treatments-xml/data/3F/08/B2/3F08B26E3F183848FF57FD6AFABAE28E.xml
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<mods:title id="7FBE2BA3335A775220DEBEB52BCF6839">New species of Omalodes and redefinition of the tribe Omalodini (Coleoptera: Histeridae: Histerinae)</mods:title>
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<taxonomicName id="70A178FB3F18384DFF57FD6AFCF0E38E" ID-CoL="7W579" authority="Leivas &amp; Degallier" authorityName="Leivas &amp; Degallier" box="[151,814,724,750]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov." subGenus="Omalodes">
<emphasis id="85D5DF6A3F18384DFF57FD6AFECEE38E" bold="true" box="[151,272,724,750]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F18384DFEE0FD6AFE46E38E" bold="true" box="[288,408,724,750]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
)
<emphasis id="85D5DF6A3F18384DFE6AFD6AFDE2E38E" bold="true" box="[426,572,724,750]" italics="true" pageId="2" pageNumber="111">atacamanus</emphasis>
Leivas &amp; Degallier
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<taxonomicNameLabel id="9EE662113F18384DFCF5FD6AFC4DE38E" box="[821,915,724,750]" pageId="2" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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locality.
</emphasis>
<collectingCountry id="CFB643E83F18384DFEFCFCA4FEA2E253" box="[316,380,794,819]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
, Província de Antofagasta, Taltal (Paposo).
</paragraph>
<paragraph id="B71E03783F18384DFF07FC80FEC7E763" blockId="2.[151,1437,794,2010]" pageId="2" pageNumber="111">
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<typeStatus id="681ABDDA3F18384DFF07FC80FEDCE237" box="[199,258,830,855]" pageId="2" pageNumber="111">Type</typeStatus>
material.
<typeStatus id="681ABDDA3F18384DFEBBFC80FE32E237" box="[379,492,830,855]" pageId="2" pageNumber="111" type="holotype">Holotype</typeStatus>
:
</emphasis>
(♂) [“
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II REGION/Tal-Tal Paposo/
<date id="C31F25B83F18384DFC11FC81FB52E237" box="[977,1164,830,855]" pageId="2" pageNumber="111" value="1997-10-24">24 Octubre 1997</date>
/Leg. V.M. Diéguez M.” “
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/V.M. Diéguez M.” “
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<emphasis id="85D5DF6A3F18384DFDF9FCDCFD74E21B" box="[569,682,866,891]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
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<emphasis id="85D5DF6A3F18384DFD7CFCDCFCA6E21B" box="[700,888,866,891]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
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)/
<taxonomicName id="70A178FB3F18384DFC49FCDCFB67E21B" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" box="[905,1209,866,891]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFC49FCDCFBD0E21B" box="[905,1038,866,891]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(
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)
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/ Det. G. Arriagada 2001”] (
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).
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(15)
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(
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) [“
<collectingCountry id="CFB643E83F18384DFD4AFC39FD3CE2FF" box="[650,738,903,927]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
III Región/Bahia Esmeralda/
<date id="C31F25B83F18384DFBFAFC38FB77E2FF" box="[1082,1193,902,927]" pageId="2" pageNumber="111" value="1999-10-23">23.X.1999</date>
/leg. Guildo Castillo” “
<taxonomicName id="70A178FB3F18384DFF63FC14FECBE2A3" box="[163,277,938,963]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFF63FC14FECBE2A3" box="[163,277,938,963]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
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<emphasis id="85D5DF6A3F18384DFEE6FC14FE3CE2A3" box="[294,482,938,963]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFE33FC14FD0BE2A3" authority="Lewis" authorityName="Lewis" box="[499,725,938,963]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFE33FC14FDA9E2A3" box="[499,631,938,963]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(Lewis)
</taxonomicName>
/ Det. G. Arriagada 2008”] (
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); same data and labels (1 ♀
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); (1 ♀
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); (1 ♀
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); (
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) [“
<collectingCountry id="CFB643E83F18384DFCCDFC71FCBBE287" box="[781,869,975,999]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
II REGION/Taltal/
<date id="C31F25B83F18384DFB97FC70FB67E287" box="[1111,1209,974,999]" pageId="2" pageNumber="111" value="85-10-22">22.10.85</date>
/leg. G. Arriagada” “
<collectionCode id="D1B09BBD3F18384DFF61FC4DFE9BE56B" box="[161,325,1011,1035]" pageId="2" pageNumber="111">COLLECION</collectionCode>
/V.M. Diéguez M.” “
<taxonomicName id="70A178FB3F18384DFD81FC4CFD6CE56B" box="[577,690,1010,1035]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFD81FC4CFD6CE56B" box="[577,690,1010,1035]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/
<taxonomicName id="70A178FB3F18384DFD7BFC4CFCE1E56B" box="[699,831,1010,1035]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFD7BFC4CFCE1E56B" box="[699,831,1010,1035]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
</taxonomicName>
/G. ARRIAGADA DET. 1997”] (
<collectionCode id="D1B09BBD3F18384DFB10FC4DFAE1E56B" box="[1232,1343,1011,1035]" pageId="2" pageNumber="111">MNHNC</collectionCode>
); (1 ♀) [“
<collectingCountry id="CFB643E83F18384DFF67FBA9FF21E54F" box="[167,255,1047,1071]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
II REGION/Taltal/
<date id="C31F25B83F18384DFE3BFBA8FD83E54F" box="[507,605,1046,1071]" pageId="2" pageNumber="111" value="85-10-22">22.10.85</date>
/leg. G. Arriagada” “
<collectionCode id="D1B09BBD3F18384DFCAAFBA9FBD0E54F" box="[874,1038,1047,1071]" pageId="2" pageNumber="111">COLLECION</collectionCode>
/V.M. Diéguez M.” “
<taxonomicName id="70A178FB3F18384DFAE3FBA8FA4BE54F" box="[1315,1429,1046,1071]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFAE3FBA8FA4BE54F" box="[1315,1429,1046,1071]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/ (
<taxonomicName id="70A178FB3F18384DFF60FB84FE82E533" box="[160,348,1082,1107]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFF60FB84FE82E533" box="[160,348,1082,1107]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFEADFB84FD49E533" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" box="[365,663,1082,1107]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFEADFB84FE2FE533" box="[365,497,1082,1107]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(
<bibRefCitation id="D3307E893F18384DFDC2FB84FD51E533" author="Lewis" box="[514,655,1082,1107]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)
</taxonomicName>
/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFC16FB85FB9BE533" box="[982,1093,1083,1107]" pageId="2" pageNumber="111">MNHNC</collectionCode>
); (
<specimenCount id="A1A7C8F13F18384DFBA2FB85FB50E533" box="[1122,1166,1083,1107]" pageId="2" pageNumber="111" type="generic">2 ♂</specimenCount>
) [“
<collectingCountry id="CFB643E83F18384DFB70FB85FAD6E533" box="[1200,1288,1083,1107]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
II REGION/ Paposo/
<date id="C31F25B83F18384DFF31FBE0FE8BE517" box="[241,341,1118,1143]" pageId="2" pageNumber="111" value="92-05-10">05.10.92</date>
/leg. G. Castillo” “
<collectionCode id="D1B09BBD3F18384DFDEDFBE1FD0FE517" box="[557,721,1119,1143]" pageId="2" pageNumber="111">COLLECION</collectionCode>
/V.M. Diéguez M.” “
<taxonomicName id="70A178FB3F18384DFC08FBE0FBE7E517" box="[968,1081,1118,1143]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFC08FBE0FBE7E517" box="[968,1081,1118,1143]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFB8AFBE0FAD8E517" box="[1098,1286,1118,1143]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFB8AFBE0FAD8E517" box="[1098,1286,1118,1143]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFAD8FBE0FEEBE5FB" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFAD8FBE0FA42E517" box="[1304,1436,1118,1143]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(
<bibRefCitation id="D3307E893F18384DFF5FFB3CFEF3E5FB" author="Lewis" box="[159,301,1154,1179]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)
</taxonomicName>
/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFDB9FB3DFD16E5FB" box="[633,712,1155,1179]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="111">DZUP</collectionCode>
); (2 ♀) [“
<collectingCountry id="CFB643E83F18384DFCF6FB3DFC50E5FB" box="[822,910,1155,1179]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
II REGION/Tal-Tal Paposo/
<date id="C31F25B83F18384DFB18FB3CFA4DE5FB" box="[1240,1427,1154,1179]" pageId="2" pageNumber="111" value="1997-10-24">24.Octubre 1997</date>
/ Leg. V.M. Diéguez M.” “
<collectionCode id="D1B09BBD3F18384DFE08FB19FDB3E5DF" box="[456,621,1191,1215]" pageId="2" pageNumber="111">COLLECION</collectionCode>
/V.M. Diéguez M.” “
<taxonomicName id="70A178FB3F18384DFCA6FB18FC09E5DF" box="[870,983,1190,1215]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFCA6FB18FC09E5DF" box="[870,983,1190,1215]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFC29FB18FB7BE5DF" box="[1001,1189,1190,1215]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFC29FB18FB7BE5DF" box="[1001,1189,1190,1215]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFB76FB18FF09E583" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFB76FB18FAE5E5DF" box="[1206,1339,1190,1215]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(
<bibRefCitation id="D3307E893F18384DFA8FFB18FF11E583" author="Lewis" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)
</taxonomicName>
/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFDD3FB75FDBCE583" box="[531,610,1227,1251]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="111">DZUP</collectionCode>
); (1 ♀) [“
<collectingCountry id="CFB643E83F18384DFD0BFB75FCFCE583" box="[715,802,1227,1251]" name="Chile" pageId="2" pageNumber="111">CHILE</collectingCountry>
II REGION/Tal-Tal Paposo/
<date id="C31F25B83F18384DFBA8FB74FAFDE583" box="[1128,1315,1226,1251]" pageId="2" pageNumber="111" value="1997-10-24">24.Octubre 1997</date>
/Leg. V.M. Diéguez M.” “
<collectionCode id="D1B09BBD3F18384DFEF9FB51FE03E467" box="[313,477,1263,1287]" pageId="2" pageNumber="111">COLLECION</collectionCode>
/V.M. Diéguez M.” “
<taxonomicName id="70A178FB3F18384DFD05FB50FCE8E467" box="[709,822,1262,1287]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFD05FB50FCE8E467" box="[709,822,1262,1287]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFC88FB50FBDAE467" box="[840,1028,1262,1287]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFC88FB50FBDAE467" box="[840,1028,1262,1287]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFBD5FB50FAE3E467" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" box="[1045,1341,1262,1287]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFBD5FB50FB44E467" box="[1045,1178,1262,1287]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
(
<bibRefCitation id="D3307E893F18384DFB68FB50FAEBE467" author="Lewis" box="[1192,1333,1262,1287]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)
</taxonomicName>
/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFEB9FAADFE0DE44B" box="[377,467,1299,1323]" pageId="2" pageNumber="111">VMDC</collectionCode>
); (1 ♀) [“Taltal/S. la Quinta/
<date id="C31F25B83F18384DFCFAFAACFC72E44B" box="[826,940,1298,1323]" pageId="2" pageNumber="111" value="1985-05-10">5.10.1985</date>
/Coll. M. Elgueta” “Bajo cactus/
<taxonomicName id="70A178FB3F18384DFAF0FAACFF35E42F" class="Magnoliopsida" family="Cactaceae" genus="Copiapoa" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="111" phylum="Tracheophyta" rank="species" species="cinerea">Copiapoa cinerea</taxonomicName>
” “
<collectionCode id="D1B09BBD3F18384DFEC7FA88FE76E42F" box="[263,424,1334,1359]" pageId="2" pageNumber="111">COLLECIÓN</collectionCode>
<collectionCode id="D1B09BBD3F18384DFE72FA88FDD6E42F" LSID="urn:lsid:biocol.org:col:34988" box="[434,520,1334,1359]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="111">MNHN</collectionCode>
/
<collectingCountry id="CFB643E83F18384DFDD0FA88FD92E42F" box="[528,588,1334,1359]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
- SANTIAGO” “
<taxonomicName id="70A178FB3F18384DFCD9FA88FC54E42F" box="[793,906,1334,1359]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFCD9FA88FC54E42F" box="[793,906,1334,1359]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFC5CFA88FB86E42F" box="[924,1112,1334,1359]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFC5CFA88FB86E42F" box="[924,1112,1334,1359]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFBA9FA88FB33E42F" box="[1129,1261,1334,1359]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFBA9FA88FB33E42F" box="[1129,1261,1334,1359]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
</taxonomicName>
/(
<bibRefCitation id="D3307E893F18384DFB3EFA88FA55E42F" author="Lewis" box="[1278,1419,1334,1359]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFE08FAE5FDE9E413" box="[456,567,1371,1395]" pageId="2" pageNumber="111">MNHNC</collectionCode>
); (1 ♀) [“
<collectingCountry id="CFB643E83F18384DFD65FAE4FD38E413" box="[677,742,1370,1395]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
Atacama/Paposo/La Rinconada/
<date id="C31F25B83F18384DFB9DFAE4FB05E413" box="[1117,1243,1370,1395]" pageId="2" pageNumber="111" value="1983-10-10">10.10.1983</date>
/leg. M. Elgueta” “
<collectionCode id="D1B09BBD3F18384DFF60FAC0FE9FE4F7" box="[160,321,1406,1431]" pageId="2" pageNumber="111">COLLECIÓN</collectionCode>
<collectionCode id="D1B09BBD3F18384DFE8DFAC0FE7DE4F7" LSID="urn:lsid:biocol.org:col:34988" box="[333,419,1406,1431]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="111">MNHN</collectionCode>
/
<collectingCountry id="CFB643E83F18384DFE6BFAC0FE39E4F7" box="[427,487,1406,1431]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
- SANTIAGO” “
<taxonomicName id="70A178FB3F18384DFD7AFAC0FCF5E4F7" box="[698,811,1406,1431]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFD7AFAC0FCF5E4F7" box="[698,811,1406,1431]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFCFCFAC0FC26E4F7" box="[828,1016,1406,1431]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFCFCFAC0FC26E4F7" box="[828,1016,1406,1431]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFBC9FAC0FB53E4F7" box="[1033,1165,1406,1431]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFBC9FAC0FB53E4F7" box="[1033,1165,1406,1431]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
</taxonomicName>
/(
<bibRefCitation id="D3307E893F18384DFB5EFAC0FAF3E4F7" author="Lewis" box="[1182,1325,1406,1431]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFEB1FA1DFE1BE4DB" box="[369,453,1443,1467]" pageId="2" pageNumber="111">CEMT</collectionCode>
); (
<specimenCount id="A1A7C8F13F18384DFE24FA1DFDCCE4DB" box="[484,530,1443,1467]" pageId="2" pageNumber="111" type="generic">2 ♂</specimenCount>
) [“
<collectingCountry id="CFB643E83F18384DFDF6FA1CFDA9E4DB" box="[566,631,1442,1467]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
Anfogasta/Tatal/
<date id="C31F25B83F18384DFC81FA1CFC60E4DB" box="[833,958,1442,1467]" pageId="2" pageNumber="111" value="1957-09-19">19.09.1957</date>
/leg. G. Kuschel” “
<collectionCode id="D1B09BBD3F18384DFB53FA1CFAEAE4DB" box="[1171,1332,1442,1467]" pageId="2" pageNumber="111">COLLECIÓN</collectionCode>
<collectionCode id="D1B09BBD3F18384DFAFFFA1DFA4AE4DB" LSID="urn:lsid:biocol.org:col:34988" box="[1343,1428,1443,1467]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="111">MNHN</collectionCode>
/
<collectingCountry id="CFB643E83F18384DFF57FA78FF0AE4BF" box="[151,212,1478,1503]" name="Chile" pageId="2" pageNumber="111">Chile</collectingCountry>
- SANTIAGO” “
<taxonomicName id="70A178FB3F18384DFE7EFA78FDEEE4BF" box="[446,560,1478,1503]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFE7EFA78FDEEE4BF" box="[446,560,1478,1503]" italics="true" pageId="2" pageNumber="111">Omalodes</emphasis>
</taxonomicName>
/(
<taxonomicName id="70A178FB3F18384DFD80FA78FD22E4BF" box="[576,764,1478,1503]" class="Insecta" family="Histeridae" genus="Diplogrammicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F18384DFD80FA78FD22E4BF" box="[576,764,1478,1503]" italics="true" pageId="2" pageNumber="111">Diplogrammicus</emphasis>
</taxonomicName>
)/
<taxonomicName id="70A178FB3F18384DFCCEFA78FC4CE4BF" box="[782,914,1478,1503]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="85D5DF6A3F18384DFCCEFA78FC4CE4BF" box="[782,914,1478,1503]" italics="true" pageId="2" pageNumber="111">intermedius</emphasis>
</taxonomicName>
/(
<bibRefCitation id="D3307E893F18384DFC63FA78FBE4E4BF" author="Lewis" box="[931,1082,1478,1503]" pageId="2" pageNumber="111" refString="Lewis, G. (1907) On new species of Histeridae and notices of others. The Annals and Magazine of Natural History, 7 (20), 480 - 484. http: // dx. doi. org / 10.1080 / 00222930709487371" type="journal article" year="1907">Lewis, 1907</bibRefCitation>
)/ Det. G. Arriagada 2001”] (
<collectionCode id="D1B09BBD3F18384DFF5EFA55FED3E763" box="[158,269,1515,1539]" pageId="2" pageNumber="111">MNHNC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FFBB50F33F18384DFF07F9B0FA5AE7F6" pageId="2" pageNumber="111" type="diagnosis">
<paragraph id="B71E03783F18384DFF07F9B0FA5AE7F6" blockId="2.[151,1437,794,2010]" pageId="2" pageNumber="111">
<emphasis id="85D5DF6A3F18384DFF07F9B0FEE4E747" bold="true" box="[199,314,1550,1575]" pageId="2" pageNumber="111">Diagnosis</emphasis>
. Frons flat (
<figureCitation id="2F9A1FFD3F18384DFE06F9B0FDD7E747" box="[454,521,1550,1575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
D); marginal stria of prosternal lobe absent (
<figureCitation id="2F9A1FFD3F18384DFBC6F9B0FB97E747" box="[1030,1097,1550,1575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
E, F); posterior margin of the elytra with longitudinal strioles (
<figureCitation id="2F9A1FFD3F18384DFDC9F98CFD92E72B" box="[521,588,1586,1611]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="2" pageNumber="111">Fig. 3</figureCitation>
E); lateral metaventral stria oblique; all tibiae with a row of setae on outer submarginal region (
<figureCitation id="2F9A1FFD3F18384DFE40F9E8FE1AE70F" box="[384,452,1622,1647]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="2" pageNumber="111">Fig. 3</figureCitation>
BD); profemora with sparse setae on the posterior surface (
<figureCitation id="2F9A1FFD3F18384DFBB9F9E8FB62E70F" box="[1145,1212,1622,1647]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
EF); propygidium and pygidium with punctuation similar to the dorsum (
<figureCitation id="2F9A1FFD3F18384DFD3BF9C3FCE3E7F6" box="[763,829,1661,1686]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="2" pageNumber="111">Fig. 3</figureCitation>
E); 9th tergite without lateral projections (Fig. 4 M).
</paragraph>
</subSubSection>
<subSubSection id="FFBB50F33F183849FF07F91FFB45E603" lastPageId="6" lastPageNumber="115" pageId="2" pageNumber="111" type="description">
<paragraph id="B71E03783F18384AFF07F91FFE34E686" blockId="2.[151,1437,794,2010]" lastBlockId="5.[151,1436,1841,2022]" lastPageId="5" lastPageNumber="114" pageId="2" pageNumber="111">
<emphasis id="85D5DF6A3F18384DFF07F91FFE8FE7DA" bold="true" box="[199,337,1697,1722]" pageId="2" pageNumber="111">Description</emphasis>
. Length (pronotum+elytra)
<quantity id="7059AE9D3F18384DFD4FF91FFCC7E7D9" box="[655,793,1697,1722]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.6" metricValueMin="5.3" pageId="2" pageNumber="111" unit="mm" value="5.949999999999999" valueMax="6.6" valueMin="5.3">5.36.6 mm</quantity>
, elytral width (humeral region):
<quantity id="7059AE9D3F18384DFB4CF91CFAC9E7D9" box="[1164,1303,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.7" metricValueMin="3.7" pageId="2" pageNumber="111" unit="mm" value="4.2" valueMax="4.7" valueMin="3.7">3.74.7 mm</quantity>
. Color dark or dark-brown. Body shape subrectangular, glabrous, covered by ground punctuation (
<figureCitation id="2F9A1FFD3F18384DFBBFF97BFB11E7BE" box="[1151,1231,1733,1758]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Figs. 2</figureCitation>
AB). Frons and clypeus flat; frontal stria inwardly curved and usually complete; supraorbital stria usually absent or rudimentary; frontoclypeal suture weakly marked only laterally (
<figureCitation id="2F9A1FFD3F18384DFD2DF8B3FCEAE646" box="[749,820,1805,1830]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
D); postoccipital stria complete not setose; fovea of postgena present. Antennal scape basally slender (
<figureCitation id="2F9A1FFD3F18384DFD0FF88FFCCCE62A" box="[719,786,1841,1866]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
D) and with subrectangular elevation at the apical region (
<figureCitation id="2F9A1FFD3F18384DFF5FF8EBFF3AE60E" box="[159,228,1877,1902]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="2" pageNumber="111">Fig. 2</figureCitation>
D, see also
<figureCitation id="2F9A1FFD3F18384DFEB9F8EBFE63E60E" box="[377,445,1877,1902]" captionStart="FIGURE 6. A" captionStartId="8.[151,250,1857,1879]" captionTargetBox="[151,1435,375,1835]" captionTargetId="figure@8.[151,1436,375,1835]" captionTargetPageId="8" captionText="FIGURE 6. A. Ebonius politus Lewis, 1885: prothorax in ventral view; B. Omalodes (Omalodes) omega (Kirby, 1818): prothorax in ventral view; C. Omalodes (Diplogrammicus) marseuli Schmidt, 1889: prothorax in ventral view; D. Ebonius politus: meso- and metathorax and first abdominal ventrite; E. Omalodes (Cornillus) binodulus (Lewis, 1910), meso- and metathorax and first abdominal ventrite; F. Omalodes (Omalodes) omega: antennal scape." httpUri="https://zenodo.org/record/236798/files/figure.png" pageId="2" pageNumber="111">Fig. 6</figureCitation>
F); fifth and sixth segments of the funicle not expanded laterally; antennal club with pseudo-sutures slightly inwardly arcuate and with a distinct apical sensorial area. Palpifer maxillary with robust setae on the external margin and without setae on the internal margin; lacinial hook absent. Submentum subpentagonal and with long setae on the surface; anterior margin of the mentum medially emarginate. Prothorax
<emphasis id="85D5DF6A3F1F384AFF57F88FFEC6E626" bold="true" box="[151,280,1841,1863]" pageId="5" pageNumber="114">FIGURE 4.</emphasis>
<taxonomicName id="70A178FB3F1F384AFEE4F88FFD4BE627" box="[292,661,1841,1863]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="114" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov." subGenus="Omalodes">
<emphasis id="85D5DF6A3F1F384AFEE4F88FFE55E627" box="[292,395,1841,1863]" italics="true" pageId="5" pageNumber="114">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F1F384AFE5FF88FFDD8E627" box="[415,518,1841,1863]" italics="true" pageId="5" pageNumber="114">Omalodes</emphasis>
)
<emphasis id="85D5DF6A3F1F384AFDDBF88CFD4BE627" box="[539,661,1842,1863]" italics="true" pageId="5" pageNumber="114">atacamanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EE662113F1F384AFD61F88CFD32E627" box="[673,748,1842,1863]" pageId="5" pageNumber="114" rank="species">
<emphasis id="85D5DF6A3F1F384AFD61F88CFD32E627" bold="true" box="[673,748,1842,1863]" pageId="5" pageNumber="114">sp.nov.</emphasis>
</taxonomicNameLabel>
, female terminalia (AF) and male terminalia (G-N). A. Bursa copulatrix, common oviduct, spermatheca and spermathecal gland; B. Basal region of bursa copulatrix, spermatheca and spermathecal gland; C, Coxites in ventral view; D. Close of apex of the coxites in ventral view; E, Coxites in dorsal view; F, Coxites in lateral view; G. Aedeagus in dorsal view; H. Aedeagus in lateral view; I. Aedeagus in ventral view; J. 8th tergite; K. 8th sternite; L, Close of the apex of 8th sternite, setae on the apex; M. 9th and 10th tergites; N. 9th sternite. Scale: A, C, EK, M, N (
<quantity id="7059AE9D3F1F384AFF5EF86EFF30E686" box="[158,238,2000,2022]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="114" unit="mm" value="0.5">0.5 mm</quantity>
) and B, D, L (
<quantity id="7059AE9D3F1F384AFEBFF86EFE01E686" box="[383,479,2000,2022]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="114" unit="mm" value="0.25">0.25 mm</quantity>
).
</paragraph>
<caption id="E3DE53F03F19384CFF57F8A6FA9CE60B" httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="3" pageNumber="112" targetBox="[151,1436,193,1792]" targetPageId="3">
<paragraph id="B71E03783F19384CFF57F8A6FA9CE60B" blockId="3.[151,1436,1816,1899]" pageId="3" pageNumber="112">
<emphasis id="85D5DF6A3F19384CFF57F8A6FECBE64D" bold="true" box="[151,277,1816,1838]" pageId="3" pageNumber="112">FIGURE 2.</emphasis>
Male of
<taxonomicName id="70A178FB3F19384CFEBAF8A6FD3AE64E" box="[378,740,1816,1838]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="112" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov." subGenus="Omalodes">
<emphasis id="85D5DF6A3F19384CFEBAF8A6FE3FE64E" box="[378,481,1816,1838]" italics="true" pageId="3" pageNumber="112">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F19384CFE32F8A6FD87E64E" box="[498,601,1816,1838]" italics="true" pageId="3" pageNumber="112">Omalodes</emphasis>
)
<emphasis id="85D5DF6A3F19384CFDAAF8A7FD3AE64E" box="[618,740,1817,1838]" italics="true" pageId="3" pageNumber="112">atacamanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EE662113F19384CFD2DF8A6FCE6E64D" box="[749,824,1816,1837]" pageId="3" pageNumber="112" rank="species">
<emphasis id="85D5DF6A3F19384CFD2DF8A6FCE6E64D" bold="true" box="[749,824,1816,1837]" pageId="3" pageNumber="112">sp.nov.</emphasis>
</taxonomicNameLabel>
A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1mm.
</paragraph>
</caption>
<caption id="E3DE53F03F1E384BFF57F8BAFB25E659" httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="4" pageNumber="113" targetBox="[151,1436,194,1768]" targetPageId="4">
<paragraph id="B71E03783F1E384BFF57F8BAFB25E659" blockId="4.[151,1436,1796,1849]" pageId="4" pageNumber="113">
<emphasis id="85D5DF6A3F1E384BFF57F8BAFECCE679" bold="true" box="[151,274,1796,1818]" pageId="4" pageNumber="113">FIGURE 3.</emphasis>
<taxonomicName id="70A178FB3F1E384BFED8F8BAFDA3E67A" box="[280,637,1796,1818]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="113" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov." subGenus="Omalodes">
<emphasis id="85D5DF6A3F1E384BFED8F8BAFEA1E67A" box="[280,383,1796,1818]" italics="true" pageId="4" pageNumber="113">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F1E384BFE4DF8BAFE2AE67A" box="[397,500,1796,1818]" italics="true" pageId="4" pageNumber="113">Omalodes</emphasis>
)
<emphasis id="85D5DF6A3F1E384BFDC3F8BBFDA3E67A" box="[515,637,1797,1818]" italics="true" pageId="4" pageNumber="113">atacamanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EE662113F1E384BFD43F8BAFD0AE679" box="[643,724,1796,1817]" pageId="4" pageNumber="113" rank="species">
<emphasis id="85D5DF6A3F1E384BFD43F8BAFD0AE679" bold="true" box="[643,724,1796,1817]" pageId="4" pageNumber="113">sp. nov.</emphasis>
</taxonomicNameLabel>
A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1mm.
</paragraph>
</caption>
<caption id="E3DE53F03F1C3849FF57FDE0FF35E3F2" httpUri="https://zenodo.org/record/236797/files/figure.png" pageId="6" pageNumber="115" targetBox="[306,1286,204,575]" targetPageId="6">
<paragraph id="B71E03783F1C3849FF57FDE0FF35E3F2" blockId="6.[151,1437,606,658]" pageId="6" pageNumber="115">
<emphasis id="85D5DF6A3F1C3849FF57FDE0FECDE313" bold="true" box="[151,275,606,628]" pageId="6" pageNumber="115">FIGURE 5.</emphasis>
A. Pronotum of
<taxonomicName id="70A178FB3F1C3849FE7EFDE0FC5FE314" authority="Kirby, 1818" authorityName="Kirby" authorityYear="1818" box="[446,897,606,628]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="omega" subGenus="Omalodes">
<emphasis id="85D5DF6A3F1C3849FE7EFDE0FDFBE314" box="[446,549,606,628]" italics="true" pageId="6" pageNumber="115">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F1C3849FDF4FDE0FD44E314" box="[564,666,606,628]" italics="true" pageId="6" pageNumber="115">Omalodes</emphasis>
)
<emphasis id="85D5DF6A3F1C3849FD6AFDE1FD30E314" box="[682,750,607,628]" italics="true" pageId="6" pageNumber="115">omega</emphasis>
(Kirby, 1818)
</taxonomicName>
; B. Pronotum of
<taxonomicName id="70A178FB3F1C3849FBEFFDE0FA8AE314" authority="Lewis, 1885" authorityName="Lewis" authorityYear="1885" box="[1071,1364,606,628]" class="Insecta" family="Histeridae" genus="Ebonius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="85D5DF6A3F1C3849FBEFFDE0FB13E314" box="[1071,1229,606,628]" italics="true" pageId="6" pageNumber="115">Ebonius politus</emphasis>
Lewis, 1885
</taxonomicName>
. Scale: 0.5 mm.
</paragraph>
</caption>
<paragraph id="B71E03783F1C3849FF57FD04FC2FE4E7" blockId="6.[151,1437,698,2035]" pageId="6" pageNumber="115">
with deep cavity to insert the antennal club; prosternal lobe not longer than 1/3 of the total length of the prosternum, anterior margin rounded or feebly truncated, marginal prosternal stria absent, lateral extension of the prosternal lobe without projections; lateral marginal prosternal stria weak and marked only next to procoxae; prosternal keel with transverse elevation medially (
<figureCitation id="2F9A1FFD3F1C3849FD1CFC9BFCF5E25E" box="[732,811,805,830]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Figs. 2</figureCitation>
EF), anterior half with two parallel carinal prosternal striae, posterior margin acuminate (
<figureCitation id="2F9A1FFD3F1C3849FDEFFCF4FDAAE202" box="[559,628,842,867]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Fig. 2</figureCitation>
E), posterior gland openings located below the lateral marginal striae (
<figureCitation id="2F9A1FFD3F1C3849FF5FFCD3FF3DE2E6" box="[159,227,877,902]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Fig. 2</figureCitation>
E). Pronotum with lateral posterior extremity not rounded, posterior margin with obtuse angle (
<figureCitation id="2F9A1FFD3F1C3849FAEAFCD3FAB0E2E6" box="[1322,1390,877,902]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Fig. 3</figureCitation>
A); marginal pronotal stria usually complete; lateral pronotal stria complete. Elytra with posterior and sutural regions not depressed; posteriorly emarginate (
<figureCitation id="2F9A1FFD3F1C3849FD8BFC06FD47E2B1" box="[587,665,952,977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Figs. 2</figureCitation>
A, C) and with longitudinal strioles; 1st dorsal elytral stria complete, 2nd dorsal elytral stria variable on its length, 3rd dorsal stria short, not exceeding the anterior half of elytra, 4th and 5th dorsal striae variable on its length, sutural elytral stria absent (
<figureCitation id="2F9A1FFD3F1C3849FCB9FBBDFC1EE57C" box="[889,960,1027,1052]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Fig. 2</figureCitation>
A), inner subhumeral stria absent, outer subhumeral stria variable on its length, epipleural region with one stria not reaching the posterior margin. Mesoventrite, anterior margin strongly emarginate medially, laterally with shallow excavations for receiving the anterior trochanters; marginal mesoventral stria interrupted medially; discal mesoventral stria absent (
<figureCitation id="2F9A1FFD3F1C3849FAEDFBD1FAACE5E8" box="[1325,1394,1135,1160]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Fig. 2</figureCitation>
F). Metaventrite with lateral stria oblique, postmesocoxal stria absent (
<figureCitation id="2F9A1FFD3F1C3849FC46FB2DFC16E5CC" box="[902,968,1171,1196]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Fig. 2</figureCitation>
B). Profemora with setae on the posterior surface (
<figureCitation id="2F9A1FFD3F1C3849FF3CFB09FE90E5B0" box="[252,334,1207,1232]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Figs. 2</figureCitation>
EF); all tibiae with a row of setae on outer margin and submarginal region (
<figureCitation id="2F9A1FFD3F1C3849FB39FB09FA95E5B0" box="[1273,1355,1207,1232]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Figs. 3</figureCitation>
BD). Protibiae truncated with a weak emargination at apex; inner row of setae ending in an apical cluster (
<figureCitation id="2F9A1FFD3F1C3849FAEAFB65FAB1E594" box="[1322,1391,1243,1268]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Fig. 3</figureCitation>
B); tarsal cavity slightly sinuous; internal region of tarsal cavity with a differentiated sulcus with setae. Stria of 1st visible abdominal sternite marked only laterally and sometimes rudimentary. Propygidium and pygidium with punctuation similar to the dorsum, the first one with punctuation slightly thicker on the sides (
<figureCitation id="2F9A1FFD3F1C3849FB7CFAF4FB21E403" box="[1212,1279,1354,1379]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Fig. 3</figureCitation>
E). Pygidium longer than half propygidium length and with regular punctuation (
<figureCitation id="2F9A1FFD3F1C3849FC46FAD0FC16E4E7" box="[902,968,1390,1415]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Fig. 3</figureCitation>
E).
</paragraph>
<paragraph id="B71E03783F1C3849FF07FA2BFD00E7CF" blockId="6.[151,1437,698,2035]" pageId="6" pageNumber="115">
Male terminalia. 10th tergite cordiform and composed of two regions more sclerotized but sometimes with separation not clear; 9th tergite without lateral projections (Fig. 4 M); 9th sternite “U”-shaped (Fig. 4 N); 8th sternite composed of two sclerites, with setae on the lateral-posterior region (Figs. 4 KL); 8th tergite with basal membrane attachment line, basal margin emarginate (Fig. 4 J). Aedeagus, basal piece about 1/3 parameres length, with a dorso-lateral projection, anterior margin in dorsal view strongly emarginate on the middle (Figs. 4 G, H). Parameres with posterior margin medially emarginate; in lateral view subcylindrical on anterior region (
<figureCitation id="2F9A1FFD3F1C3849FB7EF9F0FADCE707" box="[1214,1282,1614,1639]" captionStart="FIGURE 3" captionStartId="4.[151,250,1796,1818]" captionTargetBox="[151,1436,194,1768]" captionTargetId="figure@4.[151,1436,193,1774]" captionTargetPageId="4" captionText="FIGURE 3. Omalodes (Omalodes) atacamanus sp. nov. A, Pronotum and apical region of elytra; B. Protibia; C, Mesotibia; D. Metatibia; E. Apex of elytra, propygidium and pygidium. (sto) with longitudinal strioles of elytra. Scale: 1 mm." httpUri="https://zenodo.org/record/236796/files/figure.png" pageId="6" pageNumber="115">Fig. 3</figureCitation>
H), posterior region slightly narrowed; ventrally fused on anterior region by a weakly sclerotized region, posterior region with setae, with the extremity rounded and oblique sides.
</paragraph>
<paragraph id="B71E03783F1C3849FF07F904FB45E603" blockId="6.[151,1437,698,2035]" pageId="6" pageNumber="115">Female terminalia. Spermatheca with constraining rings, longer than wide and inserted on the anterior extremity of bursa copulatrix (Figs. 4 AB). Spermathecal gland connected at the base of spermatheca and bigger than it, duct of spermathecal gland with constraining rings (Figs. 4 AB). Common oviduct inserted on the posterior region of the bursa (Fig. 4 A). Coxites without a subapical lateral projection next to the cavity for insertion of stylus; in lateral view with internal and external margins carinate (Figs. 4 CE).</paragraph>
</subSubSection>
<subSubSection id="FFBB50F33F1C3848FF07F8D0FC29E35C" lastPageId="7" lastPageNumber="116" pageId="6" pageNumber="115" type="discussion">
<paragraph id="B71E03783F1C3848FF07F8D0FE92E05F" blockId="6.[151,1437,698,2035]" lastBlockId="7.[151,1436,151,1006]" lastPageId="7" lastPageNumber="116" pageId="6" pageNumber="115">
<emphasis id="85D5DF6A3F1C3849FF07F8D0FEEAE6E7" bold="true" box="[199,308,1902,1927]" pageId="6" pageNumber="115">Remarks</emphasis>
. The frontal stria can be anteriorly discontinuous; supraorbital usually developed, but incomplete; frontoclypeal suture very discrete (imperceptible in some specimens). In the male, the suture between the prosternal keel and the lobe can have two glands opening at the middle (
<figureCitation id="2F9A1FFD3F1C3849FC04F808FBCFE6AF" box="[964,1041,1974,1999]" captionStart="FIGURE 2" captionStartId="3.[151,250,1816,1838]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@3.[151,1436,193,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male of Omalodes (Omalodes) atacamanus sp. nov. A. Habitus in dorsal view; B. Habitus in ventral view; C. Habitus in lateral view; D. Head and anterior margin of pronotum; E. Head, prosternum and mesoventrite in lateroventral view; F. Prosternum and mesoventrite. (bs) base of scape; (se) setae of profemur; (re) subrectangular elevation. Scale: 1 mm." httpUri="https://zenodo.org/record/236795/files/figure.png" pageId="6" pageNumber="115">Figs. 2</figureCitation>
EF), but it is not consistent in the species; the carinal stria of the prosternal keel may vary in length. Pronotal marginal stria may be slightly interrupted in the middle of the anterior margin of pronotum. Between the first and third dorsal striae there may be several short and random striae; second elytral dorsal stria can be complete or absent on the posterior half or posterior third; fourth and fifth dorsal striae can be short or absent; outer subhumeral stria is usually discontinuous next to apical extremity of humeral stria. The sulcus on internal region of tarsal cavity can be formed by strong and close punctures.
</paragraph>
<paragraph id="B71E03783F1D3848FF07FEF2FC29E35C" blockId="7.[151,1436,151,1006]" pageId="7" pageNumber="116">
<emphasis id="85D5DF6A3F1D3848FF07FEF2FE31E005" bold="true" box="[199,495,332,357]" pageId="7" pageNumber="116">Geographic distribution.</emphasis>
The new species is known only from the Antofagasta Province (Taltal, Paposo and Bahia Esmeralda) along the arid coastal region of
<collectingCountry id="CFB643E83F1D3848FD22FED1FCFFE0E8" box="[738,801,367,392]" name="Chile" pageId="7" pageNumber="116">Chile</collectingCountry>
, in the Atacama biogeographic province (described by OBrien 1971) which belongs to the South American Transition Zone (
<bibRefCitation id="D3307E893F1D3848FC33FE2AFB7EE0CD" author="Morrone" box="[1011,1184,404,429]" pageId="7" pageNumber="116" refString="Morrone, J. J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean Islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467 - 494. http: // dx. doi. org / 10.1146 / annurev. ento. 50.071803.130447" type="journal article" year="2006">Morrone 2006</bibRefCitation>
;
<bibRefCitation id="D3307E893F1D3848FB71FE2AFB30E0CC" author="Morrone" box="[1201,1262,404,428]" pageId="7" pageNumber="116" refString="Morrone, J. J. (2014) Biogeographic regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. http: // dx. doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">2014</bibRefCitation>
). This zone is characterized by the overlapping areas of Neotropical and Andean biotic components. The Atacama biogeographic province has an entomological fauna related with that from Coquimbo (belonging to Central Chilean Subregion of Andean Region) (
<bibRefCitation id="D3307E893F1D3848FE9FFDBEFDD6E378" author="Morrone" box="[351,520,511,536]" pageId="7" pageNumber="116" refString="Morrone, J. J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean Islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467 - 494. http: // dx. doi. org / 10.1146 / annurev. ento. 50.071803.130447" type="journal article" year="2006">Morrone 2006</bibRefCitation>
). Several taxa of Magnoliophyta (plants), Arthropoda (including
<taxonomicName id="70A178FB3F1D3848FB27FE41FABAE378" box="[1255,1380,511,536]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) and Vertebrata are reported as endemic to the Atacama province (
<bibRefCitation id="D3307E893F1D3848FC83FD9AFC35E35C" author="Morrone" box="[835,1003,548,572]" pageId="7" pageNumber="116" refString="Morrone, J. J. (2014) Biogeographic regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. http: // dx. doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FFBB50F33F1D3848FF07FDF9FF19E2AA" pageId="7" pageNumber="116" type="biology_ecology">
<paragraph id="B71E03783F1D3848FF07FDF9FF19E2AA" blockId="7.[151,1436,151,1006]" pageId="7" pageNumber="116">
<emphasis id="85D5DF6A3F1D3848FF07FDF9FEF9E300" bold="true" box="[199,295,583,608]" pageId="7" pageNumber="116">Biology.</emphasis>
The adults and larvae of
<taxonomicName id="70A178FB3F1D3848FDA7FDF9FCB3E300" box="[615,877,583,608]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov.">
<emphasis id="85D5DF6A3F1D3848FDA7FDF9FCB3E300" box="[615,877,583,608]" italics="true" pageId="7" pageNumber="116">Omalodes atacamanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EE662113F1D3848FCBAFDF9FC04E300" box="[890,986,583,608]" pageId="7" pageNumber="116" rank="species">
<emphasis id="85D5DF6A3F1D3848FCBAFDF9FC04E300" bold="true" box="[890,986,583,608]" pageId="7" pageNumber="116">sp. nov.</emphasis>
</taxonomicNameLabel>
were found within
<taxonomicName id="70A178FB3F1D3848FB0DFDF6FE62E3E4" authority="(Philippi) Britton &amp; Rose" authorityName="(Philippi) Britton &amp; Rose" class="Magnoliopsida" family="Cactaceae" genus="Copiapoa" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="116" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="85D5DF6A3F1D3848FB0DFDF6FA42E300" box="[1229,1436,584,608]" italics="true" pageId="7" pageNumber="116">Copiapoa cinerea</emphasis>
(Philippi) Britton &amp; Rose
</taxonomicName>
(Plantae:
<taxonomicName id="70A178FB3F1D3848FDF3FDD2FD78E3E4" box="[563,678,620,644]" class="Magnoliopsida" family="Cactaceae" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="116" phylum="Tracheophyta" rank="family">Cactaceae</taxonomicName>
) feeding on the larvae of
<taxonomicName id="70A178FB3F1D3848FC15FDD2FBADE3E5" box="[981,1139,620,645]" class="Insecta" family="Drosophilidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="family">Drosophilidae</taxonomicName>
and
<taxonomicName id="70A178FB3F1D3848FB70FDD2FAFFE3E5" box="[1200,1313,620,645]" class="Insecta" family="Syrphidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="family">Syrphidae</taxonomicName>
(
<taxonomicName id="70A178FB3F1D3848FAF4FDD2FA42E3E5" box="[1332,1436,620,645]" class="Insecta" family="Bombyliidae" genus="Volucella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F1D3848FAF4FDD2FA42E3E5" box="[1332,1436,620,645]" italics="true" pageId="7" pageNumber="116">Volucella</emphasis>
</taxonomicName>
sp.) (
<bibRefCitation id="D3307E893F1D3848FF14FD31FE53E3C8" author="Arriagada" box="[212,397,655,680]" pageId="7" pageNumber="116" refString="Arriagada, S. G. (1985) Notas sobre Histeridos Neotropicales (Coleoptera: Histeridae). Revista Chilena de Entomologia, 12, 105 - 112." type="journal article" year="1985">Arriagada 1985</bibRefCitation>
;
<bibRefCitation id="D3307E893F1D3848FE5DFD31FE06E3C8" author="Arriagada" box="[413,472,655,680]" pageId="7" pageNumber="116" refString="Arriagada, S. G. (1986) Histeridos Chilenos (Coleoptera: Histeridae). Primera Parte. Revista Chilena de Entomologia, 14, 71 - 80." type="journal article" year="1986">1986</bibRefCitation>
, cited it as
<taxonomicName id="70A178FB3F1D3848FDABFD31FB43E3C8" authority="Lewis" authorityName="Lewis" box="[619,1181,655,680]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="intermedius" subGenus="Diplogrammicus">
<emphasis id="85D5DF6A3F1D3848FDABFD31FD02E3C8" box="[619,732,655,680]" italics="true" pageId="7" pageNumber="116">Omalodes</emphasis>
(
<emphasis id="85D5DF6A3F1D3848FD32FD31FC70E3C8" box="[754,942,655,680]" italics="true" pageId="7" pageNumber="116">Diplogrammicus</emphasis>
)
<emphasis id="85D5DF6A3F1D3848FC03FD31FB99E3C8" box="[963,1095,655,680]" italics="true" pageId="7" pageNumber="116">intermedius</emphasis>
Lewis
</taxonomicName>
). All specimens were collected in September and October in the BWk region (cold desert) where the mean annual temperature is &lt;18°C (
<bibRefCitation id="D3307E893F1D3848FF5EFD67FEC9E392" author="Peel" box="[158,279,729,754]" pageId="7" pageNumber="116" refString="Peel, M. C., Finlayson, B. L. &amp; MacMahon, T. A. (2007) Update world map of Koppen-Geiger climate classification. Hydrology and Earth System Sciences, 11, 1633 - 1644. http: // dx. doi. org / 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">Peel 2007</bibRefCitation>
). Living in cold desert environments in South
<collectingCountry id="CFB643E83F1D3848FCF6FD67FC48E392" box="[822,918,729,754]" name="United States of America" pageId="7" pageNumber="116">America</collectingCountry>
appears to be a unique feature of
<taxonomicName id="70A178FB3F1D3848FAEAFD67FEC0E275" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="atacamanus" status="sp. nov.">
<emphasis id="85D5DF6A3F1D3848FAEAFD67FEC0E275" italics="true" pageId="7" pageNumber="116">Omalodes atacamanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EE662113F1D3848FEE9FD43FE58E276" box="[297,390,765,790]" pageId="7" pageNumber="116" rank="species">
<emphasis id="85D5DF6A3F1D3848FEE9FD43FE58E276" bold="true" box="[297,390,765,790]" pageId="7" pageNumber="116">sp. nov.</emphasis>
</taxonomicNameLabel>
among the
<taxonomicName id="70A178FB3F1D3848FDD4FD43FD51E276" box="[532,655,765,790]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="tribe" tribe="Omalodini">Omalodini</taxonomicName>
, since other species of
<taxonomicName id="70A178FB3F1D3848FC63FD43FBCBE276" box="[931,1045,765,790]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F1D3848FC63FD43FBCBE276" box="[931,1045,765,790]" italics="true" pageId="7" pageNumber="116">Omalodes</emphasis>
</taxonomicName>
and
<taxonomicName id="70A178FB3F1D3848FB92FD43FB70E276" box="[1106,1198,765,790]" class="Insecta" family="Histeridae" genus="Ebonius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F1D3848FB92FD43FB70E276" box="[1106,1198,765,790]" italics="true" pageId="7" pageNumber="116">Ebonius</emphasis>
</taxonomicName>
occupy tropical and subtropical areas, with only a few species of
<taxonomicName id="70A178FB3F1D3848FD5CFC9FFCD0E25A" box="[668,782,801,826]" class="Insecta" family="Histeridae" genus="Omalodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="85D5DF6A3F1D3848FD5CFC9FFCD0E25A" box="[668,782,801,826]" italics="true" pageId="7" pageNumber="116">Omalodes</emphasis>
</taxonomicName>
found in xeric areas in New
<collectingCountry id="CFB643E83F1D3848FB9FFC9FFB6AE25A" box="[1119,1204,801,826]" name="Mexico" pageId="7" pageNumber="116">Mexico</collectingCountry>
and Arizona (
<collectingCountry id="CFB643E83F1D3848FA99FC9CFA4AE25A" box="[1369,1428,802,826]" name="United States of America" pageId="7" pageNumber="116">USA</collectingCountry>
) (
<bibRefCitation id="D3307E893F1D3848FF5EFCFBFE14E23E" author="Kovarik" box="[158,458,837,862]" pageId="7" pageNumber="116" refString="Kovarik, P. W. &amp; Caterino, M. S. (2001) Histeridae. In: Arnett, R. H. Jr. &amp; Thomas, M. C. (Eds.) American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 214 - 227." type="book" year="2001">Kovarik &amp; Caterino 2001</bibRefCitation>
). The presence of setae on the outer submarginal region of anterior, middle and hind tibiae, as well as on the surface of the femur represent possible adaptations to sandy environments in order to increase the surface of contact of the beetle's body with the sand, and thus assist in the vertical movement in sandy soil.
</paragraph>
</subSubSection>
<subSubSection id="FFBB50F33F1D3848FF07FC6BFABAE28E" box="[199,1380,981,1006]" pageId="7" pageNumber="116" type="etymology">
<paragraph id="B71E03783F1D3848FF07FC6BFABAE28E" blockId="7.[151,1436,151,1006]" box="[199,1380,981,1006]" pageId="7" pageNumber="116">
<emphasis id="85D5DF6A3F1D3848FF07FC6BFE98E28E" bold="true" box="[199,326,981,1006]" pageId="7" pageNumber="116">Etymology</emphasis>
. The species name refers to the desert region of Atacama, where this species has been found.
</paragraph>
</subSubSection>
</treatment>
</document>