427 lines
42 KiB
XML
427 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1113.85448" ID-Pensoft-Pub="1313-2970-1113-167" ID-Pensoft-UUID="3DF554A1DDD95950A5C7336AB4FAB97C" ID-ZooBank="AF6603ADBD0347EEAFD005459C3CCC9E" ModsDocID="1313-2970-1113-167" checkinTime="1658252669484" checkinUser="pensoft" docAuthor="Martynov, Alexander V., Sivaruban, T., Palatov, Dmitry M., Srinivasan, Pandiarajan, Barathy, S., Isack, Rajasekaran & Sartori, Michel" docDate="2022" docId="CC90A06F686C53918E0A26CFF2F627DB" docLanguage="en" docName="ZooKeys 1113: 167-197" docOrigin="ZooKeys 1113" docPubDate="2022-07-18" docSource="http://dx.doi.org/10.3897/zookeys.1113.85448" docTitle="Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack 2022, sp. nov." docType="treatment" docUuid="060B341B-E366-4FC0-96B4-C81BE326FAF4" docUuidSource="ZooBank" docVersion="1" id="3DF554A1DDD95950A5C7336AB4FAB97C" lastPageNumber="167" masterDocId="3DF554A1DDD95950A5C7336AB4FAB97C" masterDocTitle="Contribution to the knowledge of Teloganodidae (Ephemeroptera, Ephemerelloidea) of India" masterLastPageNumber="197" masterPageNumber="167" pageNumber="167" updateTime="1658252669484" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Contribution to the knowledge of Teloganodidae (Ephemeroptera, Ephemerelloidea) of India</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Martynov, Alexander V.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6506-5134</mods:nameIdentifier>
|
|
<mods:affiliation>National Museum of Natural History, National Academy of Sciences of Ukraine, Bohdan Khmelnytsky str., 15, 01030, Kyiv, Ukraine</mods:affiliation>
|
|
<mods:nameIdentifier type="email">centroptilum@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sivaruban, T.</mods:namePart>
|
|
<mods:affiliation>PG & Research Department of Zoology, The American College, Madurai- 625002, India</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Palatov, Dmitry M.</mods:namePart>
|
|
<mods:affiliation>Independent researcher, Lviv, Ukraine</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8118-3256</mods:nameIdentifier>
|
|
<mods:affiliation>PG & Research Department of Zoology, The American College, Madurai- 625002, India</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Barathy, S.</mods:namePart>
|
|
<mods:affiliation>Department of Zoology, Fatima College, Madurai- 625018, India</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Isack, Rajasekaran</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9952-4335</mods:nameIdentifier>
|
|
<mods:affiliation>PG & Research Department of Zoology, The American College, Madurai- 625002, India</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sartori, Michel</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
|
|
<mods:affiliation>Musee cantonal de zoologie, Palais de Rumine, Place de la Riponne 6, 1014 Lausanne, Switzerland & Department of Ecology and Evolution, Biophore, University of Lausanne, 1015 Lausanne, Switzerland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-07-18</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>1113</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>167</mods:start>
|
|
<mods:end>197</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.1113.85448</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1113.85448</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-1113-167</mods:identifier>
|
|
<mods:identifier type="ZooBank">AF6603ADBD0347EEAFD005459C3CCC9E</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">3DF554A1DDD95950A5C7336AB4FAB97C</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:zoobank.org:act:060B341B-E366-4FC0-96B4-C81BE326FAF4" httpUri="http://treatment.plazi.org/id/CC90A06F686C53918E0A26CFF2F627DB" lastPageNumber="167" pageId="0" pageNumber="167">
|
|
<subSubSection pageId="0" pageNumber="167" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<taxonomicName LSID="https://zoobank.org/060B341B-E366-4FC0-96B4-C81BE326FAF4" authority="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityName="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes molinerii" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="molinerii" status="sp. nov.">Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="description">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">Figs 9</figureCitation>
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">, 10</figureCitation>
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">, 11</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="167">Material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<materialsCitation collectingDate="2020-10-28" collectorName="Pandiarajan Srinivasan, Isack Rajasekaran" country="India" county="Theni district" elevation="632" latitude="10.0809" location="Kottakudi River" longLatPrecision="7" longitude="77.2552" municipality="Kurangani" specimenCount="♀" stateProvince="Tamil Nadu" typeStatus="Holotype">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="167">
|
|
<typeStatus>Holotype</typeStatus>
|
|
</emphasis>
|
|
: mature
|
|
<specimenCount>♀</specimenCount>
|
|
larva,
|
|
<collectingCountry name="India">India</collectingCountry>
|
|
,
|
|
<collectingRegion country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
|
|
,
|
|
<collectingCounty>Theni district</collectingCounty>
|
|
,
|
|
<collectingMunicipality>Kurangani</collectingMunicipality>
|
|
hills,
|
|
<location LSID="urn:lsid:plazi:treatment:CC90A06F686C53918E0A26CFF2F627DB:A6B0F75D709E19FB840DEC782DE3DED2" country="India" county="Theni district" latitude="10.0809" longLatPrecision="7" longitude="77.2552" municipality="Kurangani" name="Kottakudi River" stateProvince="Tamil Nadu">Kottakudi River</location>
|
|
,
|
|
<geoCoordinate degrees="10.0809" direction="north" orientation="latitude" precision="5" value="10.0809">10.0809°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="77.2552" direction="east" orientation="longitude" precision="5" value="77.2552">77.2552°E</geoCoordinate>
|
|
,
|
|
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.32" unit="m" value="632.0">
|
|
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.32" unit="m" value="632.0">632 m</quantity>
|
|
a.s.l.
|
|
</elevation>
|
|
,
|
|
<collectingDate value="2020-10-28">28.x.2020</collectingDate>
|
|
,
|
|
<collectorName>Pandiarajan Srinivasan</collectorName>
|
|
&
|
|
<collectorName>Isack Rajasekaran</collectorName>
|
|
leg., ZSI-SRC/I/E/654
|
|
</materialsCitation>
|
|
.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="167">
|
|
<typeStatus>Paratypes</typeStatus>
|
|
</emphasis>
|
|
:
|
|
<specimenCount type="larva">5 larvae</specimenCount>
|
|
,
|
|
<materialsCitation collectingDate="2020-10-28" collectorName="Pandiarajan Srinivasan, Isack Rajasekaran" country="India" county="Theni district" elevation="632" latitude="10.0809" location="Kottakudi River" longLatPrecision="7" longitude="77.2552" municipality="Kurangani" specimenCount="5" specimenCount-larva="5" stateProvince="Tamil Nadu" typeStatus="paratype">
|
|
ibid.,
|
|
<collectingDate value="2020-10-28">28.x.2020</collectingDate>
|
|
,
|
|
<collectorName>Pandiarajan Srinivasan</collectorName>
|
|
&
|
|
<collectorName>Isack Rajasekaran</collectorName>
|
|
leg., ZSI-SRC/I/E/655 (
|
|
<specimenCount type="larva">1 larva</specimenCount>
|
|
), AMC ZN 237 (
|
|
<specimenCount type="larva">4 larvae</specimenCount>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="etymology">
|
|
<paragraph pageId="0" pageNumber="167">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">The new species is named in honour of Dr Carlos Molineri of Argentina, who contributed significantly to the study of mayflies.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="description">
|
|
<paragraph pageId="0" pageNumber="167">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Mature larva</emphasis>
|
|
.
|
|
</emphasis>
|
|
Body length 4.7-4.9 mm; cerci length subequal to body length. General coloration of the dorsal side of head dirty yellow, with dark brown to blackish maculations; thorax and abdomen dark brown to blackish, with dirty yellow maculations (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9A-D</figureCitation>
|
|
). Ventral side of body yellowish to light brown. Dorsal surface of femora with two blackish longitudinal stripes one medial along ridge and one along outer margin; also three (proximal, medial, and distal) black spots with indistinct borders along medial ridge; proximal and medial spots divided in two parts by brownish bands (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11A-C</figureCitation>
|
|
). Dorsal part of male eyes dark brown to black (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9D</figureCitation>
|
|
). Abdominal terga I-VIII with pair of submedian yellowish spots. Anterior part of tergum X yellowish (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167" start="Figure 9" startId="F9">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 9.</emphasis>
|
|
Larva of
|
|
<taxonomicName authorityName="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes molinerii" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="molinerii">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes molinerii</emphasis>
|
|
</taxonomicName>
|
|
Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A</emphasis>
|
|
total dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="167">B</emphasis>
|
|
total ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="167">C</emphasis>
|
|
head, thorax and abdomen, dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="167">D</emphasis>
|
|
head, dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="167">E</emphasis>
|
|
row of setae at outer margin of head
|
|
<emphasis bold="true" pageId="0" pageNumber="167">F</emphasis>
|
|
head and thorax, dorso-lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="167">G</emphasis>
|
|
tarsal claw
|
|
<emphasis bold="true" pageId="0" pageNumber="167">H</emphasis>
|
|
gill II
|
|
<emphasis bold="true" pageId="0" pageNumber="167">I</emphasis>
|
|
gill III
|
|
<emphasis bold="true" pageId="0" pageNumber="167">J</emphasis>
|
|
gill IV
|
|
<emphasis bold="true" pageId="0" pageNumber="167">K</emphasis>
|
|
gill V. Abbreviations: white arrows show tubercles.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="167">Head</emphasis>
|
|
with pair of occipital tubercles (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9D</figureCitation>
|
|
). Genae moderately developed. Lateral margin of head capsule from eye to labrum insertion with row of long, forked near the base, stout setae with pointed apices (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9E</figureCitation>
|
|
). Antennae length 1.25 times head width, flagellum with 11 segments. Head covered with scattered short hair-like setae and short stout setae with slightly divergent margins.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Mouthparts</emphasis>
|
|
.
|
|
</emphasis>
|
|
Labrum compact, width/length ratio 2.51-2.53; with smooth anterior emargination; dorsal surface with transversal row of scattered, stout, hair-like setae (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">10C, D</figureCitation>
|
|
); anterior area and margin of labrum densely covered with differently sized feathered setae. Mandibles slender with long, stout, hair-like seta in the middle of the outer margin (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">10A, B</figureCitation>
|
|
). Number of teeth of both mandibular outer incisors cannot be determined precisely due to their wear in type specimens. Right mandible inner incisor with two teeth; prostheca reduced, with the appearance of a cluster of thin setae; small row of five long, stout, hair-like setae below mola and some short setae above mola. Left mandible inner incisor with two teeth inserted transversely, one smaller and rounded and other one larger and rounded; prostheca small; no setae below and above mola. Maxilla (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">10F, G</figureCitation>
|
|
) slender, shape of canine impossible to determine (completely worn); two indented dentisetae and three long setae on inner apical part and cluster of long, simple setae at crown; inner margin at the base of lacinia, with two feathered, long setae; maxillary palp highly reduced to protuberance. Hypopharynx (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">10E, H</figureCitation>
|
|
) with long, feathered setae on the rounded apexes of superlinguae and very short setae on lingua. Labial palp (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, B mandibles C labrum D stout setae of transversal row of labrum E hypopharynx F maxilla G apical part of maxilla H superlingua I labium J labial palp." figureDoi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167">10I, J</figureCitation>
|
|
) three-segmented, slightly constricted towards apex; articulation between segments clearly visible; segment III elongate and rounded apically, length/width ratio at base 2.0-2.1. Surface and margins of segments I and II covered with scattered long, stout, hair-like setae; segment III bare. Submentum well developed laterally. Glossae and paraglossae short and broad, rounded apically, their apexes densely covered with differently sized, feathered, stout setae; outer margins of paraglossae covered with long feathered setae.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1113.85448.figure10" httpUri="https://binary.pensoft.net/fig/716508" pageId="0" pageNumber="167" start="Figure 10" startId="F10">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 10.</emphasis>
|
|
Larva of
|
|
<taxonomicName authorityName="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes molinerii" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="molinerii">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes molinerii</emphasis>
|
|
</taxonomicName>
|
|
Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
mandibles
|
|
<emphasis bold="true" pageId="0" pageNumber="167">C</emphasis>
|
|
labrum
|
|
<emphasis bold="true" pageId="0" pageNumber="167">D</emphasis>
|
|
stout setae of transversal row of labrum
|
|
<emphasis bold="true" pageId="0" pageNumber="167">E</emphasis>
|
|
hypopharynx
|
|
<emphasis bold="true" pageId="0" pageNumber="167">F</emphasis>
|
|
maxilla
|
|
<emphasis bold="true" pageId="0" pageNumber="167">G</emphasis>
|
|
apical part of maxilla
|
|
<emphasis bold="true" pageId="0" pageNumber="167">H</emphasis>
|
|
superlingua
|
|
<emphasis bold="true" pageId="0" pageNumber="167">I</emphasis>
|
|
labium
|
|
<emphasis bold="true" pageId="0" pageNumber="167">J</emphasis>
|
|
labial palp.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Thorax</emphasis>
|
|
.
|
|
</emphasis>
|
|
Pronotum with three pairs of tubercles: SMs, SLs, and Ls; tubercles with a few short, rounded setae. Mesonotum with three pairs of tubercles: two pairs of SMMs, a pair of LAs, and unpaired MP (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9C, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Forefemur broad, ca 1.3 times longer than wide (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11A, D</figureCitation>
|
|
); outer margin covered with a row of long, stout, hair-like setae (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11G</figureCitation>
|
|
); submarginal row of setae composed of scattered, short, stout setae with rounded apices; basal half of inner margin with row of long, stout, hair-like setae; distal half almost without setae. Transverse row on the dorsal surface made of about 30 long, pointed apically stout setae (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11E, F</figureCitation>
|
|
). Dorsal surface of fore femur covered with scattered, short, stout setae and thin, hair-like setae. Dorsal surface of fore tibia with solitary hair-like setae and oblique regular row of long, stout, hair-like setae; outer margins of tibia with regular row of long, stout, hair-like setae.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">Figure 11.</emphasis>
|
|
Larva of
|
|
<taxonomicName authorityName="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes molinerii" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="molinerii">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes molinerii</emphasis>
|
|
</taxonomicName>
|
|
Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes
|
|
<emphasis bold="true" pageId="0" pageNumber="167">A, D</emphasis>
|
|
fore femur
|
|
<emphasis bold="true" pageId="0" pageNumber="167">B</emphasis>
|
|
middle femur
|
|
<emphasis bold="true" pageId="0" pageNumber="167">C</emphasis>
|
|
hind femur
|
|
<emphasis bold="true" pageId="0" pageNumber="167">E, F</emphasis>
|
|
stout setae of transversal row on forefemur
|
|
<emphasis bold="true" pageId="0" pageNumber="167">G</emphasis>
|
|
outer margin of fore femur
|
|
<emphasis bold="true" pageId="0" pageNumber="167">H</emphasis>
|
|
abdomen
|
|
<emphasis bold="true" pageId="0" pageNumber="167">I</emphasis>
|
|
eggs, light microscopy
|
|
<emphasis bold="true" pageId="0" pageNumber="167">J</emphasis>
|
|
egg, SEM microscopy
|
|
<emphasis bold="true" pageId="0" pageNumber="167">K</emphasis>
|
|
cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">H</emphasis>
|
|
); 0.02 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">J</emphasis>
|
|
); 0.01 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="167">K</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Middle and hind femora, in contrast to fore femur, more slender, ca 1.8-2 times longer than wide, with denser submarginal row of short, stout setae (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11B, C</figureCitation>
|
|
). Outer and inner margins covered with a regular row of long, stout, hair-like setae. Outer margin of middle and hind tibiae with a regular row of long, stout, hair-like setae. Setation of dorsal surface of middle and hind tibiae similar to those of fore leg.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Tarsal claw moderately hooked, bearing 3-6 medial denticles, 1-2 subapical denticles (if two subapical denticles present, they are situated each on opposite sides of claw) and a row of 3-4 subapical setae on dorsal and ventral sides (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Abdomen</emphasis>
|
|
.
|
|
</emphasis>
|
|
All terga with median tubercles that bear short, stout setae with slightly divergent margins. Median tubercles moderately developed on terga IV-VIII, and slightly marked on terga I-III, IX, and X (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11H</figureCitation>
|
|
). Posterolateral projections moderately developed on segments VI-IX, and slightly marked on segments II-V. Submedian and sublateral areas of terga VI and VII with scattered stout setae with divided apices and a few small, rounded stout setae.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Gills on segments II-V (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy, Isack, sp. nov., paratypes A total dorsal view B total ventral view C head, thorax and abdomen, dorsal view D head, dorsal view E row of setae at outer margin of head F head and thorax, dorso-lateral view G tarsal claw H gill II I gill III J gill IV K gill V. Abbreviations: white arrows show tubercles." figureDoi="10.3897/zookeys.1113.85448.figure9" httpUri="https://binary.pensoft.net/fig/716507" pageId="0" pageNumber="167">9H-K</figureCitation>
|
|
); gill II with dorsal lamella operculate, oval and with entire margin; gills III-V with dorsal lamella incised medially.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">Cerci length subequal to the body length; posterior margin of proximal half segments with elongate, stout setae with rounded apices; posterior margin of distal half segments with long, spine-like setae on the lateral margins; length of the stout setae less than length of corresponding segment. Paracercus absent.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="167">Egg.</emphasis>
|
|
Ovoid, ca 100-110
|
|
<normalizedToken originalValue="μm">μm</normalizedToken>
|
|
long, with numerous micropyles. Egg with one polar cap, on the opposite pole with a cluster of 18-20 spines (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larva of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov., paratypes A, D fore femur B middle femur C hind femur E, F stout setae of transversal row on forefemur G outer margin of fore femur H abdomen I eggs, light microscopy J egg, SEM microscopy K cluster of spines on pole of egg. Abbreviations: sII - abdominal segment II, sX - abdominal segment X. Scale bars: 0.2 mm (H); 0.02 mm (J); 0.01 mm (K)." figureDoi="10.3897/zookeys.1113.85448.figure11" httpUri="https://binary.pensoft.net/fig/716509" pageId="0" pageNumber="167">11I-K</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<emphasis bold="true" pageId="0" pageNumber="167">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Winged stages</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="distribution">
|
|
<paragraph pageId="0" pageNumber="167">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">Western Ghats (Tamil Nadu, India).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="habitat">
|
|
<paragraph pageId="0" pageNumber="167">Habitats.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
The larvae of
|
|
<taxonomicName genus="D." lsidName="D. molinerii" pageId="0" pageNumber="167" rank="species" species="molinerii">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">D. molinerii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. inhabit cobble and pebble substrates of rivers with a strong current (Fig.
|
|
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Type habitats A, B, D, E of new species and cages (Martynov's construction) C used for Dudgeodes selvakumari Martynov & Palatov, sp. nov. winged stages rearing A, B, D type habitat of Dudgeodes selvakumari Martynov & Palatov, sp. nov. C Martynov-designed grow nets for mayfly winged stages rearing E type habitat of Dudgeodes molinerii Sivaruban, Martynov, Srinivasan, Barathy & Isack, sp. nov. and Teloganodes barathyae Sivaruban, Martynov, Srinivasan & Isack, sp. nov. Abbreviations: arrows show microhabitats with the highest density of a new species' larvae." figureDoi="10.3897/zookeys.1113.85448.figure16" httpUri="https://binary.pensoft.net/fig/716514" pageId="0" pageNumber="167">16E</figureCitation>
|
|
), where there is no significant anthropogenic stress. Water temperatures range between 20 and 22 °C and pH ranges between 7.1 and 7.4. This species was caught with other mayflies such as
|
|
<taxonomicName authorityName="Srinivasan, Sivaruban, Barathy & Isack" authorityYear="2022" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" lsidName="Clypeocaenis malzacheri" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="malzacheri">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Clypeocaenis malzacheri</emphasis>
|
|
</taxonomicName>
|
|
Srinivasan, Sivaruban, Barathy & Isack, 2022 (
|
|
<taxonomicName class="Insecta" family="Caenidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="family">Caenidae</taxonomicName>
|
|
),
|
|
<taxonomicName authorityName="Sivaruban, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" lsidName="Nigrobaetis klugei" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="klugei">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Nigrobaetis klugei</emphasis>
|
|
</taxonomicName>
|
|
Sivaruban, Srinivasan, Barathy & Isack, 2022 (
|
|
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
|
),
|
|
<taxonomicName authorityName="Peters & Edmunds" authorityYear="1970" class="Insecta" family="Leptophlebiidae" genus="Notophlebia" kingdom="Animalia" lsidName="Notophlebia" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Notophlebia</emphasis>
|
|
</taxonomicName>
|
|
sp. (
|
|
<taxonomicName class="Insecta" family="Leptophlebiidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
|
|
), and
|
|
<taxonomicName authorityName="Muller-Liebenau & Hubbard" authorityYear="1985" baseAuthorityName="Muller-Liebenau & Hubbard" baseAuthorityYear="1985" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" lsidName="Tenuibaetis frequentus" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="frequentus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Tenuibaetis frequentus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
|
|
& Hubbard, 1985) (
|
|
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="167" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="167">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
<taxonomicName authorityName="Sivaruban, Martynov, Srinivasan, Barathy & Isack" authorityYear="2022" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes molinerii" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="molinerii">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes molinerii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from other
|
|
<taxonomicName authorityName="Sartori" authorityYear="2008" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes</emphasis>
|
|
</taxonomicName>
|
|
species by the following combination of characters.
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Larva</emphasis>
|
|
: (i) dorsal part of male eyes dark brown to black; (ii) head with pair of small occipital tubercles; (iii) antennae length 1.25 times head width, flagellum with 11 segments; (iv) labrum with transversal row of scattered, stout, hair-like setae; (v) forefemur with transverse row of about 30 long, apically pointed, stout setae; (vi) tarsal claw bearing 3-6 medial denticles, and 1-2 subapical denticles (if two, they are on opposite sides of claw), and 3-4 subapical setae on dorsal and ventral sides; (vii) pronotum bears three pairs of tubercles: SMs, SLs, and Ls; mesonotum bears three pairs of tubercles: two pairs of SMMs, a pair of LAs, and unpaired MP; (viii) median tubercles moderately developed on terga IV-VIII, and slightly marked on terga I-III, IX and X; (ix) posterolateral projections moderately developed on segments VI-IX, and slightly marked on segments II-V.
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Egg</emphasis>
|
|
: (i) egg with cluster of 18-20 spines present on pole opposite to polar cap; (ii) surface without microgranules.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="167">
|
|
Larval stage of this new species can be easily distinguished from other Indian
|
|
<taxonomicName authorityName="Sartori" authorityYear="2008" class="Insecta" family="Teloganodidae" genus="Dudgeodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dudgeodes" order="Ephemeroptera" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="167">Dudgeodes</emphasis>
|
|
</taxonomicName>
|
|
by: (i) presence of tubercles on head; (ii) number of tubercles on pro- and mesonotum; (iii) shape of femora; (iv) setation of forefemur; (v) size and shape median tubercles on abdomen.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |