294 lines
36 KiB
XML
294 lines
36 KiB
XML
<document id="DD526CC8911774BE3CA1B5DBFF248B76" ID-CLB-Dataset="32093" ID-DOI="10.11646/zootaxa.4320.2.1" ID-GBIF-Dataset="f3a915c9-a35b-454a-9090-c8792904d8d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="891714" ID-ZooBank="F3E61Ad4-9740-4B21-8Aea-619Abaae3580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505461295665" checkinUser="plazi" docAuthor="Leo, Piero" docDate="2017" docId="CB1787FD3D1DFF43CCD9FE745CC844A3" docLanguage="en" docName="zootaxa.4320.2.1.pdf.imf" docOrigin="Zootaxa 4320 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplia (Hoplia) messapia Uliana, Liberto" docType="treatment" docVersion="7" lastPageNumber="211" masterDocId="372EFF853D14FF49CC4EFFE05D10450F" masterDocTitle="On the taxonomy of the Italian endemic Hoplia dubia (Rossi, 1792), with the description of two new species (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="224" masterPageNumber="201" pageNumber="210" updateTime="1698468510115" updateUser="ExternalLinkService">
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<mods:title id="595B52FE94CC54DFAEF8FEE91FBEAF80">On the taxonomy of the Italian endemic Hoplia dubia (Rossi, 1792), with the description of two new species (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:name id="8F11285DB560C60E725FE229C6275740" type="personal">
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<mods:namePart id="584DE72FA4FD2F0B2051EA9785D25384">Leo, Piero</mods:namePart>
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<mods:title id="F143CD429A7912FDD7A65602C52CAC3D">Zootaxa</mods:title>
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<mods:part id="38A6A7264832ECA8B184679E84BEDC98">
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<mods:date id="7546992D61CCFEC3FFDF1348BF0D5EB7">2017</mods:date>
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<mods:number id="E2FE0C18CB9790973333B5E1214DA079">2017-09-15</mods:number>
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<mods:detail id="7B2EB572D5A9B1F30B179C13F397BD53" type="volume">
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<mods:number id="09D6B4BE51A41AFF774A573C8F897B81">4320</mods:number>
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<mods:identifier id="5806957EBE826BA93B4486CEBA83C447" type="CLB-Dataset">32093</mods:identifier>
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<mods:identifier id="A003BF8EE9E0B4FEDD216494A6C67468" type="DOI">10.11646/zootaxa.4320.2.1</mods:identifier>
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<treatment id="CB1787FD3D1DFF43CCD9FE745CC844A3" ID-DOI="http://doi.org/10.5281/zenodo.6022138" ID-GBIF-Taxon="134089777" ID-Zenodo-Dep="6022138" LSID="urn:lsid:plazi:treatment:CB1787FD3D1DFF43CCD9FE745CC844A3" httpUri="http://treatment.plazi.org/id/CB1787FD3D1DFF43CCD9FE745CC844A3" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210">
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<subSubSection id="0BA465603D1DFF40CCD9FE745F5544C0" pageId="9" pageNumber="210" type="nomenclature">
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<paragraph id="430136EB3D1DFF40CCD9FE745EB844A1" blockId="9.[151,936,404,463]" box="[151,936,404,430]" pageId="9" pageNumber="210">
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<heading id="184981873D1DFF40CCD9FE745EB844A1" bold="true" box="[151,936,404,430]" fontSize="11" level="1" pageId="9" pageNumber="210" reason="1">
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<emphasis id="71CAEAF93D1DFF40CCD9FE745EB844A1" bold="true" box="[151,936,404,430]" pageId="9" pageNumber="210">
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<taxonomicName id="84BE4D683D1DFF40CCD9FE745F8744A1" authority="Uliana, Liberto" authorityName="Uliana, Liberto" box="[151,663,404,430]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="messapia" subGenus="Hoplia">
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<emphasis id="71CAEAF93D1DFF40CCD9FE745DFA44A1" bold="true" box="[151,234,404,430]" italics="true" pageId="9" pageNumber="210">Hoplia</emphasis>
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||
(
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||
<emphasis id="71CAEAF93D1DFF40CCB5FE745C5E44A1" bold="true" box="[251,334,404,430]" italics="true" pageId="9" pageNumber="210">Hoplia</emphasis>
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||
)
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<emphasis id="71CAEAF93D1DFF40CD11FE745CDE44A1" bold="true" box="[351,462,404,430]" italics="true" pageId="9" pageNumber="210">messapia</emphasis>
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Uliana, Liberto
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</taxonomicName>
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, and Leo, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="430136EB3D1DFF40CCD9FE565F5544C0" blockId="9.[151,936,404,463]" box="[151,581,438,463]" pageId="9" pageNumber="210">
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||
(
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<figureCitation id="DB852A6E3D1DFF40CCD1FE565C0744C0" box="[159,279,438,463]" captionStart="FIGURES 9 – 11" captionStartId="10.[151,264,1959,1981]" captionTargetBox="[164,1410,451,1889]" captionTargetId="figure@10.[151,1436,439,1949]" captionTargetPageId="10" captionText="FIGURES 9 – 11. Habitus of Hoplia messapia in dorsal and lateral view. 9, male (Cesine, Le Vernole); 10, female (Torre Mozza); 11, male (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891722/files/figure.png" pageId="9" pageNumber="210">Figs. 9–11</figureCitation>
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,
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<figureCitation id="DB852A6E3D1DFF40CD6DFE565C7C44C0" box="[291,364,438,463]" captionStart="FIGURES 12 – 17" captionStartId="12.[151,264,1882,1904]" captionTargetBox="[151,1436,900,1861]" captionTargetId="figure@12.[151,1436,900,1861]" captionTargetPageId="12" captionText="FIGURES 12 – 17. Hoplia dubia species group. Details of scale vestiture of dorsal side (12, 14, 16) and pygidium (13, 15, 17), male specimens. 12 – 13, Hoplia dubia (Marina di Vecchiano); 14 – 15, H. meridiana (Policoro); 16 – 17, H. messapia (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891724/files/figure.png" pageId="9" pageNumber="210">16–17</figureCitation>
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,
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<figureCitation id="DB852A6E3D1DFF40CD39FE575CD044C0" box="[375,448,439,463]" captionStart="FIGURES 18 – 23" captionStartId="13.[151,264,983,1005]" captionTargetBox="[151,1436,193,962]" captionTargetId="figure@13.[151,1436,193,962]" captionTargetPageId="13" captionText="FIGURES 18 – 23. Hoplia dubia species group. Details of scutellar area in oblique (18, 20, 22) and dorsal (19, 21, 23) view, male specimens. 18 – 19, Hoplia dubia (Marina di Vecchiano); 20 – 21, H. meridiana (Policoro); 22 – 23, H. messapia (Torre Mozza). Note the difference between scutellar setae (thin in H. dubia, thick in H. meridiana and H. messapia) and the differences on the relation between the setae of scutellum and periscutellar setae of elytra (reciprocally similar for length and thickness in H. dubia and H. meridiana, different in H. messapia as the scutellar setae are short and thick and the elytral setae long and thin)." httpUri="https://zenodo.org/record/891726/files/figure.png" pageId="9" pageNumber="210">22–23</figureCitation>
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,
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<figureCitation id="DB852A6E3D1DFF40CD85FE565CFA44C0" box="[459,490,438,463]" captionStart="FIGURES 24 – 26" captionStartId="14.[151,264,1816,1838]" captionTargetBox="[151,1372,646,1750]" captionTargetId="figure@14.[151,1436,589,1795]" captionTargetPageId="14" captionText="FIGURES 24 – 26. Hoplia dubia species group. Comparison between the shape of protoracic tarsi (right leg, ventral view), length uniform. 24, Hoplia dubia (Marina di Vecchiano); 25, H. meridiana (R. N. O. Bosco Pantano di Policoro); 26, H. messapia (Cesine, Le Vernole). A stouter structure, with broader tarsomeres and claw, is evident in H. meridiana (courtesy of Daniele Baiocchi and Lara Gallina)." httpUri="https://zenodo.org/record/891728/files/figure.png" pageId="9" pageNumber="210">26</figureCitation>
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,
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<figureCitation id="DB852A6E3D1DFF40CDBBFE575F2D44C0" box="[501,573,439,463]" captionStart="FIGURES 34 – 38" captionStartId="18.[151,264,1939,1961]" captionTargetBox="[212,1389,193,1921]" captionTargetId="figure@18.[212,1389,193,1921]" captionTargetPageId="18" captionText="FIGURES 34 – 38. Hoplia messapia in nature from Vernole, Le Cesine. 34, male on a Tamarix inflorescence, 20. V. 2011 (courtesy of Maurizio Pavesi); 35 – 37, males clustering on the top of grasses and on dry Juncus infructescences, 21. V. 2011 (courtesy of Maurizio Bollino); 38, habitat of the species, 7. VI. 2003 (courtesy of Giuseppe De Matteis)." httpUri="https://zenodo.org/record/891736/files/figure.png" pageId="9" pageNumber="210">34–37</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0BA465603D1DFF43CCD9FE1E58704412" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210" type="description">
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<paragraph id="430136EB3D1DFF40CCD9FE1E5EC44734" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
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||
<materialsCitation id="F3D63CB63D1DFF40CCD9FE1E5FB14718" ID-GBIF-Occurrence="1638471433" box="[151,673,510,535]" location="Specimens" pageId="9" pageNumber="210" specimenCount="417" specimenCount-female="16" specimenCount-male="401">
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<emphasis id="71CAEAF93D1DFF40CCD9FE1E5C894718" bold="true" box="[151,409,510,535]" pageId="9" pageNumber="210">
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<location id="466160303D1DFF40CCD9FE1E5C054718" LSID="urn:lsid:plazi:treatment:CB1787FD3D1DFF43CCD9FE745CC844A3:466160303D1DFF40CCD9FE1E5C054718" box="[151,277,510,535]" name="Specimens" pageId="9" pageNumber="210">Specimens</location>
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||
examined.
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||
</emphasis>
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||
A total of
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||
<specimenCount id="55B8FD623D1DFF40CE50FE1F5F4F4718" box="[542,607,511,535]" pageId="9" pageNumber="210" type="male">401♂</specimenCount>
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||
,
|
||
<specimenCount id="55B8FD623D1DFF40CE20FE1E5FB14718" box="[622,673,510,535]" pageId="9" pageNumber="210" type="female">16♀</specimenCount>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="F3D63CB63D1DFF40CEE1FE1E5FAF4734" ID-GBIF-Occurrence="1638471434" collectingDate="2011-05-21" collectionCode="MSNVE" collectorName="Vernole, R. N. & M. Bollino" country="Italy" latitude="40.35" location="Puglia" longLatPrecision="1163" longitude="18.316668" pageId="9" pageNumber="210" specimenCount="1" specimenCount-male="1" stateProvince="Puglia" typeStatus="holotype">
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||
<typeStatus id="9C0588493D1DFF40CEE1FE1E5E0C4718" box="[687,796,510,535]" pageId="9" pageNumber="210" type="holotype">
|
||
<emphasis id="71CAEAF93D1DFF40CEE1FE1E5E0C4718" bold="true" box="[687,796,510,535]" pageId="9" pageNumber="210">Holotype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectingCountry id="3BA9767B3D1DFF40CF60FE1E5E7A4718" box="[814,874,510,535]" name="Italy" pageId="9" pageNumber="210">Italia</collectingCountry>
|
||
,
|
||
<collectingRegion id="817AF8093D1DFF40CF37FE1E5ED04718" box="[889,960,510,535]" country="Italy" name="Puglia" pageId="9" pageNumber="210">Puglia</collectingRegion>
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||
(LE),
|
||
<collectorName id="EE4B533D3D1DFF40C85EFE1E59BD4719" box="[1040,1197,510,535]" pageId="9" pageNumber="210">Vernole, R.N.</collectorName>
|
||
Le Cesine,
|
||
<geoCoordinate id="268A502C3D1DFF40C975FE1F588C4718" box="[1339,1436,511,535]" direction="north" orientation="latitude" pageId="9" pageNumber="210" precision="925" value="40.35">40°21’N</geoCoordinate>
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<geoCoordinate id="268A502C3D1DFF40CCD9FDC35DE74734" box="[151,247,547,571]" direction="east" orientation="longitude" pageId="9" pageNumber="210" precision="925" value="18.316668">18°19’E</geoCoordinate>
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||
,
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||
<collectingDate id="2744E9C33D1DFF40CD4DFDC35C664734" box="[259,374,547,571]" pageId="9" pageNumber="210" value="2011-05-21">
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||
<date id="3700102B3D1DFF40CD4DFDC35C664734" box="[259,374,547,571]" pageId="9" pageNumber="210" value="2011-05-21">21.V.2011</date>
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||
</collectingDate>
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||
, leg.
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||
<collectorName id="EE4B533D3D1DFF40CDFCFDC25F364734" box="[434,550,546,571]" pageId="9" pageNumber="210">M.Bollino</collectorName>
|
||
<specimenCount id="55B8FD623D1DFF40CE63FDC35F544735" box="[557,580,547,570]" pageId="9" pageNumber="210" type="male">♂</specimenCount>
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||
,
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||
<collectionCode id="25AFAE2E3D1DFF40CE1EFDC35FA74734" box="[592,695,547,571]" pageId="9" pageNumber="210">MSNVE</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<emphasis id="71CAEAF93D1DFF40CE87FDC25E514734" bold="true" box="[713,833,546,571]" pageId="9" pageNumber="210">Paratypes</emphasis>
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||
: see
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||
<tableCitation id="0E3C03503D1DFF40CF35FDC25EC04734" box="[891,976,546,571]" captionStart="TABLE 4" captionStartId="21.[151,239,446,469]" captionText="TABLE 4. Hoplia messapia, list of examined specimens (holotype and paratypes). For standardization of labels data, see Materials and methods." pageId="9" pageNumber="210">Table 4</tableCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="430136EB3D1DFF40CC89FDA65C5C478C" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
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||
<emphasis id="71CAEAF93D1DFF40CC89FDA65FA44750" bold="true" box="[199,692,582,607]" pageId="9" pageNumber="210">Integrative and comparative description.</emphasis>
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For variable characters, the condition of the
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||
<typeStatus id="9C0588493D1DFF40C8F7FDA6580B4750" box="[1209,1307,582,607]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
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||
is between square brackets.
|
||
</paragraph>
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||
<paragraph id="430136EB3D1DFF40CC89FD6E5F6B47A8" blockId="9.[151,1437,510,2011]" box="[199,635,654,679]" pageId="9" pageNumber="210">
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||
<emphasis id="71CAEAF93D1DFF40CC89FD6E5C1B47A8" bold="true" box="[199,267,654,679]" pageId="9" pageNumber="210">Male.</emphasis>
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||
Habitus as in
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<figureCitation id="DB852A6E3D1DFF40CDE5FD6E5F3247A8" box="[427,546,654,679]" captionStart="FIGURES 9 – 11" captionStartId="10.[151,264,1959,1981]" captionTargetBox="[164,1410,451,1889]" captionTargetId="figure@10.[151,1436,439,1949]" captionTargetPageId="10" captionText="FIGURES 9 – 11. Habitus of Hoplia messapia in dorsal and lateral view. 9, male (Cesine, Le Vernole); 10, female (Torre Mozza); 11, male (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891722/files/figure.png" pageId="9" pageNumber="210">Figs. 9, 11</figureCitation>
|
||
,
|
||
<figureCitation id="DB852A6E3D1DFF40CE60FD6F5F6847A8" box="[558,632,655,679]" captionStart="FIGURES 34 – 38" captionStartId="18.[151,264,1939,1961]" captionTargetBox="[212,1389,193,1921]" captionTargetId="figure@18.[212,1389,193,1921]" captionTargetPageId="18" captionText="FIGURES 34 – 38. Hoplia messapia in nature from Vernole, Le Cesine. 34, male on a Tamarix inflorescence, 20. V. 2011 (courtesy of Maurizio Pavesi); 35 – 37, males clustering on the top of grasses and on dry Juncus infructescences, 21. V. 2011 (courtesy of Maurizio Bollino); 38, habitat of the species, 7. VI. 2003 (courtesy of Giuseppe De Matteis)." httpUri="https://zenodo.org/record/891736/files/figure.png" pageId="9" pageNumber="210">34–37</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89FD535EA847E1" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89FD535DEB47C5" box="[199,251,691,714]" italics="true" pageId="9" pageNumber="210">Size.</emphasis>
|
||
Larger than
|
||
<taxonomicName id="84BE4D683D1DFF40CDDEFD545CE447C4" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[400,500,690,715]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40CDDEFD545CE447C4" box="[400,500,690,715]" italics="true" pageId="9" pageNumber="210">H. dubia</emphasis>
|
||
</taxonomicName>
|
||
, similar to
|
||
<taxonomicName id="84BE4D683D1DFF40CE37FD545E1247C4" box="[633,770,690,715]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridian">
|
||
<emphasis id="71CAEAF93D1DFF40CE37FD545E1247C4" box="[633,770,690,715]" italics="true" pageId="9" pageNumber="210">H. meridian</emphasis>
|
||
</taxonomicName>
|
||
a. PBL:
|
||
<quantity id="84469B0E3D1DFF40CF2DFD525EFC47C5" box="[867,1004,690,715]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.6" metricValueMin="6.1" pageId="9" pageNumber="210" unit="mm" value="6.85" valueMax="7.6" valueMin="6.1">6.1–7.6 mm</quantity>
|
||
[6.7], average
|
||
<quantity id="84469B0E3D1DFF40C8D4FD5259E547C5" box="[1178,1269,690,715]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="210" unit="mm" value="6.7">6.7 mm</quantity>
|
||
; TBL:
|
||
<quantity id="84469B0E3D1DFF40C906FD525DD347E1" metricMagnitude="-3" metricUnit="m" metricValue="8.399999999999999" metricValueMax="9.2" metricValueMin="7.6" pageId="9" pageNumber="210" unit="mm" value="8.399999999999999" valueMax="9.2" valueMin="7.6">7.6–9.2 mm</quantity>
|
||
[7.7], average:
|
||
<quantity id="84469B0E3D1DFF40CD3DFD375CDC47E1" box="[371,460,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="9" pageNumber="210" unit="mm" value="8.3">8.3 mm</quantity>
|
||
; MBW:
|
||
<quantity id="84469B0E3D1DFF40CE65FD375FA247E1" box="[555,690,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.9" metricValueMin="3.7" pageId="9" pageNumber="210" unit="mm" value="4.3" valueMax="4.9" valueMin="3.7">3.7–4.9 mm</quantity>
|
||
[4.3], average
|
||
<quantity id="84469B0E3D1DFF40CF14FD375EA347E1" box="[858,947,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="210" unit="mm" value="4.2">4.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89FD1A582F461C" blockId="9.[151,1437,510,2011]" box="[199,1343,762,787]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89FD1A5C17461C" box="[199,263,762,787]" italics="true" pageId="9" pageNumber="210">Color</emphasis>
|
||
. Elytra light brown [as in
|
||
<typeStatus id="9C0588493D1DFF40CE63FD1A5F82461C" box="[557,658,762,787]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
] to brown, specimens with dark or black elytra not observed.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89FCFE59154020" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89FCFE5C1F4638" box="[199,271,798,823]" italics="true" pageId="9" pageNumber="210">Scales</emphasis>
|
||
(
|
||
<figureCitation id="DB852A6E3D1DFF40CD6EFCFE5CB94638" box="[288,425,798,823]" captionStart="FIGURES 12 – 17" captionStartId="12.[151,264,1882,1904]" captionTargetBox="[151,1436,900,1861]" captionTargetId="figure@12.[151,1436,900,1861]" captionTargetPageId="12" captionText="FIGURES 12 – 17. Hoplia dubia species group. Details of scale vestiture of dorsal side (12, 14, 16) and pygidium (13, 15, 17), male specimens. 12 – 13, Hoplia dubia (Marina di Vecchiano); 14 – 15, H. meridiana (Policoro); 16 – 17, H. messapia (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891724/files/figure.png" pageId="9" pageNumber="210">Figs. 16–17</figureCitation>
|
||
). Shape: in general, scales squatter than in
|
||
<taxonomicName id="84BE4D683D1DFF40CFEAFCFF59184638" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[932,1032,798,823]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40CFEAFCFF59184638" box="[932,1032,798,823]" italics="true" pageId="9" pageNumber="210">H. dubia</emphasis>
|
||
</taxonomicName>
|
||
and quite similar to
|
||
<taxonomicName id="84BE4D683D1DFF40C8B3FCFF58854638" box="[1277,1429,798,823]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40C8B3FCFF58854638" box="[1277,1429,798,823]" italics="true" pageId="9" pageNumber="210">H. meridiana</emphasis>
|
||
</taxonomicName>
|
||
, although larger. On the head, elliptical to narrowly lanceolate, not bristle-like. On pronotum, scutellum and elytra broadly elliptical or oval to almost round, usually with broadly rounded apex [as in
|
||
<typeStatus id="9C0588493D1DFF40C830FC8659F64670" box="[1150,1254,870,895]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
]. On pygidium, elliptical or drop shaped (with narrowly acute base) to lanceolate, tapering at the apex, on average slender than on pronotum. Color: on head, nacreous to pale green or cyan. Pronotum dichromatic, with color changing quite abruptly from the discal area to the edges. Discal area yellow to ochre or light brown. Marginal areas, at least near the angles and the lateral margins but sometimes along broad stripes (maximum extension broader than in
|
||
<taxonomicName id="84BE4D683D1DFF40C9CFFC175C1A413C" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40C9CFFC175C1A413C" italics="true" pageId="9" pageNumber="210">H. meridiana</emphasis>
|
||
</taxonomicName>
|
||
) [not in
|
||
<typeStatus id="9C0588493D1DFF40CD25FBFA5CC3413C" box="[363,467,1050,1075]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
], nacreous to cyan. An irregular row of similar scales is present along the basal margin of elytra, the rest of their surface and scutellum with scales similar to those on the pronotal disc. On propygidium, pygidium, ventral side, and legs, similar to the margins of pronotum. Arrangement: distinctly raised from the integument. On head present and sparse on frons and vertex. On pronotum, basal "whirls" observed in
|
||
<taxonomicName id="84BE4D683D1DFF40C97FFB6758854190" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[1329,1429,1158,1183]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40C97FFB6758854190" box="[1329,1429,1158,1183]" italics="true" pageId="9" pageNumber="210">H. dubia</emphasis>
|
||
</taxonomicName>
|
||
, with marginal scales directed forward, are present in some specimens [as in
|
||
<typeStatus id="9C0588493D1DFF40C816FB4A59D041CC" box="[1112,1216,1194,1219]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
], absent in others (corresponding scales are directed towards the midline). On elytra, poorly spaced: in the discal area the transversal distance between two scales is roughly less than the width of one scale to nothing. The clothing therefore looks denser than in the other two species and masks a larger part of the integument.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89FADB596F40B1" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89FADB5C14405D" box="[199,260,1339,1362]" italics="true" pageId="9" pageNumber="210">Setae</emphasis>
|
||
. In general, longer than
|
||
<taxonomicName id="84BE4D683D1DFF40CE63FADC5FD7405C" box="[557,711,1338,1363]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40CE63FADC5FD7405C" box="[557,711,1338,1363]" italics="true" pageId="9" pageNumber="210">H. meridiana</emphasis>
|
||
</taxonomicName>
|
||
and similar to
|
||
<taxonomicName id="84BE4D683D1DFF40CFCCFADC5EF9405C" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[898,1001,1338,1363]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40CFCCFADC5EF9405C" box="[898,1001,1338,1363]" italics="true" pageId="9" pageNumber="210">H. dubia</emphasis>
|
||
</taxonomicName>
|
||
(except scutellum). Longest setae of pronotum about 0.3–0.4x the length of pronotum and 0.9–1.3x the width of scutellum (
|
||
<figureCitation id="DB852A6E3D1DFF40C8C9FABE59CB4078" box="[1159,1243,1374,1399]" captionStart="FIGURES 9 – 11" captionStartId="10.[151,264,1959,1981]" captionTargetBox="[164,1410,451,1889]" captionTargetId="figure@10.[151,1436,439,1949]" captionTargetPageId="10" captionText="FIGURES 9 – 11. Habitus of Hoplia messapia in dorsal and lateral view. 9, male (Cesine, Le Vernole); 10, female (Torre Mozza); 11, male (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891722/files/figure.png" pageId="9" pageNumber="210">Fig. 11</figureCitation>
|
||
,
|
||
<figureCitation id="DB852A6E3D1DFF40C8A4FABE58194078" box="[1258,1289,1374,1399]" captionStart="FIGURES 12 – 17" captionStartId="12.[151,264,1882,1904]" captionTargetBox="[151,1436,900,1861]" captionTargetId="figure@12.[151,1436,900,1861]" captionTargetPageId="12" captionText="FIGURES 12 – 17. Hoplia dubia species group. Details of scale vestiture of dorsal side (12, 14, 16) and pygidium (13, 15, 17), male specimens. 12 – 13, Hoplia dubia (Marina di Vecchiano); 14 – 15, H. meridiana (Policoro); 16 – 17, H. messapia (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891724/files/figure.png" pageId="9" pageNumber="210">16</figureCitation>
|
||
). Length not uniform, decreasing towards the base, but less than in
|
||
<taxonomicName id="84BE4D683D1DFF40CEB7FA645E9E4094" box="[761,910,1410,1435]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40CEB7FA645E9E4094" box="[761,910,1410,1435]" italics="true" pageId="9" pageNumber="210">H. meridiana</emphasis>
|
||
</taxonomicName>
|
||
and only in the medial area, not at the sides of the base. In the medial basal third setae are shorter, measuring 0.5–0.6x the longest setae.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89FA2A5E844324" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
Scutellum with setae shorth and thick, bristle-like, evidently different from those along the base of the pronotum and of the elytra, which are longer and thinner (
|
||
<figureCitation id="DB852A6E3D1DFF40CF66FA0E5EA34308" box="[808,947,1518,1543]" captionStart="FIGURES 18 – 23" captionStartId="13.[151,264,983,1005]" captionTargetBox="[151,1436,193,962]" captionTargetId="figure@13.[151,1436,193,962]" captionTargetPageId="13" captionText="FIGURES 18 – 23. Hoplia dubia species group. Details of scutellar area in oblique (18, 20, 22) and dorsal (19, 21, 23) view, male specimens. 18 – 19, Hoplia dubia (Marina di Vecchiano); 20 – 21, H. meridiana (Policoro); 22 – 23, H. messapia (Torre Mozza). Note the difference between scutellar setae (thin in H. dubia, thick in H. meridiana and H. messapia) and the differences on the relation between the setae of scutellum and periscutellar setae of elytra (reciprocally similar for length and thickness in H. dubia and H. meridiana, different in H. messapia as the scutellar setae are short and thick and the elytral setae long and thin)." httpUri="https://zenodo.org/record/891726/files/figure.png" pageId="9" pageNumber="210">Figs. 22–23</figureCitation>
|
||
). Their length is much shorter than half the length of the scutellum, and close to
|
||
<date id="3700102B3D1DFF40CE7DF9F35F5F4324" box="[563,591,1555,1579]" pageId="9" pageNumber="210">2x</date>
|
||
the length of a scale, or less.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F9D65C45437C" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">Elytral setae hair-like, sparsely mixed in length, long setae present: setae measuring about 1.5-6.0 times the length of a scale.</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F99E5E8A4229" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89F99E5C474398" box="[199,343,1662,1687]" italics="true" pageId="9" pageNumber="210">Morphology.</emphasis>
|
||
Clypeus about 2.3–2.8 [2.5] times wider than long (average: 2.5). Sides of pronotum not regularly curved: straight to slightly concave [slightly concave] in the anterior half, slightly convex to slighty concave in the posterior half [straight], the two halves usually parted by a broadly rounded angle or a sinuation. Sutural region of elytra, compared to the rest of the surface, flat to moderately raised, which is the most common condition [as in
|
||
<typeStatus id="9C0588493D1DFF40CD02F8EE5CA54228" box="[332,437,1806,1831]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
|
||
]. Pygidium W/L =1.35, as in
|
||
<taxonomicName id="84BE4D683D1DFF40CEB3F8EF5E834228" box="[765,915,1806,1831]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40CEB3F8EF5E014229" box="[765,785,1807,1830]" italics="true" pageId="9" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="71CAEAF93D1DFF40CF6EF8EE5E834228" box="[800,915,1806,1831]" italics="true" pageId="9" pageNumber="210">meridiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F8D25E384260" blockId="9.[151,1437,510,2011]" pageId="9" pageNumber="210">
|
||
Protibia, on average, slightly broader than
|
||
<taxonomicName id="84BE4D683D1DFF40CEE1F8D45E044244" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[687,788,1842,1867]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40CEE1F8D45FD34244" box="[687,707,1844,1867]" italics="true" pageId="9" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="71CAEAF93D1DFF40CE9AF8D25E044244" box="[724,788,1842,1867]" italics="true" pageId="9" pageNumber="210">dubia</emphasis>
|
||
</taxonomicName>
|
||
and more similar to
|
||
<taxonomicName id="84BE4D683D1DFF40C84BF8D459B04244" box="[1029,1184,1842,1867]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="meridiana">
|
||
<emphasis id="71CAEAF93D1DFF40C84BF8D459094244" box="[1029,1049,1844,1867]" italics="true" pageId="9" pageNumber="210">H</emphasis>
|
||
.
|
||
<emphasis id="71CAEAF93D1DFF40C867F8D259B04244" box="[1065,1184,1842,1867]" italics="true" pageId="9" pageNumber="210">meridiana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="71CAEAF93D1DFF40C8EEF8D259B44244" box="[1184,1188,1842,1867]" italics="true" pageId="9" pageNumber="210">.</emphasis>
|
||
Average condition of teeth is intermediate between that of the other two species.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F89B5E44429C" blockId="9.[151,1437,510,2011]" box="[199,852,1914,1939]" pageId="9" pageNumber="210">
|
||
Protarsomere slender, similar to that of
|
||
<taxonomicName id="84BE4D683D1DFF40CECCF89C5FF4429C" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[642,740,1914,1939]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="71CAEAF93D1DFF40CECCF89C5FF4429C" box="[642,740,1914,1939]" italics="true" pageId="9" pageNumber="210">H. dubia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DB852A6E3D1DFF40CEBDF89A5E57429C" box="[755,839,1914,1939]" captionStart="FIGURES 24 – 26" captionStartId="14.[151,264,1816,1838]" captionTargetBox="[151,1372,646,1750]" captionTargetId="figure@14.[151,1436,589,1795]" captionTargetPageId="14" captionText="FIGURES 24 – 26. Hoplia dubia species group. Comparison between the shape of protoracic tarsi (right leg, ventral view), length uniform. 24, Hoplia dubia (Marina di Vecchiano); 25, H. meridiana (R. N. O. Bosco Pantano di Policoro); 26, H. messapia (Cesine, Le Vernole). A stouter structure, with broader tarsomeres and claw, is evident in H. meridiana (courtesy of Daniele Baiocchi and Lara Gallina)." httpUri="https://zenodo.org/record/891728/files/figure.png" pageId="9" pageNumber="210">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F87E5F0542B8" blockId="9.[151,1437,510,2011]" box="[199,533,1950,1975]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89F87E5C3542B8" bold="true" box="[199,293,1950,1975]" pageId="9" pageNumber="210">Female.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="DB852A6E3D1DFF40CD8BF87E5F0042B8" box="[453,528,1950,1975]" captionStart="FIGURES 9 – 11" captionStartId="10.[151,264,1959,1981]" captionTargetBox="[164,1410,451,1889]" captionTargetId="figure@10.[151,1436,439,1949]" captionTargetPageId="10" captionText="FIGURES 9 – 11. Habitus of Hoplia messapia in dorsal and lateral view. 9, male (Cesine, Le Vernole); 10, female (Torre Mozza); 11, male (Cesine, Le Vernole)." httpUri="https://zenodo.org/record/891722/files/figure.png" pageId="9" pageNumber="210">Fig 10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1DFF40CC89F822592042D4" blockId="9.[151,1437,510,2011]" box="[199,1072,1986,2011]" pageId="9" pageNumber="210">
|
||
<emphasis id="71CAEAF93D1DFF40CC89F8225C1D42D4" box="[199,269,1986,2011]" italics="true" pageId="9" pageNumber="210">Color.</emphasis>
|
||
Tarsi light brown. Specimens with lighter tibia or femora not observed.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1EFF43CC89FF775EEE45A0" blockId="10.[151,1436,151,429]" box="[199,1022,151,176]" pageId="10" pageNumber="211">
|
||
<emphasis id="71CAEAF93D1EFF43CC89FF775C1F45BF" box="[199,271,151,176]" italics="true" pageId="10" pageNumber="211">Scales</emphasis>
|
||
. Color not different from that of the male, including the pronotum.
|
||
</paragraph>
|
||
<paragraph id="430136EB3D1EFF43CC89FF5C58704412" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="211">
|
||
<emphasis id="71CAEAF93D1EFF43CC89FF5C5C4745DA" box="[199,343,188,213]" italics="true" pageId="10" pageNumber="211">Morphology.</emphasis>
|
||
Elytra different from that of the male: sides more curved, narrowing in the distal part and therefore overall shape shifting from subquadrate to more-or-less oval. Overall difference from the condition of the male is variable, from scarcely perceivable to distinct. Pygidium more convex than in males, ratio W/L: 1.26.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BA465603D1EFF43CC89FEC75ED2446B" pageId="10" pageNumber="211" type="etymology">
|
||
<paragraph id="430136EB3D1EFF43CC89FEC75ED2446B" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="211">
|
||
<emphasis id="71CAEAF93D1EFF43CC89FEC75CB5444F" bold="true" box="[199,421,295,320]" pageId="10" pageNumber="211">Derivatio nominis.</emphasis>
|
||
From the latin adjective "messapius", meaning "of Messapia", in reference to the ancient name of southern
|
||
<collectingRegion id="817AF8093D1EFF43CD2EFEAC5CBC446A" box="[352,428,332,357]" country="Italy" name="Puglia" pageId="10" pageNumber="211">Apulia</collectingRegion>
|
||
(nowadays Salento), where this species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BA465603D1EFF43CC89FE8F5CC844A3" pageId="10" pageNumber="211" type="distribution">
|
||
<paragraph id="430136EB3D1EFF43CC89FE8F5CC844A3" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="211">
|
||
<emphasis id="71CAEAF93D1EFF43CC89FE8F5C704487" bold="true" box="[199,352,367,392]" pageId="10" pageNumber="211">Distribution.</emphasis>
|
||
<taxonomicName id="84BE4D683D1EFF43CD29FE8F5F354487" box="[359,549,367,392]" class="Insecta" family="Melolonthidae" genus="Hoplia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="messapia">
|
||
<emphasis id="71CAEAF93D1EFF43CD29FE8F5F354487" box="[359,549,367,392]" italics="true" pageId="10" pageNumber="211">Hoplia messapia</emphasis>
|
||
</taxonomicName>
|
||
is scattered along the coasts of Salento (southern
|
||
<collectingRegion id="817AF8093D1EFF43C818FE8F59B74487" box="[1110,1191,367,392]" country="Italy" name="Puglia" pageId="10" pageNumber="211">Apulia</collectingRegion>
|
||
), on both the Adriatic and the Ionic sides (
|
||
<figureCitation id="DB852A6E3D1EFF43CD39FE745CDC44A3" box="[375,460,404,429]" captionStart="FIGURE 27" captionStartId="15.[151,250,1729,1751]" captionTargetBox="[151,1436,193,1707]" captionTargetId="figure@15.[151,1436,193,1707]" captionTargetPageId="15" captionText="FIGURE 27. Known distribution of the species of the Hoplia dubia species group." httpUri="https://zenodo.org/record/891730/files/figure.png" pageId="10" pageNumber="211">Fig. 27</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |