumber="194">15.8</td>
<td id="711D5DC65D790076FB7AFC8CFB425F7B" box="[1193,1235,780,801]" gridcol="3" gridrow="13" pageId="9" pageNumber="194">16.4</td>
<td id="711D5DC65D790076FAEAFC8CFAF85F7B" box="[1337,1385,780,801]" gridcol="4" gridrow="13" pageId="9" pageNumber="194">16.0</td>
</tr>
<tr id="32CC34BA5D790076FF44FCB2FAF85F1D" box="[151,1385,818,839]" gridrow="14" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FCB2FDA65F1D" box="[151,567,818,839]" gridcol="0" gridrow="14" pageId="9" pageNumber="194">P1H—Pectoral-fin height</th>
<td id="711D5DC65D790076FCBDFCB2FC095F1D" box="[878,920,818,839]" gridcol="1" gridrow="14" pageId="9" pageNumber="194">8.6</td>
<td id="711D5DC65D790076FBF5FCB2FBC15F1D" box="[1062,1104,818,839]" gridcol="2" gridrow="14" pageId="9" pageNumber="194">8.3</td>
<td id="711D5DC65D790076FB7AFCB2FB425F1D" box="[1193,1235,818,839]" gridcol="3" gridrow="14" pageId="9" pageNumber="194">8.8</td>
<td id="711D5DC65D790076FAEAFCB2FAF85F1D" box="[1337,1385,818,839]" gridcol="4" gridrow="14" pageId="9" pageNumber="194">8.6</td>
</tr>
<tr id="32CC34BA5D790076FF44FCDCFAF85F2B" box="[151,1385,860,881]" gridrow="15" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FCDCFDA65F2B" box="[151,567,860,881]" gridcol="0" gridrow="15" pageId="9" pageNumber="194">P1P—Pectoral-fin posterior margin</th>
<td id="711D5DC65D790076FCBDFCDCFC095F2B" box="[878,920,860,881]" gridcol="1" gridrow="15" pageId="9" pageNumber="194">14.8</td>
<td id="711D5DC65D790076FBF5FCDCFBC15F2B" box="[1062,1104,860,881]" gridcol="2" gridrow="15" pageId="9" pageNumber="194">14.0</td>
<td id="711D5DC65D790076FB7AFCDCFB425F2B" box="[1193,1235,860,881]" gridcol="3" gridrow="15" pageId="9" pageNumber="194">15.5</td>
<td id="711D5DC65D790076FAEAFCDCFAF85F2B" box="[1337,1385,860,881]" gridcol="4" gridrow="15" pageId="9" pageNumber="194">14.9</td>
</tr>
<tr id="32CC34BA5D790076FF44FC02FAF85FCD" box="[151,1385,898,919]" gridrow="16" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FC02FDA65FCD" box="[151,567,898,919]" gridcol="0" gridrow="16" pageId="9" pageNumber="194">P1I—Pectoral-fin inner margin</th>
<td id="711D5DC65D790076FCBDFC02FC095FCD" box="[878,920,898,919]" gridcol="1" gridrow="16" pageId="9" pageNumber="194">4.8</td>
<td id="711D5DC65D790076FBF5FC02FBC15FCD" box="[1062,1104,898,919]" gridcol="2" gridrow="16" pageId="9" pageNumber="194">4.4</td>
<td id="711D5DC65D790076FB7AFC02FB425FCD" box="[1193,1235,898,919]" gridcol="3" gridrow="16" pageId="9" pageNumber="194">4.6</td>
<td id="711D5DC65D790076FAEAFC02FAF85FCD" box="[1337,1385,898,919]" gridcol="4" gridrow="16" pageId="9" pageNumber="194">4.5</td>
</tr>
<tr id="32CC34BA5D790076FF44FC28FAF85FE7" box="[151,1385,936,957]" gridrow="17" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FC28FDA65FE7" box="[151,567,936,957]" gridcol="0" gridrow="17" pageId="9" pageNumber="194">P2L—Pelvic-fin length</th>
<td id="711D5DC65D790076FCBDFC28FC095FE7" box="[878,920,936,957]" gridcol="1" gridrow="17" pageId="9" pageNumber="194">12.4</td>
<td id="711D5DC65D790076FBF5FC28FBC15FE7" box="[1062,1104,936,957]" gridcol="2" gridrow="17" pageId="9" pageNumber="194">13.3</td>
<td id="711D5DC65D790076FB7AFC28FB425FE7" box="[1193,1235,936,957]" gridcol="3" gridrow="17" pageId="9" pageNumber="194">13.5</td>
<td id="711D5DC65D790076FAEAFC28FAF85FE7" box="[1337,1385,936,957]" gridcol="4" gridrow="17" pageId="9" pageNumber="194">13.4</td>
</tr>
<tr id="32CC34BA5D790076FF44FC4EFAF85FB9" box="[151,1385,974,995]" gridrow="18" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FC4EFDA65FB9" box="[151,567,974,995]" gridcol="0" gridrow="18" pageId="9" pageNumber="194">P2A—Pelvic-fin anterior margin</th>
<td id="711D5DC65D790076FCBDFC4EFC095FB9" box="[878,920,974,995]" gridcol="1" gridrow="18" pageId="9" pageNumber="194">8.0</td>
<td id="711D5DC65D790076FBF5FC4EFBC15FB9" box="[1062,1104,974,995]" gridcol="2" gridrow="18" pageId="9" pageNumber="194">7.2</td>
<td id="711D5DC65D790076FB7AFC4EFB425FB9" box="[1193,1235,974,995]" gridcol="3" gridrow="18" pageId="9" pageNumber="194">7.5</td>
<td id="711D5DC65D790076FAEAFC4EFAF85FB9" box="[1337,1385,974,995]" gridcol="4" gridrow="18" pageId="9" pageNumber="194">7.4</td>
</tr>
<tr id="32CC34BA5D790076FF44FC74FAF85853" box="[151,1385,1012,1033]" gridrow="19" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FC74FDA65853" box="[151,567,1012,1033]" gridcol="0" gridrow="19" pageId="9" pageNumber="194">P2B—Pelvic-fin base</th>
<td id="711D5DC65D790076FCBDFC74FC095853" box="[878,920,1012,1033]" gridcol="1" gridrow="19" pageId="9" pageNumber="194">4.9</td>
<td id="711D5DC65D790076FBF5FC74FBC15853" box="[1062,1104,1012,1033]" gridcol="2" gridrow="19" pageId="9" pageNumber="194">5.8</td>
<td id="711D5DC65D790076FB7AFC74FB425853" box="[1193,1235,1012,1033]" gridcol="3" gridrow="19" pageId="9" pageNumber="194">5.9</td>
<td id="711D5DC65D790076FAEAFC74FAF85853" box="[1337,1385,1012,1033]" gridcol="4" gridrow="19" pageId="9" pageNumber="194">5.9</td>
</tr>
<tr id="32CC34BA5D790076FF44FB9AFAF85875" box="[151,1385,1050,1071]" gridrow="20" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FB9AFDA65875" box="[151,567,1050,1071]" gridcol="0" gridrow="20" pageId="9" pageNumber="194">P2H—Pelvic-fin height</th>
<td id="711D5DC65D790076FCBDFB9AFC095875" box="[878,920,1050,1071]" gridcol="1" gridrow="20" pageId="9" pageNumber="194">5.9</td>
<td id="711D5DC65D790076FBF5FB9AFBC15875" box="[1062,1104,1050,1071]" gridcol="2" gridrow="20" pageId="9" pageNumber="194">5.8</td>
<td id="711D5DC65D790076FB7AFB9AFB425875" box="[1193,1235,1050,1071]" gridcol="3" gridrow="20" pageId="9" pageNumber="194">6.5</td>
<td id="711D5DC65D790076FAEAFB9AFAF85875" box="[1337,1385,1050,1071]" gridcol="4" gridrow="20" pageId="9" pageNumber="194">6.1</td>
</tr>
<tr id="32CC34BA5D790076FF44FBC4FAF85803" box="[151,1385,1092,1113]" gridrow="21" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FBC4FDA65803" box="[151,567,1092,1113]" gridcol="0" gridrow="21" pageId="9" pageNumber="194">P2P—Pelvic-fin posterior margin length</th>
<td id="711D5DC65D790076FCBDFBC4FC095803" box="[878,920,1092,1113]" gridcol="1" gridrow="21" pageId="9" pageNumber="194">8.3</td>
<td id="711D5DC65D790076FBF5FBC4FBC15803" box="[1062,1104,1092,1113]" gridcol="2" gridrow="21" pageId="9" pageNumber="194">8.8</td>
<td id="711D5DC65D790076FB7AFBC4FB425803" box="[1193,1235,1092,1113]" gridcol="3" gridrow="21" pageId="9" pageNumber="194">9.9</td>
<td id="711D5DC65D790076FAEAFBC4FAF85803" box="[1337,1385,1092,1113]" gridcol="4" gridrow="21" pageId="9" pageNumber="194">9.2</td>
</tr>
<tr id="32CC34BA5D790076FF44FBEEFAF858D9" box="[151,1385,1134,1155]" gridrow="22" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FBEEFDA658D9" box="[151,567,1134,1155]" gridcol="0" gridrow="22" pageId="9" pageNumber="194">P2I—Pelvic-fin inner margin length</th>
<td id="711D5DC65D790076FCBDFBEEFC0958D9" box="[878,920,1134,1155]" gridcol="1" gridrow="22" pageId="9" pageNumber="194">7.4</td>
<td id="711D5DC65D790076FBF5FBEEFBC158D9" box="[1062,1104,1134,1155]" gridcol="2" gridrow="22" pageId="9" pageNumber="194">7.6</td>
<td id="711D5DC65D790076FB7AFBEEFB4258D9" box="[1193,1235,1134,1155]" gridcol="3" gridrow="22" pageId="9" pageNumber="194">8.9</td>
<td id="711D5DC65D790076FAEAFBEEFAF858D9" box="[1337,1385,1134,1155]" gridcol="4" gridrow="22" pageId="9" pageNumber="194">8.2</td>
</tr>
<tr id="32CC34BA5D790076FF44FB14FAF858F3" box="[151,1385,1172,1193]" gridrow="23" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FB14FDA658F3" box="[151,567,1172,1193]" gridcol="0" gridrow="23" pageId="9" pageNumber="194">P2S—Pelvic-fin span</th>
<td id="711D5DC65D790076FCBDFB14FC0958F3" box="[878,920,1172,1193]" gridcol="1" gridrow="23" pageId="9" pageNumber="194">19.4</td>
<td id="711D5DC65D790076FBF5FB14FBC158F3" box="[1062,1104,1172,1193]" gridcol="2" gridrow="23" pageId="9" pageNumber="194">18.9</td>
<td id="711D5DC65D790076FB7AFB14FB4258F3" box="[1193,1235,1172,1193]" gridcol="3" gridrow="23" pageId="9" pageNumber="194">20.0</td>
<td id="711D5DC65D790076FAEAFB14FAF858F3" box="[1337,1385,1172,1193]" gridcol="4" gridrow="23" pageId="9" pageNumber="194">19.4</td>
</tr>
<tr id="32CC34BA5D790076FF44FB3AFAF85895" box="[151,1385,1210,1231]" gridrow="24" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FB3AFDA65895" box="[151,567,1210,1231]" gridcol="0" gridrow="24" pageId="9" pageNumber="194">CLO—Clasper outer length</th>
<td id="711D5DC65D790076FCBDFB3AFC095895" box="[878,920,1210,1231]" gridcol="1" gridrow="24" pageId="9" pageNumber="194">3.2</td>
<td id="711D5DC65D790076FBF5FB3AFBC15895" box="[1062,1104,1210,1231]" gridcol="2" gridrow="24" pageId="9" pageNumber="194">0.0</td>
<td id="711D5DC65D790076FB7AFB3AFB425895" box="[1193,1235,1210,1231]" gridcol="3" gridrow="24" pageId="9" pageNumber="194">4.0</td>
<td id="711D5DC65D790076FAEAFB3AFAF85895" box="[1337,1385,1210,1231]" gridcol="4" gridrow="24" pageId="9" pageNumber="194">2.0</td>
</tr>
<tr id="32CC34BA5D790076FF44FB60FAF858AF" box="[151,1385,1248,1269]" gridrow="25" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FB60FDA658AF" box="[151,567,1248,1269]" gridcol="0" gridrow="25" pageId="9" pageNumber="194">CLI—Clasper inner length</th>
<td id="711D5DC65D790076FCBDFB60FC0958AF" box="[878,920,1248,1269]" gridcol="1" gridrow="25" pageId="9" pageNumber="194">7.3</td>
<td id="711D5DC65D790076FBF5FB60FBC158AF" box="[1062,1104,1248,1269]" gridcol="2" gridrow="25" pageId="9" pageNumber="194">0.0</td>
<td id="711D5DC65D790076FB7AFB60FB4258AF" box="[1193,1235,1248,1269]" gridcol="3" gridrow="25" pageId="9" pageNumber="194">9.2</td>
<td id="711D5DC65D790076FAEAFB60FAF858AF" box="[1337,1385,1248,1269]" gridcol="4" gridrow="25" pageId="9" pageNumber="194">4.6</td>
</tr>
<tr id="32CC34BA5D790076FF44FA86FAF85941" box="[151,1385,1286,1307]" gridrow="26" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FA86FDA65941" box="[151,567,1286,1307]" gridcol="0" gridrow="26" pageId="9" pageNumber="194">CLB—Clasper base width</th>
<td id="711D5DC65D790076FCBDFA86FC095941" box="[878,920,1286,1307]" gridcol="1" gridrow="26" pageId="9" pageNumber="194">0.6</td>
<td id="711D5DC65D790076FBF5FA86FBC15941" box="[1062,1104,1286,1307]" gridcol="2" gridrow="26" pageId="9" pageNumber="194">0.0</td>
<td id="711D5DC65D790076FB7AFA86FB425941" box="[1193,1235,1286,1307]" gridcol="3" gridrow="26" pageId="9" pageNumber="194">0.6</td>
<td id="711D5DC65D790076FAEAFA86FAF85941" box="[1337,1385,1286,1307]" gridcol="4" gridrow="26" pageId="9" pageNumber="194">0.3</td>
</tr>
<tr id="32CC34BA5D790076FF44FAACFAF8591B" box="[151,1385,1324,1345]" gridrow="27" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FAACFDA6591B" box="[151,567,1324,1345]" gridcol="0" gridrow="27" pageId="9" pageNumber="194">D1L—First dorsal-fin length</th>
<td id="711D5DC65D790076FCBDFAACFC09591B" box="[878,920,1324,1345]" gridcol="1" gridrow="27" pageId="9" pageNumber="194">11.7</td>
<td id="711D5DC65D790076FBF5FAACFBC1591B" box="[1062,1104,1324,1345]" gridcol="2" gridrow="27" pageId="9" pageNumber="194">11.3</td>
<td id="711D5DC65D790076FB7AFAACFB42591B" box="[1193,1235,1324,1345]" gridcol="3" gridrow="27" pageId="9" pageNumber="194">12.9</td>
<td id="711D5DC65D790076FAEAFAACFAF8591B" box="[1337,1385,1324,1345]" gridcol="4" gridrow="27" pageId="9" pageNumber="194">12.0</td>
</tr>
<tr id="32CC34BA5D790076FF44FAD6FAF85931" box="[151,1385,1366,1387]" gridrow="28" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FAD6FDA65931" box="[151,567,1366,1387]" gridcol="0" gridrow="28" pageId="9" pageNumber="194">D1A—First dorsal-fin anterior margin</th>
<td id="711D5DC65D790076FCBDFAD6FC095931" box="[878,920,1366,1387]" gridcol="1" gridrow="28" pageId="9" pageNumber="194">13.6</td>
<td id="711D5DC65D790076FBF5FAD6FBC15931" box="[1062,1104,1366,1387]" gridcol="2" gridrow="28" pageId="9" pageNumber="194">13.6</td>
<td id="711D5DC65D790076FB7AFAD6FB425931" box="[1193,1235,1366,1387]" gridcol="3" gridrow="28" pageId="9" pageNumber="194">15.4</td>
<td id="711D5DC65D790076FAEAFAD6FAF85931" box="[1337,1385,1366,1387]" gridcol="4" gridrow="28" pageId="9" pageNumber="194">14.3</td>
</tr>
<tr id="32CC34BA5D790076FF44FAFCFAF859CB" box="[151,1385,1404,1425]" gridrow="29" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FAFCFDA659CB" box="[151,567,1404,1425]" gridcol="0" gridrow="29" pageId="9" pageNumber="194">D1B—First dorsal-fin base</th>
<td id="711D5DC65D790076FCBDFAFCFC0959CB" box="[878,920,1404,1425]" gridcol="1" gridrow="29" pageId="9" pageNumber="194">5.9</td>
<td id="711D5DC65D790076FBF5FAFCFBC159CB" box="[1062,1104,1404,1425]" gridcol="2" gridrow="29" pageId="9" pageNumber="194">6.0</td>
<td id="711D5DC65D790076FB7AFAFCFB4259CB" box="[1193,1235,1404,1425]" gridcol="3" gridrow="29" pageId="9" pageNumber="194">6.7</td>
<td id="711D5DC65D790076FAEAFAFCFAF859CB" box="[1337,1385,1404,1425]" gridcol="4" gridrow="29" pageId="9" pageNumber="194">6.3</td>
</tr>
<tr id="32CC34BA5D790076FF44FA22FAF859ED" box="[151,1385,1442,1463]" gridrow="30" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FA22FDA659ED" box="[151,567,1442,1463]" gridcol="0" gridrow="30" pageId="9" pageNumber="194">D1H—First dorsal-fin height</th>
<td id="711D5DC65D790076FCBDFA22FC0959ED" box="[878,920,1442,1463]" gridcol="1" gridrow="30" pageId="9" pageNumber="194">9.5</td>
<td id="711D5DC65D790076FBF5FA22FBC159ED" box="[1062,1104,1442,1463]" gridcol="2" gridrow="30" pageId="9" pageNumber="194">8.9</td>
<td id="711D5DC65D790076FB7AFA22FB4259ED" box="[1193,1235,1442,1463]" gridcol="3" gridrow="30" pageId="9" pageNumber="194">9.5</td>
<td id="711D5DC65D790076FAEAFA22FAF859ED" box="[1337,1385,1442,1463]" gridcol="4" gridrow="30" pageId="9" pageNumber="194">9.2</td>
</tr>
<tr id="32CC34BA5D790076FF44FA4CFAF859BB" box="[151,1385,1484,1505]" gridrow="31" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FA4CFDA659BB" box="[151,567,1484,1505]" gridcol="0" gridrow="31" pageId="9" pageNumber="194">D1P—First dorsal-fin posterior margin</th>
<td id="711D5DC65D790076FCBDFA4CFC0959BB" box="[878,920,1484,1505]" gridcol="1" gridrow="31" pageId="9" pageNumber="194">9.3</td>
<td id="711D5DC65D790076FBF5FA4CFBC159BB" box="[1062,1104,1484,1505]" gridcol="2" gridrow="31" pageId="9" pageNumber="194">9.1</td>
<td id="711D5DC65D790076FB7AFA4CFB4259BB" box="[1193,1235,1484,1505]" gridcol="3" gridrow="31" pageId="9" pageNumber="194">9.7</td>
<td id="711D5DC65D790076FAEAFA4CFAF859BB" box="[1337,1385,1484,1505]" gridcol="4" gridrow="31" pageId="9" pageNumber="194">9.4</td>
</tr>
<tr id="32CC34BA5D790076FF44FA76FAF85A51" box="[151,1385,1526,1547]" gridrow="32" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44FA76FDA65A51" box="[151,567,1526,1547]" gridcol="0" gridrow="32" pageId="9" pageNumber="194">D1I—First dorsal-fin inner margin</th>
<td id="711D5DC65D790076FCBDFA76FC095A51" box="[878,920,1526,1547]" gridcol="1" gridrow="32" pageId="9" pageNumber="194">5.8</td>
<td id="711D5DC65D790076FBF5FA76FBC15A51" box="[1062,1104,1526,1547]" gridcol="2" gridrow="32" pageId="9" pageNumber="194">5.6</td>
<td id="711D5DC65D790076FB7AFA76FB425A51" box="[1193,1235,1526,1547]" gridcol="3" gridrow="32" pageId="9" pageNumber="194">6.4</td>
<td id="711D5DC65D790076FAEAFA76FAF85A51" box="[1337,1385,1526,1547]" gridcol="4" gridrow="32" pageId="9" pageNumber="194">5.9</td>
</tr>
<tr id="32CC34BA5D790076FF44F99CFAF85A6B" box="[151,1385,1564,1585]" gridrow="33" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F99CFDA65A6B" box="[151,567,1564,1585]" gridcol="0" gridrow="33" pageId="9" pageNumber="194">D2L—Second dorsal-fin length</th>
<td id="711D5DC65D790076FCBDF99CFC095A6B" box="[878,920,1564,1585]" gridcol="1" gridrow="33" pageId="9" pageNumber="194">9.5</td>
<td id="711D5DC65D790076FBF5F99CFBC15A6B" box="[1062,1104,1564,1585]" gridcol="2" gridrow="33" pageId="9" pageNumber="194">9.3</td>
<td id="711D5DC65D790076FB7AF99CFB425A6B" box="[1193,1235,1564,1585]" gridcol="3" gridrow="33" pageId="9" pageNumber="194">10.6</td>
<td id="711D5DC65D790076FAEAF99CFAF85A6B" box="[1337,1385,1564,1585]" gridcol="4" gridrow="33" pageId="9" pageNumber="194">9.9</td>
</tr>
<tr id="32CC34BA5D790076FF44F9C6FAF85A01" box="[151,1385,1606,1627]" gridrow="34" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F9C6FDA65A01" box="[151,567,1606,1627]" gridcol="0" gridrow="34" pageId="9" pageNumber="194">D2A—Second dorsal-fin anterior margin</th>
<td id="711D5DC65D790076FCBDF9C6FC095A01" box="[878,920,1606,1627]" gridcol="1" gridrow="34" pageId="9" pageNumber="194">11.4</td>
<td id="711D5DC65D790076FBF5F9C6FBC15A01" box="[1062,1104,1606,1627]" gridcol="2" gridrow="34" pageId="9" pageNumber="194">11.4</td>
<td id="711D5DC65D790076FB7AF9C6FB425A01" box="[1193,1235,1606,1627]" gridcol="3" gridrow="34" pageId="9" pageNumber="194">12.4</td>
<td id="711D5DC65D790076FAEAF9C6FAF85A01" box="[1337,1385,1606,1627]" gridcol="4" gridrow="34" pageId="9" pageNumber="194">11.8</td>
</tr>
<tr id="32CC34BA5D790076FF44F9ECFAF85ADB" box="[151,1385,1644,1665]" gridrow="35" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F9ECFDA65ADB" box="[151,567,1644,1665]" gridcol="0" gridrow="35" pageId="9" pageNumber="194">D2B—Second dorsal-fin base</th>
<td id="711D5DC65D790076FCBDF9ECFC095ADB" box="[878,920,1644,1665]" gridcol="1" gridrow="35" pageId="9" pageNumber="194">4.7</td>
<td id="711D5DC65D790076FBF5F9ECFBC15ADB" box="[1062,1104,1644,1665]" gridcol="2" gridrow="35" pageId="9" pageNumber="194">4.9</td>
<td id="711D5DC65D790076FB7AF9ECFB425ADB" box="[1193,1235,1644,1665]" gridcol="3" gridrow="35" pageId="9" pageNumber="194">5.3</td>
<td id="711D5DC65D790076FAEAF9ECFAF85ADB" box="[1337,1385,1644,1665]" gridcol="4" gridrow="35" pageId="9" pageNumber="194">5.1</td>
</tr>
<tr id="32CC34BA5D790076FF44F912FAF85AFD" box="[151,1385,1682,1703]" gridrow="36" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F912FDA65AFD" box="[151,567,1682,1703]" gridcol="0" gridrow="36" pageId="9" pageNumber="194">D2H—Second dorsal-fin height</th>
<td id="711D5DC65D790076FCBDF912FC095AFD" box="[878,920,1682,1703]" gridcol="1" gridrow="36" pageId="9" pageNumber="194">7.8</td>
<td id="711D5DC65D790076FBF5F912FBC15AFD" box="[1062,1104,1682,1703]" gridcol="2" gridrow="36" pageId="9" pageNumber="194">6.9</td>
<td id="711D5DC65D790076FB7AF912FB425AFD" box="[1193,1235,1682,1703]" gridcol="3" gridrow="36" pageId="9" pageNumber="194">7.9</td>
<td id="711D5DC65D790076FAEAF912FAF85AFD" box="[1337,1385,1682,1703]" gridcol="4" gridrow="36" pageId="9" pageNumber="194">7.3</td>
</tr>
<tr id="32CC34BA5D790076FF44F93CFAF85A8B" box="[151,1385,1724,1745]" gridrow="37" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F93CFDA65A8B" box="[151,567,1724,1745]" gridcol="0" gridrow="37" pageId="9" pageNumber="194">D2P—Second dorsal-fin posterior margin</th>
<td id="711D5DC65D790076FCBDF93CFC095A8B" box="[878,920,1724,1745]" gridcol="1" gridrow="37" pageId="9" pageNumber="194">7.5</td>
<td id="711D5DC65D790076FBF5F93CFBC15A8B" box="[1062,1104,1724,1745]" gridcol="2" gridrow="37" pageId="9" pageNumber="194">7.0</td>
<td id="711D5DC65D790076FB7AF93CFB425A8B" box="[1193,1235,1724,1745]" gridcol="3" gridrow="37" pageId="9" pageNumber="194">7.4</td>
<td id="711D5DC65D790076FAEAF93CFAF85A8B" box="[1337,1385,1724,1745]" gridcol="4" gridrow="37" pageId="9" pageNumber="194">7.2</td>
</tr>
<tr id="32CC34BA5D790076FF44F966FAF85AA1" box="[151,1385,1766,1787]" gridrow="38" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F966FDA65AA1" box="[151,567,1766,1787]" gridcol="0" gridrow="38" pageId="9" pageNumber="194">D2I—Second dorsal-fin inner margin</th>
<td id="711D5DC65D790076FCBDF966FC095AA1" box="[878,920,1766,1787]" gridcol="1" gridrow="38" pageId="9" pageNumber="194">4.9</td>
<td id="711D5DC65D790076FBF5F966FBC15AA1" box="[1062,1104,1766,1787]" gridcol="2" gridrow="38" pageId="9" pageNumber="194">4.6</td>
<td id="711D5DC65D790076FB7AF966FB425AA1" box="[1193,1235,1766,1787]" gridcol="3" gridrow="38" pageId="9" pageNumber="194">5.4</td>
<td id="711D5DC65D790076FAEAF966FAF85AA1" box="[1337,1385,1766,1787]" gridcol="4" gridrow="38" pageId="9" pageNumber="194">5.0</td>
</tr>
<tr id="32CC34BA5D790076FF44F88CFAF85B7B" box="[151,1385,1804,1825]" gridrow="39" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F88CFDA65B7B" box="[151,567,1804,1825]" gridcol="0" gridrow="39" pageId="9" pageNumber="194">CDM—Dorsal caudal margin</th>
<td id="711D5DC65D790076FCBDF88CFC095B7B" box="[878,920,1804,1825]" gridcol="1" gridrow="39" pageId="9" pageNumber="194">15.4</td>
<td id="711D5DC65D790076FBF5F88CFBC15B7B" box="[1062,1104,1804,1825]" gridcol="2" gridrow="39" pageId="9" pageNumber="194">15.4</td>
<td id="711D5DC65D790076FB7AF88CFB425B7B" box="[1193,1235,1804,1825]" gridcol="3" gridrow="39" pageId="9" pageNumber="194">16.9</td>
<td id="711D5DC65D790076FAEAF88CFAF85B7B" box="[1337,1385,1804,1825]" gridcol="4" gridrow="39" pageId="9" pageNumber="194">16.1</td>
</tr>
<tr id="32CC34BA5D790076FF44F8B6FAF85B11" box="[151,1385,1846,1867]" gridrow="40" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F8B6FDA65B11" box="[151,567,1846,1867]" gridcol="0" gridrow="40" pageId="9" pageNumber="194">CPV—Preventral caudal margin</th>
<td id="711D5DC65D790076FCBDF8B6FC095B11" box="[878,920,1846,1867]" gridcol="1" gridrow="40" pageId="9" pageNumber="194">10.8</td>
<td id="711D5DC65D790076FBF5F8B6FBC15B11" box="[1062,1104,1846,1867]" gridcol="2" gridrow="40" pageId="9" pageNumber="194">9.6</td>
<td id="711D5DC65D790076FB7AF8B6FB425B11" box="[1193,1235,1846,1867]" gridcol="3" gridrow="40" pageId="9" pageNumber="194">11.1</td>
<td id="711D5DC65D790076FAEAF8B6FAF85B11" box="[1337,1385,1846,1867]" gridcol="4" gridrow="40" pageId="9" pageNumber="194">10.5</td>
</tr>
<tr id="32CC34BA5D790076FF44F8E0FAF85B2F" box="[151,1385,1888,1909]" gridrow="41" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F8E0FDA65B2F" box="[151,567,1888,1909]" gridcol="0" gridrow="41" pageId="9" pageNumber="194">CPL—Lower postventral caudal margin</th>
<td id="711D5DC65D790076FCBDF8E0FC095B2F" box="[878,920,1888,1909]" gridcol="1" gridrow="41" pageId="9" pageNumber="194">3.3</td>
<td id="711D5DC65D790076FBF5F8E0FBC15B2F" box="[1062,1104,1888,1909]" gridcol="2" gridrow="41" pageId="9" pageNumber="194">3.3</td>
<td id="711D5DC65D790076FB7AF8E0FB425B2F" box="[1193,1235,1888,1909]" gridcol="3" gridrow="41" pageId="9" pageNumber="194">4.3</td>
<td id="711D5DC65D790076FAEAF8E0FAF85B2F" box="[1337,1385,1888,1909]" gridcol="4" gridrow="41" pageId="9" pageNumber="194">3.7</td>
</tr>
<tr id="32CC34BA5D790076FF44F80AFAF85BC5" box="[151,1385,1930,1951]" gridrow="42" pageId="9" pageNumber="194">
<th id="711D5DC65D790076FF44F80AFDA65BC5" box="[151,567,1930,1951]" gridcol="0" gridrow="42" pageId="9" pageNumber="194">CPU—Upper postventral caudal margin</th>
<td id="711D5DC65D790076FCBDF80AFC095BC5" box="[878,920,1930,1951]" gridcol="1" gridrow="42" pageId="9" pageNumber="194">9.5</td>
<td id="711D5DC65D790076FBF5F80AFBC15BC5" box="[1062,1104,1930,1951]" gridcol="2" gridrow="42" pageId="9" pageNumber="194">9.4</td>
<td id="711D5DC65D790076FB7AF80AFB425BC5" box="[1193,1235,1930,1951]" gridcol="3" gridrow="42" pageId="9" pageNumber="194">10.3</td>
<td id="711D5DC65D790076FAEAF80AFAF85BC5" box="[1337,1385,1930,1951]" gridcol="4" gridrow="42" pageId="9" pageNumber="194">9.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8C4336F85D7AFF83FF14FE13FCF85FDA" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="195">
<emphasis id="BE88EAEA5D7AFF83FF14FE13FE545DF6" bold="true" box="[199,453,403,428]" pageId="10" pageNumber="195">Colour (when fresh).</emphasis>
Based on
<typeStatus id="5347885A5D7AFF83FD97FE14FD375DF7" box="[580,678,404,429]" pageId="10" pageNumber="195" type="holotype">holotype</typeStatus>
NMMBP16274 unless stated otherwise: Dorsal surface of body mostly medium to dark greenish brown (becoming greyish on preservation); paler, yellowish brown laterally beside rostral cartilage on snout and along hind margins of pectoral and pelvic fins; black pectoral marking absent, instead region plain or with 1 (
<quantity id="4B049B1D5D7AFF83FDD3FD80FDDB5E42" box="[512,586,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="10" pageNumber="195" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
<typeStatus id="5347885A5D7AFF83FD87FD80FD505E42" box="[596,705,512,536]" pageId="10" pageNumber="195" type="paratype">paratypes</typeStatus>
) small, diffuse-edged white spots; a few similar, isolated white spots near insertion of pectoral fins; single white spot above origin of pelvic fin; a few white spots at bases of dorsal fins (sometimes also beneath their inner margins); a well-developed, single row of small white spots on flank originating over origin of pelvic fin, then coalescing beneath first dorsal fin to form a narrow white stripe; stripe extending along dorsolateral surface of each side of tail to anterior part of caudal peduncle; dorsal and caudal fins yellowish brown, paler than body, tips dusky; orbit white with a broad black bar on its upper surface; suborbit pale; interorbit with a white spots on each side near obit and with or without a dark medium blotch. Ventral surface almost uniformly white, strongly contrasted with dorsal surface; anterior half of snout either side of rostral shaft with broad, irregular, semicircular, blackish patch (faint is some preserved material); dark patch also between nostrils in NMMBP16275; pectoral-fin with narrow dusky tips. In preservative, pale areas on dorsal surface becoming more strongly demarcated from darker areas adjacent.
</paragraph>
<caption id="D88366705D7AFF83FF44F822FE725B81" httpUri="https://zenodo.org/record/219793/files/figure.png" pageId="10" pageNumber="195" targetBox="[339,1246,940,1926]" targetPageId="10">
<paragraph id="8C4336F85D7AFF83FF44F822FE725B81" blockId="10.[151,1437,1954,2011]" pageId="10" pageNumber="195">
<emphasis id="BE88EAEA5D7AFF83FF44F822FE9D5BED" bold="true" box="[151,268,1954,1975]" pageId="10" pageNumber="195">FIGURE 8.</emphasis>
Distribution of
<taxonomicName id="4BFC4D7B5D7AFF83FE75F822FD3E5BED" box="[422,687,1954,1975]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="195" phylum="Chordata" rank="species" species="immaculatus" status="sp. nov.">
<emphasis id="BE88EAEA5D7AFF83FE75F822FD3E5BED" box="[422,687,1954,1975]" italics="true" pageId="10" pageNumber="195">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5BB57915D7AFF83FD66F822FC905BED" box="[693,769,1954,1975]" pageId="10" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
Yellow star—holotype; yellow dots—market sites for other material. Map extracted from Google Earth.
</paragraph>
</caption>
<paragraph id="8C4336F85D7BFF82FF14FF17FC925D47" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
<emphasis id="BE88EAEA5D7BFF82FF14FF17FF6F5CEA" bold="true" box="[199,254,151,176]" pageId="11" pageNumber="196">Size.</emphasis>
Reaches at least
<quantity id="4B049B1D5D7BFF82FE00FF18FD8D5CF5" box="[467,540,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="9.9" pageId="11" pageNumber="196" unit="cm" value="99.0">99 cm</quantity>
TL (non-type NMMBP15684), but this specimen and all five other males examined (
<quantity id="4B049B1D5D7BFF82FEC0FF3CFE125C8E" box="[275,387,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="8.9" metricValueMin="4.1" pageId="11" pageNumber="196" unit="cm" value="65.0" valueMax="89.0" valueMin="41.0">4189 cm</quantity>
TL) were immature. The dorsal and caudal fins were well developed in NMMBP15684 (see
<figureCitation id="14C72A7D5D7BFF82FF44FF5FFF665CA2" box="[151,247,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1899,1921]" captionTargetBox="[767,1436,475,1874]" captionTargetId="figure@5.[767,1372,964,1460]" captionTargetPageId="5" captionText="FIGURE 4. Rhynchobatus immaculatus sp. nov. fin shape and development. Immature male holotype (NMMB P 16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMB P 15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin." httpUri="https://zenodo.org/record/219789/files/figure.png" pageId="11" pageNumber="196">Figure 4</figureCitation>
), so it is likely to be a medium-size wedgefish with a maximum length less than
<quantity id="4B049B1D5D7BFF82FB52FF5FFB495CAD" box="[1153,1240,223,248]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="196" unit="cm" value="150.0">150 cm</quantity>
. An investigation of size structure in the Taiwanese population is needed.
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D7BFF82FF14FEA7FBE55D3F" pageId="11" pageNumber="196" type="distribution">
<paragraph id="8C4336F85D7BFF82FF14FEA7FBE55D3F" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
<emphasis id="BE88EAEA5D7BFF82FF14FEA7FEF15D1A" bold="true" box="[199,352,295,320]" pageId="11" pageNumber="196">Distribution.</emphasis>
Known from seas adjacent
<collectingCountry id="F4EB76685D7BFF82FD63FEA7FC975D1A" box="[688,774,295,320]" name="Taiwan" pageId="11" pageNumber="196">Taiwan</collectingCountry>
. Possibly more widespread locally in the western North Pacific, but positive confirmation is needed as these fishes are frequently mis-identified.
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D7BFF82FF14FEF0FB135D8A" pageId="11" pageNumber="196" type="etymology">
<paragraph id="8C4336F85D7BFF82FF14FEF0FB135D8A" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
<emphasis id="BE88EAEA5D7BFF82FF14FEF0FEDF5DD2" bold="true" box="[199,334,368,392]" pageId="11" pageNumber="196">Etymology.</emphasis>
The epithet, immaculate, is a combination of the Latin
<emphasis id="BE88EAEA5D7BFF82FC1EFEF0FC785DDD" box="[973,1001,368,391]" italics="true" pageId="11" pageNumber="196">im</emphasis>
(not) and
<emphasis id="BE88EAEA5D7BFF82FBB1FEF0FB245DD2" box="[1122,1205,368,392]" italics="true" pageId="11" pageNumber="196">macula</emphasis>
(spot, mark), and is based on the lack of a dark pectoral marking (rather than any white spots) which is otherwise present in small individuals of other nominal species of
<taxonomicName id="4BFC4D7B5D7BFF82FD81FE38FD605D8A" box="[594,753,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="genus">
<emphasis id="BE88EAEA5D7BFF82FD81FE38FD605D8A" box="[594,753,440,464]" italics="true" pageId="11" pageNumber="196">Rhynchobatus</emphasis>
</taxonomicName>
. Vernacular: Taiwanese Wedgefish.
</paragraph>
</subSubSection>
<subSubSection id="C4E665735D7BFF82FF14FE5CFD685ABA" pageId="11" pageNumber="196" type="discussion">
<paragraph id="8C4336F85D7BFF82FF14FE5CFBBF5EF2" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
<emphasis id="BE88EAEA5D7BFF82FF14FE5CFEAA5DAE" bold="true" box="[199,315,476,500]" pageId="11" pageNumber="196">Remarks.</emphasis>
The Taiwanese Wedgefish,
<taxonomicName id="4BFC4D7B5D7BFF82FDA7FE5CFC385DAE" box="[628,937,476,500]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FDA7FE5CFC385DAE" box="[628,937,476,500]" italics="true" pageId="11" pageNumber="196">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
, undergoes ontogenetic changes in fin shape that typify development in many other chondrichthyan fishes. The dorsal fins change shape to become taller and more erect as they develop (see
<figureCitation id="14C72A7D5D7BFF82FDD0FDA4FDF45E66" box="[515,613,548,573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1899,1921]" captionTargetBox="[767,1436,475,1874]" captionTargetId="figure@5.[767,1372,964,1460]" captionTargetPageId="5" captionText="FIGURE 4. Rhynchobatus immaculatus sp. nov. fin shape and development. Immature male holotype (NMMB P 16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMB P 15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin." httpUri="https://zenodo.org/record/219789/files/figure.png" pageId="11" pageNumber="196">Figure 4</figureCitation>
). The first dorsal fin becomes more pointed apically (
<figureCitation id="14C72A7D5D7BFF82FB15FDA4FAA25E66" box="[1222,1331,548,573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1899,1921]" captionTargetBox="[767,1436,475,1874]" captionTargetId="figure@5.[767,1372,964,1460]" captionTargetPageId="5" captionText="FIGURE 4. Rhynchobatus immaculatus sp. nov. fin shape and development. Immature male holotype (NMMB P 16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMB P 15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin." httpUri="https://zenodo.org/record/219789/files/figure.png" pageId="11" pageNumber="196">Figures 4</figureCitation>
a, d), and the caudal-fin lobes become relatively longer and narrower (
<figureCitation id="14C72A7D5D7BFF82FCE9FDC7FC365E3A" box="[826,935,583,608]" captionStart="FIGURE 4" captionStartId="5.[151,250,1899,1921]" captionTargetBox="[767,1436,475,1874]" captionTargetId="figure@5.[767,1372,964,1460]" captionTargetPageId="5" captionText="FIGURE 4. Rhynchobatus immaculatus sp. nov. fin shape and development. Immature male holotype (NMMB P 16274, 730 mm TL, recently preserved): A. Lateral view of first dorsal fin, C. Lateral view of second dorsal fin, E. Lateral view of caudal fin. Immature male non-type (NMMB P 15684, 992 mm TL, preserved): B. Lateral view of first dorsal fin, D. Lateral view of second dorsal fin, F. Lateral view of caudal fin." httpUri="https://zenodo.org/record/219789/files/figure.png" pageId="11" pageNumber="196">Figures 4</figureCitation>
c, f). These changes, which take place within a relatively narrow size range (immature males NMMBP16274 and NMMBP15684 are
<quantity id="4B049B1D5D7BFF82FB55FDECFB735EDE" box="[1158,1250,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="196" unit="mm" value="730.0">730 mm</quantity>
and
<quantity id="4B049B1D5D7BFF82FAC4FDECFAE55EDE" box="[1303,1396,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="9.92" pageId="11" pageNumber="196" unit="mm" value="992.0">992 mm</quantity>
TL respectively) in
<taxonomicName id="4BFC4D7B5D7BFF82FE98FD11FE685EF2" box="[331,505,656,680]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FE98FD11FE685EF2" box="[331,505,656,680]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
, are likely to be associated with onset of maturity.
</paragraph>
<paragraph id="8C4336F85D7BFF82FF14FD34FCF258FA" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
<taxonomicName id="4BFC4D7B5D7BFF82FF14FD34FE6A5E96" box="[199,507,692,716]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FF14FD34FE6A5E96" box="[199,507,692,716]" italics="true" pageId="11" pageNumber="196">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
differs from other wedgefishes occurring in the western North Pacific, primarily in vertebral count and coloration. It has a very high vertebral count matched only by
<taxonomicName id="4BFC4D7B5D7BFF82FB9EFD59FB715EAA" box="[1101,1248,728,752]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="djiddensis">
<emphasis id="BE88EAEA5D7BFF82FB9EFD59FB715EAA" box="[1101,1248,728,752]" italics="true" pageId="11" pageNumber="196">R. djiddensis</emphasis>
</taxonomicName>
from the Indian Ocean; total free centra of
<taxonomicName id="4BFC4D7B5D7BFF82FE02FD7DFD135F4E" box="[465,642,764,788]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FE02FD7DFD135F4E" box="[465,642,764,788]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
165170 vs. typically more than
<quantity id="4B049B1D5D7BFF82FBDFFD7CFBC75F4F" box="[1036,1110,764,789]" metricMagnitude="0" metricUnit="m" metricValue="4.4196" pageId="11" pageNumber="196" unit="in" value="174.0">174 in</quantity>
<taxonomicName id="4BFC4D7B5D7BFF82FBB3FD7DFB625F4E" box="[1120,1267,764,788]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="djiddensis">
<emphasis id="BE88EAEA5D7BFF82FBB3FD7DFB625F4E" box="[1120,1267,764,788]" italics="true" pageId="11" pageNumber="196">R. djiddensis</emphasis>
</taxonomicName>
. Another high count species
<taxonomicName id="4BFC4D7B5D7BFF82FEEFFCA1FE415F62" box="[316,464,800,824]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="australiae">
<emphasis id="BE88EAEA5D7BFF82FEEFFCA1FE415F62" box="[316,464,800,824]" italics="true" pageId="11" pageNumber="196">R. australiae</emphasis>
</taxonomicName>
, has been confirmed from
<collectingCountry id="F4EB76685D7BFF82FCC2FC9FFCF25F62" box="[785,867,799,824]" name="Taiwan" pageId="11" pageNumber="196">Taiwan</collectingCountry>
(HUMZ109480, female
<quantity id="4B049B1D5D7BFF82FB47FCA0FB645F6D" box="[1172,1269,800,824]" metricMagnitude="-1" metricUnit="m" metricValue="7.9" pageId="11" pageNumber="196" unit="mm" value="790.0">790 mm</quantity>
TL; total free centra 161) but, like
<emphasis id="BE88EAEA5D7BFF82FE43FCC5FDBD5F06" box="[400,556,836,860]" italics="true" pageId="11" pageNumber="196">
<taxonomicName id="4BFC4D7B5D7BFF82FE43FCC5FDB95F06" box="[400,552,836,860]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="djiddensis">R. djiddensis</taxonomicName>
,
</emphasis>
has a more complex colour pattern and a black pectoral marking in small individuals.
<taxonomicName id="4BFC4D7B5D7BFF82FEF6FCE8FDCF5FDA" box="[293,606,872,896]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FEF6FCE8FDCF5FDA" box="[293,606,872,896]" italics="true" pageId="11" pageNumber="196">Rhynchobatus immaculatus</emphasis>
</taxonomicName>
also appears to be a smaller wedgefish than either
<taxonomicName id="4BFC4D7B5D7BFF82FB11FCE9FAC65FDA" box="[1218,1367,872,896]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="australiae">
<emphasis id="BE88EAEA5D7BFF82FB11FCE9FAC65FDA" box="[1218,1367,872,896]" italics="true" pageId="11" pageNumber="196">R. australiae</emphasis>
</taxonomicName>
or
<taxonomicName id="4BFC4D7B5D7BFF82FA57FCE9FE985FFE" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="djiddensis">
<emphasis id="BE88EAEA5D7BFF82FA57FCE9FE985FFE" italics="true" pageId="11" pageNumber="196">R. djiddensis</emphasis>
</taxonomicName>
, which both attain about
<quantity id="4B049B1D5D7BFF82FDE7FC0CFD055FFE" box="[564,660,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="196" unit="mm" value="300.0">300 mm</quantity>
TL (Compagno &amp; Last, 1999) and
<quantity id="4B049B1D5D7BFF82FBE5FC0CFB075FFE" box="[1078,1174,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="196" unit="mm" value="310.0">310 mm</quantity>
TL (Compagno
<emphasis id="BE88EAEA5D7BFF82FA89FC0DFA045FFE" box="[1370,1429,908,932]" italics="true" pageId="11" pageNumber="196">et al.</emphasis>
, 1989) respectively. An undescribed species from the
<collectingCountry id="F4EB76685D7BFF82FCF1FC2FFCC45F92" box="[802,853,943,968]" name="India" pageId="11" pageNumber="196">Indo</collectingCountry>
Malay Archipelago,
<taxonomicName id="4BFC4D7B5D7BFF82FB89FC30FB685F92" box="[1114,1273,944,968]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="genus">
<emphasis id="BE88EAEA5D7BFF82FB89FC30FB685F92" box="[1114,1273,944,968]" italics="true" pageId="11" pageNumber="196">Rhynchobatus</emphasis>
</taxonomicName>
sp. 1 (sensu Compagno &amp; Last, 1999), which also has a dark, white-spotted body and lacks a dark pectoral marking, has a longer snout and much lower vertebral count (113116) than
<taxonomicName id="4BFC4D7B5D7BFF82FC89FC79FB9A584A" box="[858,1035,1016,1040]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FC89FC79FB9A584A" box="[858,1035,1016,1040]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
. All other species occurring in the region have a dark pectoral marking and lower vertebral counts:
<taxonomicName id="4BFC4D7B5D7BFF82FCB8FB9DFC60586E" box="[875,1009,1053,1076]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="springeri">
<emphasis id="BE88EAEA5D7BFF82FCB8FB9DFC60586E" box="[875,1009,1053,1076]" italics="true" pageId="11" pageNumber="196">R. springeri</emphasis>
</taxonomicName>
has 113126 free vertebral centra and
<taxonomicName id="4BFC4D7B5D7BFF82FF44FBC1FED15802" box="[151,320,1088,1112]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="palpebratus">
<emphasis id="BE88EAEA5D7BFF82FF44FBC1FED15802" box="[151,320,1088,1112]" italics="true" pageId="11" pageNumber="196">R. palpebratus</emphasis>
</taxonomicName>
has 135139 free vertebral centra (Compagno &amp; Last, 2008). Also, the first dorsal fin is more posteriorly positioned in
<taxonomicName id="4BFC4D7B5D7BFF82FE60FBE5FDF25826" box="[435,611,1124,1148]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FE60FBE5FDF25826" box="[435,611,1124,1148]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
(predorsal length 4849% TL vs. 4245% (mean 43%) TL and 4348% (mean 46%) TL in
<taxonomicName id="4BFC4D7B5D7BFF82FEBEFB09FE6358C5" box="[365,498,1160,1184]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="springeri">
<emphasis id="BE88EAEA5D7BFF82FEBEFB09FE6358C5" box="[365,498,1160,1184]" italics="true" pageId="11" pageNumber="196">R. springeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4BFC4D7B5D7BFF82FDF9FB09FD5E58FA" box="[554,719,1160,1184]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="palpebratus">
<emphasis id="BE88EAEA5D7BFF82FDF9FB09FD5E58FA" box="[554,719,1160,1184]" italics="true" pageId="11" pageNumber="196">R. palpebratus</emphasis>
</taxonomicName>
respectively.
</paragraph>
<paragraph id="8C4336F85D7BFF82FF14FB2CFD685ABA" blockId="11.[151,1436,151,1760]" pageId="11" pageNumber="196">
Compagno &amp; Last (1999) considered
<taxonomicName id="4BFC4D7B5D7BFF82FDACFB2CFC63589F" authority="Wang" authorityName="Wang" box="[639,1010,1196,1221]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="yentinensis">
<emphasis id="BE88EAEA5D7BFF82FDACFB2CFC35589E" box="[639,932,1196,1220]" italics="true" pageId="11" pageNumber="196">Rhynchobatus yentinensis</emphasis>
Wang
</taxonomicName>
, described from a specimen taken at nearby Wenzhou,
<collectingCountry id="F4EB76685D7BFF82FEB6FB4FFE3A58B2" box="[357,427,1231,1256]" name="China" pageId="11" pageNumber="196">China</collectingCountry>
, to be a possible synonym of
<taxonomicName id="4BFC4D7B5D7BFF82FCD6FB51FC1C58BD" box="[773,909,1232,1256]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="springeri">
<emphasis id="BE88EAEA5D7BFF82FCD6FB51FC1C58BD" box="[773,909,1232,1256]" italics="true" pageId="11" pageNumber="196">R. springeri</emphasis>
</taxonomicName>
(their former
<emphasis id="BE88EAEA5D7BFF82FBE2FB51FBD858B2" box="[1073,1097,1233,1256]" italics="true" pageId="11" pageNumber="196">R.</emphasis>
sp. 2), but later considered it more likely to be synonymous with
<taxonomicName id="4BFC4D7B5D7BFF82FDE8FB75FC4F5956" authority="Bloch &amp; Schneider, 1801" authorityName="Bloch &amp; Schneider" authorityYear="1801" box="[571,990,1268,1293]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="laevis">
<emphasis id="BE88EAEA5D7BFF82FDE8FB75FD0F5956" box="[571,670,1268,1292]" italics="true" pageId="11" pageNumber="196">R. laevis</emphasis>
(Bloch &amp; Schneider, 1801)
</taxonomicName>
(Compagno &amp; Last 2008; Eschmeyer, 2013). Wangs (1933) account of the male
<typeStatus id="5347885A5D7BFF82FD50FA97FD74596A" box="[643,741,1303,1328]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
(
<quantity id="4B049B1D5D7BFF82FD25FA98FCF55975" box="[758,868,1304,1328]" metricMagnitude="0" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="196" unit="mm" value="1010.0">1010 mm</quantity>
TL) gives an upper tooth row count of 27 (much lower the 52 rows in the smaller
<typeStatus id="5347885A5D7BFF82FDD5FABCFDF9590F" box="[518,616,1340,1365]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4BFC4D7B5D7BFF82FD5DFABDFC85590E" box="[654,788,1341,1364]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="springeri">
<emphasis id="BE88EAEA5D7BFF82FD5DFABDFC85590E" box="[654,788,1341,1364]" italics="true" pageId="11" pageNumber="196">R. springeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4BFC4D7B5D7BFF82FC99FABDFC7E590E" box="[842,1007,1340,1364]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="palpebratus">
<emphasis id="BE88EAEA5D7BFF82FC99FABDFC7E590E" box="[842,1007,1340,1364]" italics="true" pageId="11" pageNumber="196">R. palpebratus</emphasis>
</taxonomicName>
, or the 48 rows in a smaller
<typeStatus id="5347885A5D7BFF82FAFDFABDFACF590F" box="[1326,1374,1341,1365]" pageId="11" pageNumber="196">type</typeStatus>
of
<taxonomicName id="4BFC4D7B5D7BFF82FA57FABDFEB75922" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FA57FABDFEB75922" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
). The
<typeStatus id="5347885A5D7BFF82FEA6FADFFE465922" box="[373,471,1375,1400]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4BFC4D7B5D7BFF82FDD2FAE1FD0E592D" box="[513,671,1376,1400]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="yentinensis">
<emphasis id="BE88EAEA5D7BFF82FDD2FAE1FD0E592D" box="[513,671,1376,1400]" italics="true" pageId="11" pageNumber="196">R. yentinensis</emphasis>
</taxonomicName>
once existed as a mounted skin but is now missing. According to Prof. Han-Lin Wu (Shanghai Ocean University, pers comm. to Xiaoyu Kong), most
<typeStatus id="5347885A5D7BFF82FBBAFA05FB0859C7" box="[1129,1177,1413,1437]" pageId="11" pageNumber="196">type</typeStatus>
specimens in Chinese collections from the early 1900s were lost during the Second World War. However, based on Wangs description,
<taxonomicName id="4BFC4D7B5D7BFF82FA57FA29FE8559BE" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="yentinensis">
<emphasis id="BE88EAEA5D7BFF82FA57FA29FE8559BE" italics="true" pageId="11" pageNumber="196">R. yentinensis</emphasis>
</taxonomicName>
has a dark spot on the snout (vs. absent in
<taxonomicName id="4BFC4D7B5D7BFF82FCC7FA4DFC5459BE" box="[788,965,1484,1508]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FCC7FA4DFC5459BE" box="[788,965,1484,1508]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
), first dorsal-fin origin above or slightly behind the pelvic-fin origin (vs. first dorsal-fin origin well behind pelvic-fin origin), 2 spines in inner scapular rows on disc (vs. 69 spines), 2 series of white spots (vs. none) along the tail of males about
<quantity id="4B049B1D5D7BFF82FBBCF994FB525A76" box="[1135,1219,1556,1580]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="196" unit="cm" value="100.0">100 cm</quantity>
TL, and possibly a relatively shorter snout (preorbital length 2.4 times space vs. 2.62.9 times width of interorbital space). From the illustration and description of the
<typeStatus id="5347885A5D7BFF82FDF9F9DCFD1E5A2F" box="[554,655,1628,1653]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
,
<taxonomicName id="4BFC4D7B5D7BFF82FD4DF9DDFCAF5A2E" box="[670,830,1629,1652]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="yentinensis">
<emphasis id="BE88EAEA5D7BFF82FD4DF9DDFCAF5A2E" box="[670,830,1629,1652]" italics="true" pageId="11" pageNumber="196">R. yentinensis</emphasis>
</taxonomicName>
appears to be a broad-snouted species with a dark pectoral marking and low scapular spine count, suggesting that it is closer to either
<taxonomicName id="4BFC4D7B5D7BFF82FB90F901FB795AC2" box="[1091,1256,1664,1688]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="palpebratus">
<emphasis id="BE88EAEA5D7BFF82FB90F901FB795AC2" box="[1091,1256,1664,1688]" italics="true" pageId="11" pageNumber="196">R. palpebratus</emphasis>
</taxonomicName>
or
<taxonomicName id="4BFC4D7B5D7BFF82FADCF901FA045ACD" box="[1295,1429,1664,1688]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="springeri">
<emphasis id="BE88EAEA5D7BFF82FADCF901FA045ACD" box="[1295,1429,1664,1688]" italics="true" pageId="11" pageNumber="196">R. springeri</emphasis>
</taxonomicName>
, than to
<taxonomicName id="4BFC4D7B5D7BFF82FF22F925FE335AE6" box="[241,418,1700,1724]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="196" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="BE88EAEA5D7BFF82FF22F925FE335AE6" box="[241,418,1700,1724]" italics="true" pageId="11" pageNumber="196">R. immaculatus</emphasis>
</taxonomicName>
. Its placement will have to be decided using molecular analysis, if the
<typeStatus id="5347885A5D7BFF82FB30F924FAD45AE7" box="[1251,1349,1700,1725]" pageId="11" pageNumber="196" type="holotype">holotype</typeStatus>
is ever rediscovered and DNA can be extracted from the skin.
</paragraph>
</subSubSection>
</treatment>
</document>