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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.983.52630" ID-GBIF-Dataset="d3b857f7-25c1-4b21-9d39-4510ab675fca" ID-PMC="PMC7655787" ID-Pensoft-Pub="1313-2970-983-1" ID-Pensoft-UUID="C890C7FD4B2D57A8B1A062305ED42D53" ID-PubMed="33244289" ID-ZooBank="7DED4CE2934C4539945F758930C927F9" ModsDocID="1313-2970-983-1" checkinTime="1604475085656" checkinUser="pensoft" docAuthor="Smith, Sarah M., Beaver, Roger A. &amp; Cognato, Anthony I." docDate="2020" docId="8E8BA2F864366BFA349BDEEEEBE2F050" docLanguage="en" docName="ZooKeys 983: 1-442" docOrigin="ZooKeys 983" docSource="http://dx.doi.org/10.3897/zookeys.983.52630" docTitle="Coptodryas inornata Smith &amp; Beaver &amp; Cognato 2020, sp. nov." docType="treatment" docUuid="29DE1A5D-B4A5-4F5B-A798-45DE8356860A" docUuidSource="ZooBank" docVersion="4" id="C890C7FD4B2D57A8B1A062305ED42D53" lastPageNumber="1" masterDocId="C890C7FD4B2D57A8B1A062305ED42D53" masterDocTitle="A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China" masterLastPageNumber="442" masterPageNumber="1" pageNumber="1" updateTime="1668169715782" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beaver, Roger A.</mods:namePart>
<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>983</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>442</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.983.52630</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
<mods:identifier type="ZooBank">7DED4CE2934C4539945F758930C927F9</mods:identifier>
<mods:identifier type="Pensoft-UUID">C890C7FD4B2D57A8B1A062305ED42D53</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168754593" LSID="urn:lsid:zoobank.org:act:29DE1A5D-B4A5-4F5B-A798-45DE8356860A" httpUri="http://treatment.plazi.org/id/8E8BA2F864366BFA349BDEEEEBE2F050" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/29DE1A5D-B4A5-4F5B-A798-45DE8356860A" authority="Smith &amp; Beaver &amp; Cognato, 2020" authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Coptodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodryas inornata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="inornata" status="sp. nov.">Coptodryas inornata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Dorsal, lateral and declivital view of Coptodryas confusa, 1.8 - 2.2 mm (A, B, I), C. elegans syntype, 2.1 - 2.4 mm (C, D, J), C. inornata holotype, 3.1 - 3.2 mm (E, F, K), and C. mus, 2.3 - 2.4 mm (G, H, L)." figureDoi="10.3897/zookeys.983.52630.figure36" httpUri="https://binary.pensoft.net/fig/470836" pageId="0" pageNumber="1">Fig. 36E, F, K</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, female, Vietnam: Dong Nai, Cat Tien N.P.,
<geoCoordinate degrees="11.42854" direction="north" orientation="latitude" precision="1" value="11.42854">11.42854</geoCoordinate>
,
<geoCoordinate degrees="107.42544" direction="east" orientation="longitude" precision="1" value="107.42544">107.42544</geoCoordinate>
, 148 m, 23.ii.2017, VN98, A.I. Cognato, T.A. Hoang, ex 5 cm diameter branches (MSUC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
, female, as holotype (MSUC, 5; NHMUK, 5; NHMW, 5; NMNH, 5; VMNH, 5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
3.1-3.2 mm long (mean = 3.14 mm; n = 5); 2.38-2.46
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. This species is distinguished by the lack of a tuft of setae along the elytral base; declivity rounded; elytra shiny; striae and interstriae distinct; interstrial punctures confused; body lightly setose; antennal club as broad as tall; protibiae distinctly triangular; pronotum basic (type 0) when viewed laterally, basic (type 2) when viewed dorsally; and anterior margin of the pronotum without a row of serrations.
</paragraph>
<paragraph pageId="0" pageNumber="1">
This species strongly resembles
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" baseAuthorityName="Wood" baseAuthorityYear="1992" class="Insecta" family="Curculionidae" genus="Xylosandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xylosandrus formosae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="formosae">
<emphasis italics="true" pageId="0" pageNumber="1">Xylosandrus formosae</emphasis>
</taxonomicName>
which also lacks a distinct mycangial tuft (at the base of the pronotum) and both have triangular protibia.
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Coptodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodryas inornata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="inornata">
<emphasis italics="true" pageId="0" pageNumber="1">Coptodryas inornata</emphasis>
</taxonomicName>
is distinguished by the reduced scutellum, antennal club type 3 (as described for genus), and elytral base bisinuate and costate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="stat. res." baseAuthorityName="Schedl" baseAuthorityYear="1942" class="Insecta" family="Curculionidae" genus="Microperus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microperus fulvulus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fulvulus">
<emphasis italics="true" pageId="0" pageNumber="1">Microperus fulvulus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" baseAuthorityName="Wood" baseAuthorityYear="1992" class="Insecta" family="Curculionidae" genus="Xylosandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xylosandrus formosae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="formosae">
<emphasis italics="true" pageId="0" pageNumber="1">Xylosandrus formosae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(female).</emphasis>
3.1-3.2 mm long (mean = 3.14 mm; n = 5); 2.38-2.46
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Pronotum, head, antennae, and legs light brown, elytra darker brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, subshiny, punctate, punctures large, shallow, setose; punctures bearing a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum distinctly triangular, flat, flush with genae. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club longer than wide, flattened, type 4; segment 1 corneous, small, convex; segment 2 larger than segment 1, narrow, transverse, corneous; segments 1-3 present on posterior face.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
: 0.72
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. In dorsal view basic and parallel-sided, type 2, sides parallel in basal 2/3, rounded anteriorly; anterior margin without serrations. In lateral view basic, type 0, disc flat, summit at midpoint. Anterior slope with densely spaced, broad asperities, becoming lower and more strongly transverse towards summit. Disc shiny with dense minute punctures, densely setose, setae short erect hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base weakly bisinuate, posterior angles acutely rounded, almost subquadrate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytra</emphasis>
: 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.96
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum. Scutellum minute, convex, slightly raised above elytral surface. Elytral mycangium setal tuft of absent. Elytral base bisinuate, edge oblique, humeral angles rounded, parallel-sided in basal 2/3, then rounded to apex. Disc flat, shiny, striae not impressed, with small shallow punctures separated by 2-3 diameters of a puncture, setose, setae short, recumbent, hair-like; interstriae flat, minutely and confusedly punctate, setose, setae 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as strial setae, erect, hair-like. Declivity occupying approximately 2/5 of elytral length, rounded, face convex, strongly shiny; striae flat, setose, setae as described for disc, punctate, punctures similar in size to those of disc; interstriae 1-3 parallel, interstriae densely covered with long, erect hair-like setae; interstriae impunctate, densely and uniformly uniseriately granulate from base to apex, setose, setae as described for disc. Posterolateral margin distinctly carinate to interstriae 7.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
: procoxae contiguous; prosternal coxal piece short, inconspicuous. Protibiae distinctly triangular, broadest at apical 1/4; posterior face smooth; apical 1/2 of outer margin with six or seven large socketed denticles, their length longer than basal width. Meso- and metatibiae flattened; outer margins evenly rounded with 9-11 and eight large socketed denticles, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
L.
<emphasis italics="true" pageId="0" pageNumber="1">inornatus</emphasis>
= unadorned. In reference to the atypical unsculptured declivity. An adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="host">
<paragraph pageId="0" pageNumber="1">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>