301 lines
35 KiB
XML
301 lines
35 KiB
XML
<document id="9280D056EE0D028EBB399494E83C93D3" ENCODING="UTF-8" ID-GBIF-Dataset="a3834a3c-393d-4fc1-9857-4e014cb4a765" ModsDocID="z01117p001" checkinTime="1247232184893" checkinUser="thomas" docAuthor="H. H. Tan & Z. H. Sulaiman" docDate="2006" docId="EE1E971418354EBB42BC16C2380B4BFA" docLanguage="en" docName="2006_Tan_Sulaiman_gg1.xml" docOrigin="Zootaxa 1117" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:EFF4FFDA-A1B8-44EB-ACB4-F4A9623FB4B9" docTitle="Gastromyzon cranbrooki Tan & Sulaiman, 2006, new species" docType="treatment" docVersion="12" lastPageNumber="9" masterDocId="157B8C3E6EA404C176DDA767AF6D510E" masterDocTitle="Three new species of Gastromyzon (Teleostei: Balitoridae) from the Temburong River basin, Brunei Darussalam, Borneo." masterLastPageNumber="19" masterPageNumber="1" pageNumber="3" updateTime="1701314283152" updateUser="plazi">
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<mods:mods id="7C3C252810F828976AB0B1C238C53E69" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="08028597AFDAC66A01E4AB657097128D">
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<mods:title id="52C35A6D762CFF0091CF2A70B75CF301">Three new species of Gastromyzon (Teleostei: Balitoridae) from the Temburong River basin, Brunei Darussalam, Borneo.</mods:title>
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</mods:titleInfo>
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<mods:name id="AF3031DB85F06D64505232329FB80E8D" type="personal">
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<mods:role id="1DE35A9F24096DA1395CAA991B07CDAF">
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<mods:roleTerm id="1ED6CE1CE8A3DB44D9DE4D0D269FB073">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5553EFA01D0E3DE2DEF054E3B2240065">H. H. Tan</mods:namePart>
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</mods:name>
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<mods:name id="ABD56668DEED4D293BBEC786D0E3FFD4" type="personal">
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<mods:role id="218ABF2A169F8F57B8F4C3C8E2476CF0">
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<mods:roleTerm id="2F6091354D38964388E20AF8FFCDC468">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BEDC9A60F480083A0B82E1BBA924AC92">Z. H. Sulaiman</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="012AA46CC1C4335B461279C0594C5750">text</mods:typeOfResource>
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<mods:relatedItem id="110B0D8B526F6344B20550C069DD762E" type="host">
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<mods:titleInfo id="2EA97D226FB9922EEF5654756BDC9171">
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<mods:title id="906AA514ECE557CF96A52673E7B33011">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3851EF0B49D6299EDF998E07484243DF">
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<mods:date id="F97D9CADF9F91953106B018C8F17BA46">2006</mods:date>
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<mods:detail id="DDDB309D6DC83FBAB39B33DD523AE765" type="volume">
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<mods:number id="681EEDB3C368EB366134BDD06E2BBE84">1117</mods:number>
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</mods:detail>
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<mods:extent id="C289D8E751257A91811E4EE1C8FB7013" unit="page">
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<mods:start id="C727F1B812C585EBCC614B7A7D9612AE">1</mods:start>
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<mods:end id="F2744420610998E67B5D6281C8AB71ED">19</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="BDA00E5340C24A6DFF506F131ACCFAE5">
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<mods:url id="91DC73E60558EFAADDC2EEA2C5B103A8">http://www.zoobank.org/urn:lsid:zoobank.org:pub:EFF4FFDA-A1B8-44EB-ACB4-F4A9623FB4B9</mods:url>
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</mods:location>
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<mods:classification id="82701F13F8AB8107C4946EC6D4B15830">journal article</mods:classification>
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<mods:identifier id="3F79C03338A5895635E5998EC2EB0C79" type="Plazi-Custom">z01117p001</mods:identifier>
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</mods:mods>
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<treatment id="EE1E971418354EBB42BC16C2380B4BFA" ID-DOI="http://doi.org/10.5281/zenodo.6256459" ID-GBIF-Taxon="100121945" ID-Zenodo-Dep="6256459" LSID="urn:lsid:plazi:treatment:EE1E971418354EBB42BC16C2380B4BFA" httpUri="http://treatment.plazi.org/id/EE1E971418354EBB42BC16C2380B4BFA" lastPageNumber="9" pageNumber="3">
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<subSubSection id="3EC05C349A9E1A767DC486E9738D243F" type="nomenclature">
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<paragraph id="3DA6FE74C00A1C5D4411DBEF556E69BD" pageNumber="3">
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<taxonomicName id="40B52BFDB95739DE07CC462BEFBA49B7" ID-CoL="3FDR5" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="3" phylum="Chordata" rank="species" species="cranbrooki">Gastromyzon cranbrooki</taxonomicName>
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,
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<taxonomicNameLabel id="0BE69390EA502453B11FA4036C463008" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="6F215E41042D646C3AAE6CD6356CEC4F" type="description">
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<paragraph id="5A8EC6BD09D048679F0957E90AFBC077" pageNumber="3">Figs. 1-2</paragraph>
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</subSubSection>
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<subSubSection id="EE645B3312B0121A3C04EC7778635586" type="reference_group">
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<paragraph id="4B1CB6BEE17601D6111667CD610BD3D0" pageNumber="3">
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<taxonomicName id="2725BF74A3571B1C7D9DA67B97E09B71" LSID-ZBK="urn:lsid:zoobank.org:act:794A7706-9178-4564-BABC-925F845B82EF" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon borneensis Günther 1874:454 [6]" order="Cypriniformes" pageNumber="3" phylum="Chordata" rank="species" species="borneensis">Gastromyzon borneensis</taxonomicName>
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(non
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<normalizedToken id="6F08B5FE6D16C025641CFD9AB71A5A21" originalValue="Günther">Guenther</normalizedToken>
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) - Cranbrook & Edwards, 1994: 327; Choy & Chin, 1994: 761; Doi, 1997: 19 (list; in part).
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</paragraph>
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<paragraph id="74FBA3C84AC5813DD76C3129AD2D7B5C" pageNumber="3">
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<taxonomicName id="74F0F4C7F736BCF8B6E4F475B3FA2FDB" LSID-ZBK="urn:lsid:zoobank.org:act:072F9CC4-7739-44CA-A744-01FDADBADE21" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon fasciatus Inger & Chin 1961:173" order="Cypriniformes" pageNumber="3" phylum="Chordata" rank="species" species="fasciatus">Gastromyzon fasciatus</taxonomicName>
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(non Inger & Chin) - Choy & Chin, 1994: 762; Kottelat & Lim, 1995: 235 (part).
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</paragraph>
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</subSubSection>
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<subSubSection id="362AA57400E074EF1DE9365354949EE8" type="nomenclature">
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<paragraph id="B2C0D081DBDBC4F5CF9A1531EEB567E7" pageNumber="3">
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Material examined:
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<materialsCitation id="165D0BDB2359D9C2EB3E63504BBE43C6" ID-GBIF-Occurrence="923901500" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectingCountry id="4CC41174F89C789B9A6170A4668C7C8A">BRUNEI DARUSSALAM</collectingCountry>
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(
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<collectingRegion id="6DE653A13378BD4508F2AF003A54E522">Temburong district</collectingRegion>
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, Temburong River basin):
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<typeStatus id="2FBDB89AB3B0154E027C51BFC5E69E64">HOLOTYPE</typeStatus>
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:
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<collectionCode id="93D1A03E711E1216900571816C6273C9" collectionName="UBD">UBD</collectionCode>
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uncatalogued, 48.2 mm SL; Sungai Belalong, in front and near to Kuala Belalong Field Studies Centre (
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<geoCoordinate id="461062581A796152C88FCD2936494847" direction="north" orientation="latitude" value="4.5473332">04°32'50.4"N</geoCoordinate>
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<geoCoordinate id="AA432FA037F11F7A0EB5101ABCE3A79E" direction="east" orientation="longitude" value="115.15767">115°09'27.6"E</geoCoordinate>
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[80 m asl]); H. H. Tan & K. K. P. Lim, 4-
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<date id="65106A676EA3E92F2859529AFCFC8113" day="07" month="10" value="2001-10-07" year="2001">7 Oct 2001</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="5FE2ADC7F6306709F899DAC772C63504" pageNumber="3">
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<materialsCitation id="6D90CBFFC1D75094DD64C9778DF150A6" ID-GBIF-Occurrence="923901505" country="Brunei Darussalam" stateProvince="Temburong District">
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<typeStatus id="F47FEBCFEF239B9C0A9ED9D3FA123BBA">PARATYPES</typeStatus>
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:
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<collectionCode id="ADF5FD25424FF04C55940630355CE62E" collectionName="UBD">UBD</collectionCode>
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uncatalogued, 26 ex., 20.0-45.0 mm SL
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</materialsCitation>
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;
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<materialsCitation id="4AC2D8B07E1953EDEE9A69A56F8104ED" ID-GBIF-Occurrence="923901497" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="2E4FA6B8B6CB22390DDDB653BE48F2F3" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47003, 26 ex., 19.9-53.5 mm SL; same locality as holotype
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</materialsCitation>
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.
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<materialsCitation id="67DA9386EC0A988F9AEE3BF17FC5DE50" ID-GBIF-Occurrence="923901499" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="7AF5BA8495B50C0584FECEF7F25E5363" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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38757, 12 ex., 19.7-61.4 mm SL; Temburong River basin: Sungai Belalong at Kuala Belalong; K. K. P. Lim et al., 14-
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<date id="B2E66DCA38768700557F62A508689266" day="17" month="06" value="1995-06-17" year="1995">17 Jun 1995</date>
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</materialsCitation>
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.
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<materialsCitation id="EEF413464D0115312197F2CE931DA3B1" ID-GBIF-Occurrence="923901506" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="5494A47D6B7652B1A551E41FDEE77F45" collectionName="CMK">CMK</collectionCode>
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11553, 2 ex., 45.2-47.7 mm SL; Temburong River basin: Sungai Temburong, about 1 km downstream of Kuala Belalong; K. K. P. Lim et al.,
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<date id="195E09ADEEC136DECA3F6503F40D3EF8" day="16" month="06" value="1995-06-16" year="1995">16 Jun 1995</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7F95D8CEB8D5B22563EF3F5DCE175F50" type="materials_examined">
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<paragraph id="0CDA47E978527594FE3517CF4D96AAC6" pageNumber="3">
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NON-TYPE MATERIAL:
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<materialsCitation id="F191BD9C71D07F1D5126F9AD97B17B1A" ID-GBIF-Occurrence="923901504" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectingCountry id="70374F24439A9E9B8BC86859CA24568D">Brunei Darussalam</collectingCountry>
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(
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<collectingRegion id="E5CC95702EFD57FE53BA7AC6BF8BFF52">Temburong district</collectingRegion>
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: Temburong River basin):
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<collectionCode id="0A3C5DBCFDE8ACB65FA95A53EC99B1C6">UBD-SC</collectionCode>
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/92/7/F6, 4 ex., 17.9-52.8 mm SL; Sungai Belalong, Kuala Belalong Field Study Centre; S. Choy,
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<date id="8A88ABE93372660CAB5749DF8FDAA884" day="27" month="07" value="1992-07-27" year="1992">27 Jul 1992</date>
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</materialsCitation>
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.
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<materialsCitation id="01340ADD0B89D83FB5BD08971B2642CD" ID-GBIF-Occurrence="923901501" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="8735ACDA28669F68C0B0225D13DBDDD8">UBD-SC</collectionCode>
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/91/8/3, 1 ex., 75.5 mm SL; Sungai Belalong, upper Sungai Babi, past HP171; S. Choy & S. Nyawa,
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<date id="7F9451D69E4C0857F2E682FEDF42784F" day="07" month="08" value="1991-08-07" year="1991">7 Aug 1991</date>
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</materialsCitation>
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.
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<materialsCitation id="296E98A7B8664DDBECCFC686BBBCB0FE" ID-GBIF-Occurrence="923901502" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="FEFBBD1F84D950AE2FB6E8E65862E39D">UBD-SC</collectionCode>
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/92/6/2, 3 ex., 38.7-52.0 mm SL; Sungai Belalong, near Kuala Belalong Field Study Centre; S. Choy,
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<date id="6B155C2EAA24D48DC2B9A87FE1232470" day="28" month="06" value="1992-06-28" year="1992">28 Jun 1992</date>
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</materialsCitation>
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.
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<materialsCitation id="60955BC2A74C3AD23F791FC268A93E1F" ID-GBIF-Occurrence="923901507" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="9671FC6505C3DD1267DDA6985115D921" collectionName="UBD">UBD</collectionCode>
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uncatalogued, 4 ex., 45.7-58.1 mm SL; Sungai Belalong, opposite Kuala Belalong Field Study Centre intake pipe; S. Choy,
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<date id="B0ED0696B80FF309AE2719CBDE061854" day="27" month="07" value="1992-07-27" year="1992">27 Jul 1992</date>
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</materialsCitation>
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.
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<materialsCitation id="C6F9D3EAA09FE40B379BC9F59696DFFC" ID-GBIF-Occurrence="923901503" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="41E7D93E404500842D31CB460433E44E">UBD-SC</collectionCode>
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/92/7/1, 3 ex., 40.5-42.6 mm SL; Sungai Belalong, near intake pipe; A. K. Zainal & Awg. Moss,
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<date id="7DA3414B9AFB773DE120B3FB604135D7" day="27" month="07" value="1992-07-27" year="1992">27 Jul 1992</date>
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</materialsCitation>
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.
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<materialsCitation id="3123FAF257B2C4C3EFF715C2732B0CA5" ID-GBIF-Occurrence="923901508" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="BF655707EC02D4A6F3E78858D4068070">UBD-SC</collectionCode>
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/92/8/7, 6 ex., 39.9-59.2 mm SL; Sungai Belalong near Kuala Belalong Field Study Centre, near water intake pipe; S. Choy,
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<date id="B881F714682E0CE49B84D41224B74022" month="12" value="1991-12" year="1991">Dec 1991</date>
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</materialsCitation>
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.
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<materialsCitation id="90C134421753DA1BDD2B66B4C572C676" ID-GBIF-Occurrence="923901510" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="33511725CDDA14A9CB3BBB4EC2C4531C">UBD-B-</collectionCode>
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11, 11 ex., 14.6- 54.6 mm SL; Sungai Belalong at Kuala Belalong; K. K. P. Lim et al., 14-
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<date id="7CAF107AEBF2132133433FBE0F515C1A" day="17" month="06" value="1995-06-17" year="1995">17 Jun 1995</date>
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</materialsCitation>
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.
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<materialsCitation id="E63BCDF6F215E1ED441BE315E1442F2B" ID-GBIF-Occurrence="923901509" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="8E661AA3281694A38E172326451D4F38" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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31803, 1 ex., 46.7 mm SL; Sungai Belalong near Kuala Belalong Field Study Centre; S. C. Choy,
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<date id="58CBAF52AC99DD7C91D737CDFCC9E684" day="24" month="07" value="1992-07-24" year="1992">24 Jul 1992</date>
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</materialsCitation>
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.
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<materialsCitation id="3FB15B9D8A48169B2380A47513E381B4" ID-GBIF-Occurrence="923901513" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="BDB913823E716EDCD00D226B3293EAF9" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47004, 3 ex., 35.0-54.1 mm SL; Belalong River basin; Sungai Enkabang, about 15 minutes upstream of Kuala Belalong Field Studies Centre (
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<geoCoordinate id="3A20D3A95770249A83BC0463AEC75DAD" direction="north" orientation="latitude" value="4.537083">04°32'13.5"N</geoCoordinate>
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<geoCoordinate id="448F716DC7F3335BDE8AC83198BC175D" direction="east" orientation="longitude" value="115.15972">115°09'35.0"E</geoCoordinate>
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[100 m asl]); H. H. Tan & K. K. P. Lim,
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<date id="AD86AB86CC0B1983E3C4CBBC3FDE8B53" day="05" month="10" value="2001-10-05" year="2001">5 Oct 2001</date>
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</materialsCitation>
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.
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<materialsCitation id="C857107A7D179AEBF1AE5A1CECFABD8A" ID-GBIF-Occurrence="923901511" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="63B48FB67180A5742E4085F2CEC88A16" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47005, 1 ex., 51.3 mm SL; Belalong River basin; Sungai Belalong at mouth of Sungai Enkabang (
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<geoCoordinate id="45D5CA5D054DBC8ED5EC85F055BFDE0B" direction="north" orientation="latitude" value="4.537083">04°32'13.5"N</geoCoordinate>
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<geoCoordinate id="393078CF288290AD1987889D7C4B26C9" direction="east" orientation="longitude" value="115.15972">115°09'35.0"E</geoCoordinate>
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[100 m asl]); H. H. Tan et al.,
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<date id="F6C279DFD05DA7BD783D3440B84D7CC8" day="06" month="10" value="2001-10-06" year="2001">6 Oct 2001</date>
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</materialsCitation>
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.
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<materialsCitation id="7AF060BF68633B7DC298480D92F04ED0" ID-GBIF-Occurrence="923901514" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="920096A6E03CB2D633A1137923D692AC" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47006, 5 ex., 39.4-52.2 mm SL; Belalong River basin; Sungai Belalong at mouth of Sungai Esu (
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<geoCoordinate id="A916651B33705B284D11B8D0EC97C04A" direction="north" orientation="latitude" value="4.5383058">04°32'17.9"N</geoCoordinate>
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<geoCoordinate id="30DC7E0487E7468363956FEDD1911C53" direction="east" orientation="longitude" value="115.15978">115°09'35.2"E</geoCoordinate>
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); H. H. Tan et al.,
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<date id="1EA7D663404D73D8082B3E3C2D141739" day="06" month="10" value="2001-10-06" year="2001">6 Oct 2001</date>
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</materialsCitation>
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.
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<materialsCitation id="5943E8BF9F3FA622BB8DD8DB39D8C5B1" ID-GBIF-Occurrence="923901515" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="1E45B451D84E8F7E98B5A5FD51CCB7AD" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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47007, 1 ex., 48.7 mm SL; Sungai Temburong, just upstream of confluence of Sungai Temburong and Sungai Belalong; H. H. Tan et al.,
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<date id="ACFD0DD8ABDF3536E370127ABE3DB2BE" day="05" month="10" value="2001-10-05" year="2001">5 Oct 2001</date>
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</materialsCitation>
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.
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<materialsCitation id="C3EE1724EACFB1C4A80BB099164914F1" ID-GBIF-Occurrence="923901512" country="Brunei Darussalam" stateProvince="Temburong District">
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<collectionCode id="BDE9F2597EFBB9C9D90FD50468ABE51F" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
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|
47008, 2 ex., 33.5-57.5 mm SL; Belalong River basin; Sungai Engkiang, about 40 minutes upstream of Kuala Belalong Field Studies Centre (about 3-4 km upstream) (
|
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<geoCoordinate id="5BAF05E3D59188F39814A6FDADD9D5AF" direction="north" orientation="latitude" value="4.4857497">04°29'08.7"N</geoCoordinate>
|
|
<geoCoordinate id="8733661EC8381BDD96A3D8BC3F79BECF" direction="east" orientation="longitude" value="115.145386">115°08'43.4"E</geoCoordinate>
|
|
[120 m asl]); H. H. Tan & K. K. P. Lim,
|
|
<date id="9D6339AF6211DBDDBC377D095225551F" day="08" month="10" value="2001-10-08" year="2001">8 Oct 2001</date>
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|
</materialsCitation>
|
|
.
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|
</paragraph>
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|
</subSubSection>
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|
<subSubSection id="2CC57D3971F7B45FADBCE15730AB22C6" type="diagnosis">
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<paragraph id="BA441449C7F870007BDEE9F6B2362AA4" pageNumber="4">
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|
Diagnosis.
|
|
<taxonomicName id="96B3F2C0CDB0FFDBB578B4C5F7A85AC8" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="4" phylum="Chordata" rank="species" species="cranbrooki">Gastromyzon cranbrooki</taxonomicName>
|
|
differs from its congeners in having the following unique combination of characters: presence of distinct secondary rostrum; presence of complete postoral pouch; body brown, with 9-10 grey bars, head dorsum dark brown, with thin grey reticulate pattern; absence of subopercular groove; gill slit vertical; presence of sublacrymal groove; snout truncate when viewed dorsally; abdomen without scales; 56-60 scales in lateral line; pelvic fin not overlapping anal fin origin, adpressed dorsal fin falling short of level of anal fin origin. Maximum size: 75.5 mm SL (UBD-SC/91/8/3).
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</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="65D1E7B72E0700EFC75FBA6D7281AC7C" type="description">
|
|
<paragraph id="EAE316BA793D267A421B2D46A448CC5F" pageNumber="4">Description. General body shape and appearance as in Figs. 1-2. Meristic and morphometric data appear in Table 1. Head truncate when viewed in dorsal view, relatively short (26.2-29.6 % SL) and wide (26.2-30.4 % SL, 88.5-106.7 % HL), head relatively flattened (head depth 13.5-14.9 % SL, 47.4-52.4 % HL). Tubercles concentrated on anterior part of snout below secondary rostrum. Snout relatively long (65.7-70.3 % HL), with broadly rounded secondary rostrum; mature males with secondary rostrum longer than primary rostrum. Sublacrymal groove present, not visible from side. Gill slit vertical, its length less than eye diameter. Subopercular groove absent. Postoral pouch complete. No scales on belly. Pectoral and pelvic fins with serrae on anteriormost rays. Posterior part of pectoral fin just reaching anterior part of pelvic fin. Pelvic fin not reaching anal fin origin. Dorsal fin situated at about mid body (predorsal length 53.8-58.8 % SL), adpressed dorsal fin falling short of level of anal-fin origin. Deepest part of body at dorsal-fin origin (body depth at dorsal-fin origin 17.3-20.8 % SL). Anus between posterior base of fused pelvic fins and anal fin origin. Some specimens with tubercles on posterior part of lower half of body. Caudal peduncle relatively deep (8.6-9.6 % SL) and short (7.6- 10.4 % SL).</paragraph>
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<paragraph id="2C48504CEC5DBD787E8C49921EC52626" pageNumber="6">Body pigmentation and life coloration- See Fig. 1. This species was first depicted by Cranbrook & Edwards (1994: 327). Body dark brown on dorsum and sides, body with 9- 10 grey bars on side of body that are continuous across dorsum and connect with bars on opposite side; a white spot on posterior edge of every body scale; ventrum cream. Head dorsum dark brown with thin grey reticulate pattern. Eye with golden iris. Dorsal fin grey on lowermost two-thirds of rays, the interradial membranes and margin of fin hyaline. A basal black spot present on anterior part of dorsal fin. Caudal-fin base black, the fin with 4-5 black bars, fin with hyaline edge and with golden sheen in the middle section of fin. Anal-fin rays black on lowermost two-thirds; margin of fin broadly hyaline, with golden sheen. Pectoral-fin rays dark brown with 2-3 faint grey bands and hyaline margin. Pelvic axillary flap brown with yellowish margin. Pelvic-fin dark brown with 2 faint grey bands, thin hyaline margin on anterior part of fin; a thicker hyaline margin on posterior part of fin.</paragraph>
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<paragraph id="E27AE509476B3FE908945EC4355EA6B1" lastPageNumber="7" pageNumber="6">Colour in alcohol. See Fig. 2. Body black on dorsum and sides, with 9-10 grey bars extending from one side to the other side across dorsum; ventrum cream. Head dorsum black with faint thin grey reticulate pattern. Dorsal-fin rays grey on lowermost two-thirds, the interradial membranes and margin of fin hyaline. A basal black spot present on anterior part of dorsal fin. Caudal fin base black, the fin rays grey with 4-5 black bars across fin, and with interradial membranes and posterior margin hyaline. Anal-fin rays grey, with 2 black bars, and with a wide hyaline area on posterior margin of fin. Pectoral fin black, with 2-3 faint grey bands, and with a very thin hyaline posterior margin. Pelvic axillary flap black, with a thin grey margin. Pelvic fin black, with 2 faint grey bands, a very thin hyaline margin anteriorly, and a thick hyaline margin posteriorly. Juveniles with up to 11 cream bars on body, head with faint cream reticulate pattern, caudal fin with 2 black bars.</paragraph>
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</subSubSection>
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|
<subSubSection id="F791C05CC74794F534A7FEDC14B9717B" type="distribution">
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|
<paragraph id="1C2FD7C5983DDAE1855900DB35880650" pageNumber="8">
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Distribution.
|
|
<taxonomicName id="B6E03BB4564FD7D7C5D4434A5F899469" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="cranbrooki">Gastromyzon cranbrooki</taxonomicName>
|
|
is currently known only from the Temburong River basin, Brunei Darussalam (Fig. 7).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B2F956FD70316E81FE525BD1B1C077CE" type="discussion">
|
|
<paragraph id="F3DB29D481FDB380A77BA0E764DF1E11" pageNumber="8">
|
|
Remarks.
|
|
<taxonomicName id="71B88D6EFC9180A8C1C23A923CE3CF5F" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="cranbrooki">Gastromyzon cranbrooki</taxonomicName>
|
|
may be readily confused with
|
|
<taxonomicName id="44C6CD24AEACA320B5B4273FDEEB18DE" LSID-ZBK="urn:lsid:zoobank.org:act:794A7706-9178-4564-BABC-925F845B82EF" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon borneensis Günther 1874:454 [6]" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
|
, which also possesses a barred body and a secondary rostrum. In addition to characters included in the Diagnosis,
|
|
<taxonomicName id="23E2223C04B7014C223361A98039D56C" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="cranbrooki">G. cranbrooki</taxonomicName>
|
|
can be further differentiated from the other congeners of the
|
|
<taxonomicName id="1BAFDF34144A558EFDB394D9A37111A0" LSID-ZBK="urn:lsid:zoobank.org:act:794A7706-9178-4564-BABC-925F845B82EF" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon borneensis Günther 1874:454 [6]" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="borneensis">G. borneensis group</taxonomicName>
|
|
by the following characters: presence of a secondary rostrum (vs. absence in
|
|
<taxonomicName id="832657B0C85B5ADBAF5D0B37D9034832" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="monticola">G. monticola</taxonomicName>
|
|
and
|
|
<taxonomicName id="7296D1F2C32988210531C5B1B7D0B3A6" LSID-ZBK="urn:lsid:zoobank.org:act:18DED009-1FD6-4BF6-ADDC-8A48CBB97007" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ornaticauda Tan & Martin-Smith 1998:367" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="ornaticauda">G. ornaticauda</taxonomicName>
|
|
); head truncate in dorsal view (vs. rounded in
|
|
<taxonomicName id="421D0EECCADAB78B2C13E7F38301B259" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="monticola">G. monticola</taxonomicName>
|
|
); pelvic fin not overlapping anal fin origin (vs. overlapping in
|
|
<taxonomicName id="D49535E5D5C6568B0ED5E6B5320E79D9" LSID-ZBK="urn:lsid:zoobank.org:act:18DED009-1FD6-4BF6-ADDC-8A48CBB97007" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ornaticauda Tan & Martin-Smith 1998:367" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="ornaticauda">G. ornaticauda</taxonomicName>
|
|
); pectoral fin length less than in
|
|
<taxonomicName id="B3F030AC0B82FBBFC12B251FCFFF9691" LSID-ZBK="urn:lsid:zoobank.org:act:18DED009-1FD6-4BF6-ADDC-8A48CBB97007" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ornaticauda Tan & Martin-Smith 1998:367" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="ornaticauda">G. ornaticauda</taxonomicName>
|
|
(37.4-41.5, vs. 41.4-43.7 % SL); greater head width than in
|
|
<taxonomicName id="440FC5684F937FB1DB9EEE3C66DEF4D3" LSID-ZBK="urn:lsid:zoobank.org:act:18DED009-1FD6-4BF6-ADDC-8A48CBB97007" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ornaticauda Tan & Martin-Smith 1998:367" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="ornaticauda">G. ornaticauda</taxonomicName>
|
|
(26.2-30.4, vs. 21.7-25.9 % SL); greater orbit diameter than in
|
|
<taxonomicName id="9104FAF82ADDF559ED69F785EC5E7027" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="8" phylum="Chordata" rank="species" species="monticola">G. monticola</taxonomicName>
|
|
(3.7-4.5, vs. 2.9-3.5 % SL).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7DD594E428C11F886950D980E4B225FD" type="etymology">
|
|
<paragraph id="58A3B3E6FCF74016B2682B70B09E08A5" pageNumber="9">Etymology. Named for the Earl of Cranbrook, in recognition of his contributions to the study of biodiversity in Southeast Asia.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1953E807070D5DEE2DD92F1ABBBB6FA7" type="biology_ecology">
|
|
<paragraph id="CFA5414F08AC5EE97E8739A4FFE8A267" pageNumber="9">
|
|
Field notes.
|
|
<taxonomicName id="05BB2C04238392824FBF467B92FA9A21" LSID-ZBK="urn:lsid:zoobank.org:act:D41EEA6C-4212-4E99-B70E-54D86FB848BD" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon cranbrooki Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="cranbrooki">Gastromyzon cranbrooki</taxonomicName>
|
|
is the most common
|
|
<taxonomicName id="1C1D258973EF8A5653CD615B33977838" LSID-ZBK="urn:lsid:zoobank.org:act:D6E2004D-AAC6-4A97-9D32-DC5186F09138" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon Günther 1874:454" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="genus">Gastromyzon</taxonomicName>
|
|
species in the riffle zones of Sungai Belalong. Most specimens were obtained from a rocky area about 15 metres wide, just 20 metres upstream of the Kuala Belalong Field Studies Centre (Fig. 8), in shallow flowing water less than 50 cm deep, and with the pH 6.6. The substrate consists of a mix of small rocks and gravel of quartz, basalt and shale origin. Syntopic species include three other
|
|
<taxonomicName id="E42A96C6DD515D815B8AD4630A496DD7" LSID-ZBK="urn:lsid:zoobank.org:act:D6E2004D-AAC6-4A97-9D32-DC5186F09138" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon Günther 1874:454" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="genus">Gastromyzon</taxonomicName>
|
|
(
|
|
<taxonomicName id="FE006126DD19B3C9ABDB473E13C411EB" LSID-ZBK="urn:lsid:zoobank.org:act:6F3EEF4F-DE3C-4BAE-971B-B34EB98F9A98" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon aeroides Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="aeroides">G. aeroides</taxonomicName>
|
|
,
|
|
<taxonomicName id="BED7EB2F798A359FF62A3C61E962626A" LSID-ZBK="urn:lsid:zoobank.org:act:40A76FED-AE30-4C16-8A44-91CE544CFEA4" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon punctulatus Inger & Chin 1961:173" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="punctulatus">G. punctulatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="A4D0B9A6A12B3DF398CD995FDCCFEDF5" LSID-ZBK="urn:lsid:zoobank.org:act:CEC6BB92-06C1-4BB1-8E1E-1939B5547CF5" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon venustus Tan & Sulaiman 2006" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="venustus">G. venustus</taxonomicName>
|
|
) and two additional balitorid species (
|
|
<taxonomicName id="9D3BE24F8FF2EADE0C65274A9BECE2CA" family="Balitoridae" genus="Parhomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="microstoma">Parhomaloptera microstoma</taxonomicName>
|
|
and
|
|
<taxonomicName id="A249E5380F1EEA1FA32D3832D1E80008" LSID-ZBK="urn:lsid:zoobank.org:act:61CA7DD9-F636-4EE4-AD80-CC42AA7AE7FB" family="Balitoridae" genus="Neogastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neogastromyzon Popta 1905:180" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="genus">Neogastromyzon</taxonomicName>
|
|
sp.). Other syntopic species include
|
|
<taxonomicName id="753BB9A1A4CA9E6DFF41DEB07A85A64F" family="Cyprinidae" genus="Barbonymus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="collingwoodii">Barbonymus collingwoodii</taxonomicName>
|
|
,
|
|
<taxonomicName id="7A031B71082E7A96E4D66488DC9B84AA" family="Cyprinidae" genus="Hampala" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="bimaculata">Hampala bimaculata</taxonomicName>
|
|
,
|
|
<taxonomicName id="F95366436F99AF7E4D6AED6160C89ED5" family="Cyprinidae" genus="Lobocheilos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="bo">Lobocheilos cf. bo</taxonomicName>
|
|
,
|
|
<taxonomicName id="4E278B058D1D89EBE0E2728AB958A9D3" LSID-ZBK="urn:lsid:zoobank.org:act:16F97B18-9717-4EF0-9758-2D1738FBC87E" family="Cyprinidae" genus="Nematabramis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Nematabramis steindachnerii Popta 1905:179" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="steindachneri">Nematabramis steindachneri</taxonomicName>
|
|
,
|
|
<taxonomicName id="C3BEED2F4EF21D2439A184DE105CFBD6" family="Cyprinidae" genus="Paracrossocheilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="acerus">Paracrossocheilus acerus</taxonomicName>
|
|
,
|
|
<taxonomicName id="290E3485B8851764989281B130FE8C84" family="Cyprinidae" genus="Rasbora" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="agyrotaenia">Rasbora agyrotaenia</taxonomicName>
|
|
,
|
|
<taxonomicName id="5EA260776945DD27EE2327D7E0FDC282" family="Cyprinidae" genus="Tor" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="tambra">Tor tambra</taxonomicName>
|
|
(Cyprinidae),
|
|
<taxonomicName id="63F5AA12A61D48465D39B9BFDF670B62" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="9" phylum="Chordata" rank="species" species="major">Glyptothorax major</taxonomicName>
|
|
(Sisoridae),
|
|
<taxonomicName id="A9390180FAA5A7E59D1319A13D837ADA" LSID-ZBK="urn:lsid:zoobank.org:act:EDE3D933-9D70-4E6C-B4B6-4CCFE4BC1B9E" family="Gobiidae" genus="Eugnathogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Eugnathogobius Smith 1931:37" order="Perciformes" pageNumber="9" phylum="Chordata" rank="genus">Eugnathogobius</taxonomicName>
|
|
sp.,
|
|
<taxonomicName id="AA9D3420C705DE3BDF268CBF06EDEE52" LSID-ZBK="urn:lsid:zoobank.org:act:810BF1C2-E52D-4228-BF62-28BA4FB4F363" family="Gobiidae" genus="Parawaous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Parawaous Watson 1993:178" order="Perciformes" pageNumber="9" phylum="Chordata" rank="genus">Parawaous</taxonomicName>
|
|
sp. (Gobiidae), and
|
|
<taxonomicName id="19FB08374F90B9271DE612B76DED71CA" family="Mastacembelidae" genus="Macrognathus" higherTaxonomySource="CoL" kingdom="Animalia" order="Synbranchiformes" pageNumber="9" phylum="Chordata" rank="species" species="maculatus">Macrognathus maculatus</taxonomicName>
|
|
(Mastacembelidae).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |