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<document id="8D8FB6712F155F44C2DB592C185DCD55" ID-CLB-Dataset="28076" ID-DOI="10.11646/zootaxa.4504.3.5" ID-GBIF-Dataset="f0735593-cbcb-48b1-82ae-9e85178b2756" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606493" ID-ZooBank="B9A8121F-CCF9-4BC3-8CDF-CE6C28AFAC1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553584857517" checkinUser="plazi" docAuthor="Kang, Dong Won &amp; Kim, Hyung June" docDate="2018" docId="CB5787B7ED4F7872FF34FACAFBF3F89A" docLanguage="en" docName="zootaxa.4504.3.5.pdf" docOrigin="Zootaxa 4504 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clathria (Clathria) ieoensis Kang &amp; Kim 2018, sp. nov." docType="treatment" docUuid="00C9BB1F-AE09-455B-845D-357372776310" docUuidSource="ZooBank" docVersion="8" lastPageNumber="398" masterDocId="376EFFCFED49787AFFA3FFB3FFF2FF97" masterDocTitle="Two new species of the Genus Clathria (Demospongiae: Poecilosclerida: Microcionidae) from Jeju Island, Korea" masterLastPageNumber="400" masterPageNumber="390" pageNumber="396" updateTime="1698712781287" updateUser="ExternalLinkService">
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<mods:title id="E0061CB0CDC384CAAB0E68E660F242A3">Two new species of the Genus Clathria (Demospongiae: Poecilosclerida: Microcionidae) from Jeju Island, Korea</mods:title>
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<mods:namePart id="C403BD768DE91D651845134D8356A720">Kang, Dong Won</mods:namePart>
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<mods:namePart id="623E723EF6C251CA368424042B2247DC">Kim, Hyung June</mods:namePart>
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<mods:title id="930268BB442748E0EC5946651BFB9899">Zootaxa</mods:title>
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<mods:date id="98231DE94DF83551AF90D3C9D48206F2">2018</mods:date>
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<mods:number id="5A0B8C3C9911A73B7167FBB4A772915D">2018-10-25</mods:number>
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<mods:number id="C133A263B4C115E787D2910DF644E32B">4504</mods:number>
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<treatment id="CB5787B7ED4F7872FF34FACAFBF3F89A" ID-DOI="http://doi.org/10.5281/zenodo.5996663" ID-GBIF-Taxon="154495088" ID-Zenodo-Dep="5996663" LSID="urn:lsid:zoobank.org:act:00C9BB1F-AE09-455B-845D-357372776310" httpUri="http://treatment.plazi.org/id/CB5787B7ED4F7872FF34FACAFBF3F89A" lastPageId="8" lastPageNumber="398" pageId="6" pageNumber="396">
<subSubSection id="0BE4652AED4F787CFF34FACAFDBAFA03" box="[151,584,1401,1428]" pageId="6" pageNumber="396" type="nomenclature">
<paragraph id="434136A1ED4F787CFF34FACAFDBAFA03" blockId="6.[151,584,1401,1461]" box="[151,584,1401,1428]" pageId="6" pageNumber="396">
<heading id="180981CDED4F787CFF34FACAFDBAFA03" bold="true" box="[151,584,1401,1428]" fontSize="11" level="1" pageId="6" pageNumber="396" reason="1">
<emphasis id="718AEAB3ED4F787CFF34FACAFDBAFA03" bold="true" box="[151,584,1401,1428]" pageId="6" pageNumber="396">
<taxonomicName id="84FE4D22ED4F787CFF34FACAFE11FA04" ID-CoL="85K6M" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" box="[151,483,1401,1427]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="6" pageNumber="396" phylum="Porifera" rank="subGenus" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED4F787CFF34FACAFE11FA04" bold="true" box="[151,483,1401,1427]" italics="true" pageId="6" pageNumber="396">Clathria (Clathria) ieoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED4F787CFE49FAC9FDBAFA03" box="[490,584,1402,1428]" pageId="6" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="0BE4652AED4F787CFF34FA2FFE80FA22" box="[151,370,1436,1461]" pageId="6" pageNumber="396" type="description">
<paragraph id="434136A1ED4F787CFF34FA2FFE80FA22" blockId="6.[151,584,1401,1461]" box="[151,370,1436,1461]" pageId="6" pageNumber="396">
(
<figureCitation id="DBC52A24ED4F787CFF3CFA2FFEF8FA22" box="[159,266,1436,1461]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1888,1910]" captionStartId-1="8.[151,250,1580,1602]" captionTargetBox-0="[362,1225,536,1843]" captionTargetBox-1="[353,1230,208,1538]" captionTargetId-0="figure@7.[340,1247,517,1867]" captionTargetId-1="figure@8.[340,1247,193,1559]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." captionText-1="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri-0="https://zenodo.org/record/2606501/files/figure.png" httpUri-1="https://zenodo.org/record/2606503/files/figure.png" pageId="6" pageNumber="396">Figs. 56</figureCitation>
,
<tableCitation id="0E7C031AED4F787CFEB5FA2FFE9BFA22" box="[278,361,1436,1461]" captionStart="TABLE 3" captionStartId="9.[151,239,152,175]" captionText="TABLE 3. Comparison of characters between Clathria (C.) ieoenssis sp. nov. and C. (C.) murphyi Hooper, 1996." pageId="6" pageNumber="396">Table 3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BE4652AED4F787CFF34FA57FD17F9B6" pageId="6" pageNumber="396" type="materials_examined">
<paragraph id="434136A1ED4F787CFF34FA57FD17F9B6" blockId="6.[151,1437,1508,2037]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF34FA57FE8BFA6A" bold="true" box="[151,377,1508,1533]" pageId="6" pageNumber="396">Material examined</emphasis>
..
<materialsCitation id="F3963CFCED4F787CFE34FA57FD17F9B6" ID-GBIF-Occurrence="2028137926" collectingDate="2017-06-30" collectionCode="SCUBA, MABIK" collectorName="Sang-Hui Lee" country="South Korea" county="Island" elevation="15" location="Gapa-up" municipality="Seogwipo-si" pageId="6" pageNumber="396" specimenCode="IV001641547" specimenCount="1" stateProvince="Jeju" typeStatus="holotype">
<emphasis id="718AEAB3ED4F787CFE34FA57FDF9FA6A" bold="true" box="[407,523,1508,1533]" pageId="6" pageNumber="396">
<typeStatus id="9C458803ED4F787CFE34FA57FDF4FA6A" box="[407,518,1508,1533]" pageId="6" pageNumber="396" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="3BE97631ED4F787CFDB1FA56FDABFA6B" box="[530,601,1509,1532]" name="South Korea" pageId="6" pageNumber="396">Korea</collectingCountry>
,
<collectingRegion id="813AF843ED4F787CFDC6FA57FD45FA6A" box="[613,695,1508,1533]" country="South Korea" name="Jeju" pageId="6" pageNumber="396">Jeju-do</collectingRegion>
<collectingCounty id="AA204E2DED4F787CFD1CFA57FCF4FA6A" box="[703,774,1508,1533]" pageId="6" pageNumber="396">Island</collectingCounty>
,
<collectingMunicipality id="A325ACDBED4F787CFCB1FA57FC53FA6A" box="[786,929,1508,1533]" pageId="6" pageNumber="396">Seogwipo-si</collectingMunicipality>
,
<location id="4621607AED4F787CFC0EFA56FBE2FA6A" LSID="urn:lsid:plazi:treatment:CB5787B7ED4F7872FF34FACAFBF3F89A:4621607AED4F787CFC0EFA56FBE2FA6A" box="[941,1040,1509,1533]" country="South Korea" county="Island" municipality="Seogwipo-si" name="Gapa-up" pageId="6" pageNumber="396" stateProvince="Jeju">Gapa-up</location>
,
<location id="4621607AED4F787CFBBFFA57FBADFA6A" LSID="urn:lsid:plazi:treatment:CB5787B7ED4F7872FF34FACAFBF3F89A:4621607AED4F787CFBBFFA57FBADFA6A" box="[1052,1119,1508,1533]" country="South Korea" county="Island" municipality="Seogwipo-si" name="Ieodo" pageId="6" pageNumber="396" stateProvince="Jeju">Ieodo</location>
,
<elevation id="C8D3D192ED4F787CFBC9FA57FB56FA6B" box="[1130,1188,1508,1533]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="396" unit="m" value="15.0">
<quantity id="84069B44ED4F787CFBC9FA57FB56FA6B" box="[1130,1188,1508,1533]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="396" unit="m" value="15.0">15 m</quantity>
</elevation>
depth,
<collectingDate id="2704E989ED4F787CFB5BFA56FA6AFA6A" box="[1272,1432,1509,1533]" pageId="6" pageNumber="396" value="2017-06-30">
<date id="37401061ED4F787CFB5BFA56FA6AFA6A" box="[1272,1432,1509,1533]" pageId="6" pageNumber="396" value="2017-06-30">June 30, 2017</date>
</collectingDate>
,
<collectionCode id="25EFAE64ED4F787CFF34F9BAFF05F9B6" box="[151,247,1545,1569]" pageId="6" pageNumber="396">SCUBA</collectionCode>
, coll.
<collectorName id="EE0B5377ED4F787CFE99F9BBFE28F9B7" box="[314,474,1544,1569]" pageId="6" pageNumber="396">Sang-Hui Lee</collectorName>
,
<collectionCode id="25EFAE64ED4F787CFE46F9BAFDB0F9B6" box="[485,578,1545,1569]" pageId="6" pageNumber="396">MABIK</collectionCode>
<specimenCode id="13589EDAED4F787CFDEAF9BBFD17F9B6" box="[585,741,1544,1569]" collectionCode="IV" pageId="6" pageNumber="396">IV001641547</specimenCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0BE4652AED4F787DFF64F99FFD8CFE8B" lastPageId="7" lastPageNumber="397" pageId="6" pageNumber="396" type="description">
<paragraph id="434136A1ED4F787CFF64F99FFF1DF9FE" blockId="6.[151,1437,1508,2037]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F99FFEA3F9D2" bold="true" box="[199,337,1580,1605]" pageId="6" pageNumber="396">Description</emphasis>
.
<emphasis id="718AEAB3ED4F787CFEC3F99FFE57F9D2" box="[352,421,1580,1605]" italics="true" pageId="6" pageNumber="396">Shape</emphasis>
(
<figureCitation id="DBC52A24ED4F787CFE16F99FFDE4F9D2" box="[437,534,1580,1605]" captionStart="FIGURE 5" captionStartId="7.[151,250,1888,1910]" captionTargetBox="[362,1225,536,1843]" captionTargetId="figure@7.[340,1247,517,1867]" captionTargetPageId="7" captionText="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="6" pageNumber="396">Fig. 5 A</figureCitation>
): Encrusting,
<quantity id="84069B44ED4F787CFD1EF99EFD07F9D3" box="[701,757,1581,1605]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="396" unit="cm" value="2.0">2 cm</quantity>
in thickness and
<quantity id="84069B44ED4F787CFC18F99EFC01F9D3" box="[955,1011,1581,1604]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="396" unit="cm" value="7.0">7 cm</quantity>
in width, with small oscules flush on surface.
</paragraph>
<paragraph id="434136A1ED4F787CFF64F9C5FDDDF91A" blockId="6.[151,1437,1508,2037]" box="[199,559,1652,1677]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F9C5FEE5F91A" box="[199,279,1654,1677]" italics="true" pageId="6" pageNumber="396">Texture</emphasis>
: Soft and compressible.
</paragraph>
<paragraph id="434136A1ED4F787CFF64F92BFCACF926" blockId="6.[151,1437,1508,2037]" box="[199,862,1688,1713]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F92BFED2F926" box="[199,288,1688,1713]" italics="true" pageId="6" pageNumber="396">Oscules</emphasis>
:
<quantity id="84069B44ED4F787CFE96F92BFE4EF927" box="[309,444,1688,1713]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="6" pageNumber="396" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 mm</quantity>
in diameter, scattered on the surface.
</paragraph>
<paragraph id="434136A1ED4F787CFF64F90FFC10F942" blockId="6.[151,1437,1508,2037]" box="[199,994,1724,1749]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F90FFEE5F942" box="[199,279,1724,1749]" italics="true" pageId="6" pageNumber="396">Colour</emphasis>
: Red in life which gradually changes to dark brown in ethanol.
</paragraph>
<paragraph id="434136A1ED4F787CFF64F952FE0EF96F" blockId="6.[151,1437,1508,2037]" box="[199,508,1760,1785]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F952FEEFF96E" box="[199,285,1761,1785]" italics="true" pageId="6" pageNumber="396">Surface</emphasis>
: Smooth and even.
</paragraph>
<paragraph id="434136A1ED4F787CFF64F8B7FB1DF81E" blockId="6.[151,1437,1508,2037]" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F8B7FED4F88A" box="[199,294,1796,1821]" italics="true" pageId="6" pageNumber="396">Skeleton</emphasis>
(
<figureCitation id="DBC52A24ED4F787CFE96F8B7FE85F88A" box="[309,375,1796,1821]" captionStart="FIGURE 5" captionStartId="7.[151,250,1888,1910]" captionTargetBox="[362,1225,536,1843]" captionTargetId="figure@7.[340,1247,517,1867]" captionTargetPageId="7" captionText="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="6" pageNumber="396">Fig. 5</figureCitation>
BD): Ectosome membranous, hispid from connected choanosomal spicules tracts. Ectosomal skeleton with tangential layers of auxiliary styles in multispicular tracts. Choanosomal skeleton with regularly developed primary spongin fibres and plumose acanthostyles. Primary spongin fibre diameter about 3060?. Acanthostyles echinating the primary spongin fibres. Secondary spongin fibres not well developed.
</paragraph>
<paragraph id="434136A1ED4F787DFF64F827FD8CFE8B" blockId="6.[151,1437,1508,2037]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="397" pageId="6" pageNumber="396">
<emphasis id="718AEAB3ED4F787CFF64F827FED6F83A" box="[199,292,1940,1965]" italics="true" pageId="6" pageNumber="396">Spicules</emphasis>
(
<figureCitation id="DBC52A24ED4F787CFE8EF827FE83F83A" box="[301,369,1940,1965]" captionStart="FIGURE 5" captionStartId="7.[151,250,1888,1910]" captionTargetBox="[362,1225,536,1843]" captionTargetId="figure@7.[340,1247,517,1867]" captionTargetPageId="7" captionText="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="6" pageNumber="396">Fig. 5</figureCitation>
EH,
<figureCitation id="DBC52A24ED4F787CFE1DF827FDF6F83A" box="[446,516,1940,1965]" captionStart="FIGURE 6" captionStartId="8.[151,250,1580,1602]" captionTargetBox="[353,1230,208,1538]" captionTargetId="figure@8.[340,1247,193,1559]" captionTargetPageId="8" captionText="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="6" pageNumber="396">Fig. 6</figureCitation>
,
<tableCitation id="0E7C031AED4F787CFDB1F827FD9CF83A" box="[530,622,1940,1965]" captionStart="TABLE 3" captionStartId="9.[151,239,152,175]" captionText="TABLE 3. Comparison of characters between Clathria (C.) ieoenssis sp. nov. and C. (C.) murphyi Hooper, 1996." pageId="6" pageNumber="396">Table. 3</tableCitation>
):
<emphasis id="718AEAB3ED4F787CFD31F827FCD4F83A" box="[658,806,1940,1965]" italics="true" pageId="6" pageNumber="396">Megascleres.</emphasis>
Two sizes of style(
<figureCitation id="DBC52A24ED4F787CFBABF827FBB9F83A" box="[1032,1099,1940,1965]" captionStart="FIGURE 5" captionStartId="7.[151,250,1888,1910]" captionTargetBox="[362,1225,536,1843]" captionTargetId="figure@7.[340,1247,517,1867]" captionTargetPageId="7" captionText="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="6" pageNumber="396">Fig. 5</figureCitation>
EF), large and small styles, long, robust, straight or slightly curved with microspined head. Thin styles(
<figureCitation id="DBC52A24ED4F787CFBA3F80BFBC8F846" box="[1024,1082,1976,2001]" captionStart="FIGURE 6" captionStartId="8.[151,250,1580,1602]" captionTargetBox="[353,1230,208,1538]" captionTargetId="figure@8.[340,1247,193,1559]" captionTargetPageId="8" captionText="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="6" pageNumber="396">Fig.6</figureCitation>
AB), straight, long, smooth, with microspined base. Two sizes of acanthostyles(
<figureCitation id="DBC52A24ED4F787CFD79F86FFCEBF862" box="[730,793,2012,2037]" captionStart="FIGURE 5" captionStartId="7.[151,250,1888,1910]" captionTargetBox="[362,1225,536,1843]" captionTargetId="figure@7.[340,1247,517,1867]" captionTargetPageId="7" captionText="FIGURE 5. Clathria (Clathria) ieoensis sp. nov.: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle." httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="6" pageNumber="396">Fig. 5</figureCitation>
GH), large acanthostyles, long, robust, sharply pointed, evenly spined. Small acanthostyles, straight, with heavily spined bases.
<emphasis id="718AEAB3ED4E787DFBA3FF24FB62FF27" box="[1024,1168,151,176]" italics="true" pageId="7" pageNumber="397">Microscleres</emphasis>
. One size of palmate isochelae(
<figureCitation id="DBC52A24ED4E787DFEA8FF0FFEB8FF42" box="[267,330,188,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1580,1602]" captionTargetBox="[353,1230,208,1538]" captionTargetId="figure@8.[340,1247,193,1559]" captionTargetPageId="8" captionText="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="7" pageNumber="397">Fig. 6</figureCitation>
CD), small, weakly silicified, unmodified. Toxa(
<figureCitation id="DBC52A24ED4E787DFC20FF0FFC30FF42" box="[899,962,188,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1580,1602]" captionTargetBox="[353,1230,208,1538]" captionTargetId="figure@8.[340,1247,193,1559]" captionTargetPageId="8" captionText="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="7" pageNumber="397">Fig. 6</figureCitation>
EG), two size categories of wing shaped toxas, with rounded central curve and slightly reflexed arms. Accolada toxa(
<figureCitation id="DBC52A24ED4E787DFBB5FF6CFB89FF6F" box="[1046,1147,223,248]" captionStart="FIGURE 6" captionStartId="8.[151,250,1580,1602]" captionTargetBox="[353,1230,208,1538]" captionTargetId="figure@8.[340,1247,193,1559]" captionTargetPageId="8" captionText="FIGURE 6. Clathria (Clathria) ieoensis sp. nov.: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa." httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="7" pageNumber="397">Fig. 6 H</figureCitation>
), rare, thin, with sharply angular central curvature and straight arms.
</paragraph>
</subSubSection>
<subSubSection id="0BE4652AED4E787DFF64FE94FCB2FEA8" box="[199,832,295,320]" pageId="7" pageNumber="397" type="distribution">
<paragraph id="434136A1ED4E787DFF64FE94FCB2FEA8" blockId="7.[151,1436,151,464]" box="[199,832,295,320]" pageId="7" pageNumber="397">
<emphasis id="718AEAB3ED4E787DFF64FE94FEABFED7" bold="true" box="[199,345,295,320]" pageId="7" pageNumber="397">Distribution</emphasis>
. Only found in Ieodo at a depth of
<quantity id="84069B44ED4E787DFD47FE94FCCEFEA8" box="[740,828,295,320]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="7" pageNumber="397" unit="m" value="13.0" valueMax="20.0" valueMin="6.0">
6
<specimenCount id="55F8FD28ED4E787DFCA3FE94FCCEFEA8" box="[768,828,295,320]" pageId="7" pageNumber="397" type="generic">20 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE4652AED4E787DFF64FEFFFBA1FEF3" box="[199,1107,332,357]" pageId="7" pageNumber="397" type="etymology">
<paragraph id="434136A1ED4E787DFF64FEFFFBA1FEF3" blockId="7.[151,1436,151,464]" box="[199,1107,332,357]" pageId="7" pageNumber="397">
<emphasis id="718AEAB3ED4E787DFF64FEFFFEB4FEF2" bold="true" box="[199,326,332,357]" pageId="7" pageNumber="397">Etymology</emphasis>
. Named after the
<typeStatus id="9C458803ED4E787DFDACFEFEFDCDFEF2" box="[527,575,333,357]" pageId="7" pageNumber="397">type</typeStatus>
locality, Ieodo Ocean Research Station,
<collectingCountry id="3BE97631ED4E787DFBA4FEFEFBBDFEF3" box="[1031,1103,333,356]" name="South Korea" pageId="7" pageNumber="397">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0BE4652AED4E787DFF64FEDCFE87FE58" pageId="7" pageNumber="397" type="discussion">
<paragraph id="434136A1ED4E787DFF64FEDCFE87FE58" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="397">
<emphasis id="718AEAB3ED4E787DFF64FEDCFEC9FE1F" bold="true" box="[199,315,367,392]" pageId="7" pageNumber="397">Remarks.</emphasis>
<taxonomicName id="84FE4D22ED4E787DFEE1FEDCFD88FE1F" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" box="[322,634,367,392]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="397" phylum="Porifera" rank="subGenus" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED4E787DFEE1FEDCFE53FE1F" box="[322,417,367,392]" italics="true" pageId="7" pageNumber="397">Clathria</emphasis>
(
<emphasis id="718AEAB3ED4E787DFE12FEDCFDE2FE1F" box="[433,528,367,392]" italics="true" pageId="7" pageNumber="397">Clathria</emphasis>
)
<emphasis id="718AEAB3ED4E787DFD83FEC2FD88FE1F" box="[544,634,369,392]" italics="true" pageId="7" pageNumber="397">ieoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED4E787DFD22FEDCFD29FE1F" box="[641,731,367,392]" pageId="7" pageNumber="397" rank="species">
<emphasis id="718AEAB3ED4E787DFD22FEDCFD29FE1F" bold="true" box="[641,731,367,392]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
is allocated in the subgenus by having three sizes of styles and two sizes of echinating acanthostyles megascleres, with palmate isochelae, two categories of wing shape toxas and rare accolada toxas.
</paragraph>
</subSubSection>
<caption id="17816629ED4E787DFF34F8D3FA89F802" httpUri="https://zenodo.org/record/2606501/files/figure.png" pageId="7" pageNumber="397" startId="7.[151,250,1888,1910]" targetBox="[362,1225,536,1843]" targetPageId="7">
<paragraph id="434136A1ED4E787DFF34F8D3FA89F802" blockId="7.[151,1435,1888,1941]" pageId="7" pageNumber="397">
<emphasis id="718AEAB3ED4E787DFF34F8D3FEE5F8E2" bold="true" box="[151,279,1888,1910]" pageId="7" pageNumber="397">FIGURE 5.</emphasis>
<taxonomicName id="84FE4D22ED4E787DFE81F8D3FDB4F8E1" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" box="[290,582,1888,1910]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="397" phylum="Porifera" rank="subGenus" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED4E787DFE81F8D3FE8AF8E2" box="[290,376,1888,1909]" italics="true" pageId="7" pageNumber="397">Clathria</emphasis>
(
<emphasis id="718AEAB3ED4E787DFE28F8D3FE13F8E2" box="[395,481,1888,1909]" italics="true" pageId="7" pageNumber="397">Clathria</emphasis>
)
<emphasis id="718AEAB3ED4E787DFE57F8D2FDB4F8E1" box="[500,582,1889,1910]" italics="true" pageId="7" pageNumber="397">ieoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED4E787DFDF3F8D3FD57F8E2" box="[592,677,1888,1909]" pageId="7" pageNumber="397" rank="species">
<emphasis id="718AEAB3ED4E787DFDF3F8D3FD57F8E2" bold="true" box="[592,677,1888,1909]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
: (A) Image of the holotype specimen, (B) Ectosomal skeleton, (C-D) Choanosomal skeleton, (E) a. Large style, b. Small style, (F) Head of style, (G) Large acanthostyle, (H) Small acanthostyle.
</paragraph>
</caption>
<caption id="17816629ED417872FF34F99FFB5BF9F6" httpUri="https://zenodo.org/record/2606503/files/figure.png" pageId="8" pageNumber="398" startId="8.[151,250,1580,1602]" targetBox="[353,1230,208,1538]" targetPageId="8">
<paragraph id="434136A1ED417872FF34F99FFB5BF9F6" blockId="8.[151,1436,1580,1633]" pageId="8" pageNumber="398">
<emphasis id="718AEAB3ED417872FF34F99FFEE1F9D6" bold="true" box="[151,275,1580,1602]" pageId="8" pageNumber="398">FIGURE 6.</emphasis>
<taxonomicName id="84FE4D22ED417872FEB9F99FFDC5F9D5" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" box="[282,567,1580,1602]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="398" phylum="Porifera" rank="subGenus" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED417872FEB9F99FFE82F9D6" box="[282,368,1580,1601]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
(
<emphasis id="718AEAB3ED417872FE23F99FFE24F9D6" box="[384,470,1580,1601]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
)
<emphasis id="718AEAB3ED417872FE46F99EFDC5F9D5" box="[485,567,1581,1602]" italics="true" pageId="8" pageNumber="398">ieoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED417872FD9DF99EFD7DF9D5" box="[574,655,1581,1602]" pageId="8" pageNumber="398" rank="species">
<emphasis id="718AEAB3ED417872FD9DF99EFD7DF9D5" bold="true" box="[574,655,1581,1602]" pageId="8" pageNumber="398">sp. nov.</emphasis>
</taxonomicNameLabel>
: (A) Thin style, (B) Head of thin style, (C) Front view of palmate isochela, (D) Side view of palmate isochela, (E) Large toxa, (F) Small toxa, (G) End of toxa, (H) Accolada toxa.
</paragraph>
</caption>
<subSubSection id="0BE4652AED417872FF64F93BFBF3F89A" pageId="8" pageNumber="398" type="reference_group">
<paragraph id="434136A1ED417872FF64F93BFBF3F89A" blockId="8.[151,1436,1672,1805]" pageId="8" pageNumber="398">
<taxonomicName id="84FE4D22ED417872FF64F93BFDFAF937" ID-CoL="85K6M" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" box="[199,520,1672,1697]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="398" phylum="Porifera" rank="subGenus" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED417872FF64F93BFED4F936" box="[199,294,1672,1697]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
(
<emphasis id="718AEAB3ED417872FE99F93BFE6BF936" box="[314,409,1672,1697]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
)
<emphasis id="718AEAB3ED417872FE0DF93AFDFAF937" box="[430,520,1673,1696]" italics="true" pageId="8" pageNumber="398">ieoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED417872FDB7F93BFD83F937" box="[532,625,1672,1697]" pageId="8" pageNumber="398" rank="species">
<emphasis id="718AEAB3ED417872FDB7F93BFDDCF936" bold="true" box="[532,558,1672,1697]" pageId="8" pageNumber="398">sp</emphasis>
.
<emphasis id="718AEAB3ED417872FDE2F93BFD99F936" bold="true" box="[577,619,1672,1697]" pageId="8" pageNumber="398">nov</emphasis>
.
</taxonomicNameLabel>
differs from all other
<taxonomicName id="84FE4D22ED417872FCDCF93BFC2CF936" ID-CoL="84MDX" authorityName="Schmidt" authorityYear="1862" box="[895,990,1672,1697]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="398" phylum="Porifera" rank="genus">
<emphasis id="718AEAB3ED417872FCDCF93BFC2CF936" box="[895,990,1672,1697]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
</taxonomicName>
species from
<collectingCountry id="3BE97631ED417872FB24F93AFB39F937" box="[1159,1227,1673,1696]" name="South Korea" pageId="8" pageNumber="398">Korea</collectingCountry>
,
<collectingCountry id="3BE97631ED417872FB49F93AFAD8F937" box="[1258,1322,1673,1696]" name="Japan" pageId="8" pageNumber="398">Japan</collectingCountry>
based on spicule composition. Compared to Australian
<taxonomicName id="84FE4D22ED417872FD13F91FFCFDF952" ID-CoL="84MDX" authorityName="Schmidt" authorityYear="1862" box="[688,783,1708,1733]" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="398" phylum="Porifera" rank="genus">
<emphasis id="718AEAB3ED417872FD13F91FFCFDF952" box="[688,783,1708,1733]" italics="true" pageId="8" pageNumber="398">Clathria</emphasis>
</taxonomicName>
species (see
<bibRefCitation id="276F4B50ED417872FC13F91EFBB9F952" author="Hooper, J. N. A." box="[944,1099,1708,1733]" pageId="8" pageNumber="398" pagination="1 - 626" refId="ref2446" refString="Hooper, J. N. A. (1996) Revision of Microcionidae (Porifera: Poecilosclerida: Demospongiae), with description of Australian species. Memoirs of the Queensland Museum, 40, 1 - 626." type="journal article" year="1996">Hooper 1996</bibRefCitation>
), the most similar is
<taxonomicName id="84FE4D22ED417872FAE9F91EFE66F97E" ID-CoL="VS66" authority="Hooper, 1996" authorityName="Hooper" authorityYear="1996" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="399" phylum="Porifera" rank="species" species="murphyi" subGenus="Clathria">
<emphasis id="718AEAB3ED417872FAE9F91EFAAFF953" box="[1354,1373,1709,1732]" italics="true" pageId="8" pageNumber="398">C</emphasis>
. (
<emphasis id="718AEAB3ED417872FADAF91EFA7EF953" box="[1401,1420,1709,1732]" italics="true" pageId="8" pageNumber="398">C</emphasis>
.)
<emphasis id="718AEAB3ED417872FF34F963FF06F97E" box="[151,244,1744,1769]" italics="true" pageId="8" pageNumber="398">murphyi</emphasis>
<bibRefCitation id="276F4B50ED417872FF58F962FE66F97E" author="Hooper, J. N. A." box="[251,404,1744,1769]" pageId="8" pageNumber="398" pagination="1 - 626" refId="ref2446" refString="Hooper, J. N. A. (1996) Revision of Microcionidae (Porifera: Poecilosclerida: Demospongiae), with description of Australian species. Memoirs of the Queensland Museum, 40, 1 - 626." type="journal article" year="1996">
Hooper,
<quantity id="84069B44ED417872FEFFF963FE66F97E" box="[348,404,1744,1769]" metricMagnitude="1" metricUnit="m" metricValue="5.06984" pageId="8" pageNumber="398" unit="in" value="1996.0">1996</quantity>
</bibRefCitation>
</taxonomicName>
in shape, color, microsclere spicule composition (palmate isochelae, accolada toxas), but
<taxonomicName id="84FE4D22ED417872FA21F962FED6F89A" ID-CoL="VS5D" authority="Kang &amp; Kim, 2018" authorityName="Kang &amp; Kim" authorityYear="2018" class="Demospongiae" family="Microcionidae" genus="Clathria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="8" pageNumber="399" phylum="Porifera" rank="species" species="ieoensis" status="sp. nov." subGenus="Clathria">
<emphasis id="718AEAB3ED417872FA21F962FA6EF97F" box="[1410,1436,1745,1768]" italics="true" pageId="8" pageNumber="398">C.</emphasis>
(
<emphasis id="718AEAB3ED417872FF03F946FF41F89B" box="[160,179,1781,1804]" italics="true" pageId="8" pageNumber="398">C</emphasis>
.) ieoensis
</taxonomicName>
<taxonomicNameLabel id="6AB957C8ED417872FE88F947FE76F89B" box="[299,388,1780,1805]" pageId="8" pageNumber="398" rank="species">
<emphasis id="718AEAB3ED417872FE88F947FEB7F89A" bold="true" box="[299,325,1780,1805]" pageId="8" pageNumber="398">sp</emphasis>
.
<emphasis id="718AEAB3ED417872FEF7F947FE8CF89A" bold="true" box="[340,382,1780,1805]" pageId="8" pageNumber="398">nov</emphasis>
.
</taxonomicNameLabel>
differs in its megasclere spicule
<typeStatus id="9C458803ED417872FD56F946FCD7F89A" box="[757,805,1781,1805]" pageId="8" pageNumber="398">type</typeStatus>
and size (
<tableCitation id="0E7C031AED417872FC35F947FC01F89A" box="[918,1011,1780,1805]" captionStart="TABLE 3" captionStartId="9.[151,239,152,175]" captionText="TABLE 3. Comparison of characters between Clathria (C.) ieoenssis sp. nov. and C. (C.) murphyi Hooper, 1996." pageId="8" pageNumber="398">Table. 3</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>