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<document id="196174902D8B07971F330F6838B70CF7" ID-DOI="10.11646/zootaxa.3920.1.2" ID-GBIF-Dataset="0ae5e027-d588-4b5a-9835-f0b7f9be4c14" ID-ISSN="1175-5326" ID-Zenodo-Dep="236803" ID-ZooBank="6A4779B4-DAE6-4168-9CD4-0F46B871FF66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994384254" checkinUser="plazi" docAuthor="Tan, Ming Kai, Liu, Chunxiang &amp; Artchawakom, Taksin" docDate="2015" docId="38500F2CB74CFFCBFF31FAF1A5C241F2" docLanguage="en" docName="zt03920p050.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tapiena" docType="key" docVersion="9" masterDocId="C4697754B74DFFC9FFA6FFF2A05E4400" masterDocTitle="Taxonomic review of Tapiena (Orthoptera: Phaneropteridae: Phaneropterinae), with key to species and new species from Thailand" updateTime="1698562410556" updateUser="plazi">
<mods:mods id="73C56E61348BFA54E3B1AEC1553BFB20" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A2D5E640112DAA03A1498125C7441572">Taxonomic review of Tapiena (Orthoptera: Phaneropteridae: Phaneropterinae), with key to species and new species from Thailand</mods:title>
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<mods:name id="6DD98CB2B8295DFA261DA8E1A57DFDB5" type="personal">
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<mods:namePart id="52E9DA08D8FC70D2EFB799AD1AD7FDB4">Tan, Ming Kai</mods:namePart>
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<mods:namePart id="E89D58D4C82D2132876887D835908595">Liu, Chunxiang</mods:namePart>
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<mods:name id="FA2A7C382B342906E47D8D25B42FD107" type="personal">
<mods:role id="D3A7A555413B8A26C12EA9668587AB3C">
<mods:roleTerm id="630B5E4A93D38F2EC08C53355B8731AE">Author</mods:roleTerm>
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<mods:namePart id="329693CA5CFE1DBB16C47921FFEB0B07">Artchawakom, Taksin</mods:namePart>
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<mods:typeOfResource id="A9C46B9C93C7E6CAC5ABCBD669BC9CF5">text</mods:typeOfResource>
<mods:relatedItem id="6820276B3EE7D42BA12DDE22FE48823B" type="host">
<mods:titleInfo id="C9D71690B8891B7E003865951E177EA2">
<mods:title id="F9488BAA33523B6345DEBCC1D7CB8E89">Zootaxa</mods:title>
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<mods:part id="6BE3C530DE3B4BD85DCC26543BFDD837">
<mods:date id="4C1ABA6179E876F0DE4336A25671353C">2015</mods:date>
<mods:detail id="1D3BC8280F7D235CC14037444236C269" type="volume">
<mods:number id="D9644BE6BE2556718D7A4EEC279B653D">3920</mods:number>
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<mods:detail id="E4FC543241D4179F575EB59333FE7DDB" type="issue">
<mods:number id="765CB202EAE62B8E394043A1C4E70CCA">1</mods:number>
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<mods:classification id="7B8ADED83BAA5155C14D38B210EF6C23">journal volume</mods:classification>
<mods:identifier id="9B3F5ECC460D48705B162D1792655BFA" type="DOI">10.11646/zootaxa.3920.1.2</mods:identifier>
<mods:identifier id="A98C7FA730D64A76E9A9BF4058BB8A0B" type="GBIF-Dataset">0ae5e027-d588-4b5a-9835-f0b7f9be4c14</mods:identifier>
<mods:identifier id="CBE39865013D4FBD9E94B49B2F1DED09" type="ISSN">1175-5326</mods:identifier>
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<treatment id="38500F2CB74CFFCBFF31FAF1A5C241F2" ID-DOI="http://doi.org/10.5281/zenodo.5671883" ID-GBIF-Taxon="119597171" ID-Zenodo-Dep="5671883" LSID="urn:lsid:plazi:treatment:38500F2CB74CFFCBFF31FAF1A5C241F2" httpUri="http://treatment.plazi.org/id/38500F2CB74CFFCBFF31FAF1A5C241F2" lastPageId="2" pageId="1">
<subSubSection id="F8E3EDB1B74CFFC8FF31FAF1A36341AA" pageId="1" type="nomenclature">
<paragraph id="B046BE3AB74CFFC8FF31FAF1A2C4411D" blockId="1.[151,666,1283,1309]" box="[151,666,1283,1309]" pageId="1">
<heading id="EB0E0956B74CFFC8FF31FAF1A2C4411D" bold="true" box="[151,666,1283,1309]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="828D6228B74CFFC8FF31FAF1A2C4411D" bold="true" box="[151,666,1283,1309]" pageId="1">
Key to species of
<taxonomicName id="77F9C5B9B74CFFC8FECBFAF1A190411D" box="[365,462,1283,1309]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="828D6228B74CFFC8FECBFAF1A190411D" bold="true" box="[365,462,1283,1309]" italics="true" pageId="1">Tapiena</emphasis>
</taxonomicName>
(for males only)
</emphasis>
</heading>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31FAB8A36341AA" blockId="1.[151,1437,1353,1450]" pageId="1">
<taxonomicName id="77F9C5B9B74CFFC8FF31FAB8A24C4162" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[151,530,1354,1378]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="828D6228B74CFFC8FF31FAB8A1274161" box="[151,377,1354,1377]" italics="true" pageId="1">Tapiena emarginata</emphasis>
<bibRefCitation id="D468C3CBB74CFFC8FE24FAB8A24C4162" author="Karny" box="[386,530,1354,1378]" pageId="1" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="77F9C5B9B74CFFC8FDB9FAB8A47F4162" authority="Ingrisch &amp; Shishodia, 2000" authorityName="Ingrisch &amp; Shishodia" authorityYear="2000" box="[543,1057,1353,1378]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="latifolia">
<emphasis id="828D6228B74CFFC8FDB9FAB8A2844162" box="[543,730,1353,1378]" italics="true" pageId="1">Tapiena latifolia</emphasis>
Ingrisch &amp; Shishodia, 2000
</taxonomicName>
,
<taxonomicName id="77F9C5B9B74CFFC8FB88FAB8A5C24162" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[1070,1436,1354,1378]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="pentagona">
<emphasis id="828D6228B74CFFC8FB88FAB8A5584161" box="[1070,1286,1354,1377]" italics="true" pageId="1">Tapiena pentagona</emphasis>
<bibRefCitation id="D468C3CBB74CFFC8FAB6FAB8A5C24162" author="Karny" box="[1296,1436,1354,1378]" pageId="1" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="77F9C5B9B74CFFC8FF6FFA9CA21E4186" authority="Karny, 1926" authorityName="Karny" authorityYear="1926" box="[201,576,1389,1414]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="triangulata">
<emphasis id="828D6228B74CFFC8FF6FFA9CA1F74186" box="[201,425,1389,1414]" italics="true" pageId="1">Tapiena triangulata</emphasis>
Karny, 1926
</taxonomicName>
are only known for females and thus not included in this key. This key is modified from key for
<collectingCountry id="C8EEFEAAB74CFFC8FE31FA63A18441AA" box="[407,474,1425,1450]" name="China" pageId="1">China</collectingCountry>
species by
<bibRefCitation id="D468C3CBB74CFFC8FDFAFA63A36741AA" author="Liu" box="[604,825,1425,1450]" pageId="1" refString="Liu, C. - X. &amp; Kang, L. (2010) The genus Tapiena (Orthoptera: Tettigoniidae: Phaneropterinae; Holochlorini), with description of three new species from China. Zootaxa, 2328, 49 - 64." type="journal article" year="2010">Liu &amp; Kang (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8E3EDB1B74CFFCBFF31FA2BA5C241F2" lastPageId="2" pageId="1" type="key">
<keyStep id="0B0DA69FB74CFFC8FF31FA2BA5C5420B" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31FA2BA5C241EE" blockId="1.[151,1436,1497,2017]" box="[151,1436,1497,1518]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31FA2BA5C241EE" box="[151,1436,1497,1518]" pageId="1">
1. Distribution: Africa...................................................................
<taxonomicName id="77F9C5B9B74CFFC8FB64FA2BA5C241EE" authority="Bolivar, 1906" authorityName="Bolivar" authorityYear="1906" box="[1218,1436,1497,1518]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="828D6228B74CFFC8FB64FA2BA54D41EE" box="[1218,1299,1497,1518]" italics="true" pageId="1">T. minor</emphasis>
Bolívar, 1906
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31FA04A5C5420B" blockId="1.[151,1436,1497,2017]" box="[151,1435,1526,1547]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31FA04A5C5420B" box="[151,1435,1526,1547]" pageId="1">- Distribution: Asia..................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74CFFC8FF31F9E6A5C54246" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31F9E6A5C84229" blockId="1.[151,1436,1497,2017]" box="[151,1430,1556,1577]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F9E6A5C84229" box="[151,1430,1556,1577]" pageId="1">
2. Distribution:
<collectingCountry id="C8EEFEAAB74CFFC8FEC7F9E6A1CC4229" box="[353,402,1556,1577]" name="India" pageId="1">India</collectingCountry>
...............
<taxonomicName id="77F9C5B9B74CFFC8FDEFF9E6A2934229" box="[585,717,1556,1577]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="acutangula">
<emphasis id="828D6228B74CFFC8FDEFF9E6A2934229" box="[585,717,1556,1577]" italics="true" pageId="1">T. acutangula</emphasis>
</taxonomicName>
(Brunner von Wattenwyl, 1878) or
<taxonomicName id="77F9C5B9B74CFFC8FB82F9E6A5C84229" authority="Ingrisch &amp; Shishodia, 2000" authorityName="Ingrisch &amp; Shishodia" authorityYear="2000" box="[1060,1430,1556,1577]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="latifolia">
<emphasis id="828D6228B74CFFC8FB82F9E6A4D64229" box="[1060,1160,1556,1577]" italics="true" pageId="1">T. latifolia</emphasis>
Ingrisch &amp; Shishodia, 2000
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31F9C3A5C54246" blockId="1.[151,1436,1497,2017]" box="[151,1435,1585,1606]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F9C3A5C54246" box="[151,1435,1585,1606]" pageId="1">
- Distribution:
<collectingCountry id="C8EEFEAAB74CFFC8FEC7F9C3A1C44246" box="[353,410,1585,1606]" name="China" pageId="1">China</collectingCountry>
and Southeast Asia.................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74CFFC8FF31F9BCA5C5429E" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31F9BCA5C24263" blockId="1.[151,1436,1497,2017]" box="[151,1436,1614,1635]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F9BCA5C24263" box="[151,1436,1614,1635]" pageId="1">
3. Tenth abdominal tergite with apical process highly modified into a large laterally compressed plate (Figs. 1A–
<date id="C44798FAB74CFFC8FAA2F9BCA5484263" box="[1284,1302,1614,1635]" pageId="1">1I</date>
, 5F)...... 4
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31F99EA5C5429E" blockId="1.[151,1436,1497,2017]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F99EA5C5429E" pageId="1">- Tenth abdominal tergite simple, with little or no modification, often forming a simple plate (not laterally compressed) (Figs. 1J–1L, 6E, 6F)....................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74CFFC8FF31F954A5C542F6" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31F954A5C542BB" blockId="1.[151,1436,1497,2017]" box="[151,1435,1702,1723]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F954A5C542BB" box="[151,1435,1702,1723]" pageId="1">
4. Apical process of tenth abdominal tergite bilobate apically (in dorsal view) (
<figureCitation id="28C2A2BFB74CFFC8FC1AF954A3A042BB" box="[956,1022,1702,1723]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="1">Figs. 2</figureCitation>
A–2E)............................ 5
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31F936A5C542F6" blockId="1.[151,1436,1497,2017]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F936A5C542F6" pageId="1">
- Apical process of tenth abdominal tergite not furrowed or bifurcate apically, often tapering into an acute, obtuse or truncated apex (in dorsal view) (
<figureCitation id="28C2A2BFB74CFFC8FE1FF913A1A542F6" box="[441,507,1761,1782]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="1">Figs. 2</figureCitation>
F–2H, 5E).................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74CFFC8FF31F90CA5C24331" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31F90CA5C54313" blockId="1.[151,1436,1497,2017]" box="[151,1435,1790,1811]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F90CA5C54313" box="[151,1435,1790,1811]" pageId="1">
5. Subgenital plate with stylus distinctively longer than apical lobe (
<figureCitation id="28C2A2BFB74CFFC8FCF3F90CA3C84313" box="[853,918,1790,1811]" captionStart="FIGURE 3" captionStartId="5.[151,250,1365,1387]" captionTargetBox="[158,1427,201,1333]" captionTargetId="figure@5.[152,1435,193,1344]" captionTargetPageId="5" captionText="FIGURE 3. Male subgenital plates in ventral views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena longzhouensis (C), Tapiena javanica (D), Tapiena bilobata (E), Tapiena stylata (F), Tapiena cerciata (G), and Tapiena simplicis (H). Line drawings redrawn from figures from Hebard (1922), Karny (1923), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236805/files/figure.png" pageId="1">Figs. 3</figureCitation>
A, 3B).................................... 6
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31F8EEA5C24331" blockId="1.[151,1436,1497,2017]" box="[151,1436,1820,1841]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F8EEA5C24331" box="[151,1436,1820,1841]" pageId="1">
- Subgenital plate with stylus about same length as, or only slightly longer than apical lobe (
<figureCitation id="28C2A2BFB74CFFC8FBE3F8EEA4D84331" box="[1093,1158,1820,1841]" captionStart="FIGURE 3" captionStartId="5.[151,250,1365,1387]" captionTargetBox="[158,1427,201,1333]" captionTargetId="figure@5.[152,1435,193,1344]" captionTargetPageId="5" captionText="FIGURE 3. Male subgenital plates in ventral views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena longzhouensis (C), Tapiena javanica (D), Tapiena bilobata (E), Tapiena stylata (F), Tapiena cerciata (G), and Tapiena simplicis (H). Line drawings redrawn from figures from Hebard (1922), Karny (1923), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236805/files/figure.png" pageId="1">Figs. 3</figureCitation>
C–3E)................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74CFFC8FF31F8CBA5C543E1" pageId="1">
<paragraph id="B046BE3AB74CFFC8FF31F8CBA5C24389" blockId="1.[151,1436,1497,2017]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F8CBA5C24389" pageId="1">
6. Stridulatory file on left tegmen with about 240 fine teeth. Apical process of tenth abdominal tergite with short lobes at apex diverging outwards from one another; excision between lobes broadly rounded (in dorsal view) (
<figureCitation id="28C2A2BFB74CFFC8FBD8F8A4A4E6436B" box="[1150,1208,1878,1899]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="1">Fig. 2</figureCitation>
A).............................................................................................
<taxonomicName id="77F9C5B9B74CFFC8FBC8F886A5C24389" authority="Liu &amp; Kang, 2010" authorityName="Liu &amp; Kang" authorityYear="2010" box="[1134,1436,1908,1929]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="stridulous">
<emphasis id="828D6228B74CFFC8FBC8F886A4B84389" box="[1134,1254,1908,1929]" italics="true" pageId="1">T. stridulous</emphasis>
<bibRefCitation id="D468C3CBB74CFFC8FB4AF886A5C24389" author="Liu" box="[1260,1436,1908,1929]" pageId="1" refString="Liu, C. - X. &amp; Kang, L. (2010) The genus Tapiena (Orthoptera: Tettigoniidae: Phaneropterinae; Holochlorini), with description of three new species from China. Zootaxa, 2328, 49 - 64." type="journal article" year="2010">Liu &amp; Kang, 2010</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="B046BE3AB74CFFC8FF31F863A5C543E1" blockId="1.[151,1436,1497,2017]" pageId="1">
<keyLead id="0B081D0FB74CFFC8FF31F863A5C543E1" pageId="1">
- Stridulatory file on left tegmen with about 150 rather distinct fine teeth and about apical 25 indistinct teeth. Apical process of tenth abdominal tergite with short lobes at apex not diverging outwards from one another; excision between lobes truncated (in dorsal view) (
<figureCitation id="28C2A2BFB74CFFC8FEC3F83EA1C143E1" box="[357,415,1996,2017]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="1">Fig. 2</figureCitation>
B)..........................................................
<taxonomicName id="77F9C5B9B74CFFC8FBCEF83EA5C543E1" authority="Liu &amp; Xia, 1996" authorityName="Liu &amp; Xia" authorityYear="1996" box="[1128,1435,1996,2017]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="1" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="828D6228B74CFFC8FBCEF83EA4A943E1" box="[1128,1271,1996,2017]" italics="true" pageId="1">T. hainanensis</emphasis>
Liu &amp; Xia, 1996
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FF65A5C24522" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FF65A5CB44CA" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FF65A5CB44CA" pageId="2">
7. Apical process of tenth abdominal tergite distinctively forked or bilobate apically (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFB16FF65A4AB44AC" box="[1200,1269,151,172]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">Figs. 2</figureCitation>
C, 2D); in lateral view, with ventral process pointing ventrad (Figs. 1C, 1D). Cercus more slender and gently curved inwards (
<figureCitation id="28C2A2BFB74FFFCBFB56FF47A57744CA" box="[1264,1321,181,202]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
A)..... 8
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FF20A5C24522" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FF20A5C24522" pageId="2">
- Apical process of tenth abdominal tergite feebly bilobate apically (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFBA0FF20A41E44E7" box="[1030,1088,210,231]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">Fig. 2</figureCitation>
E); in lateral view, with ventral process pointing ventrad-posteriorly (Fig. 1E). Cercus more robust with interior margin more concave (
<figureCitation id="28C2A2BFB74FFFCBFB06FF1DA4874504" box="[1184,1241,239,260]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
B).............................................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB3FFEFFA5C24522" authority="Liu, 2004" authorityName="Liu" authorityYear="2004" box="[1177,1436,269,290]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="longzhouensis">
<emphasis id="828D6228B74FFFCBFB3FFEFFA5674522" box="[1177,1337,269,290]" italics="true" pageId="2">T. longzhouensis</emphasis>
Liu, 2004
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FED8A5C245B4" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FED8A5C2457A" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FED8A5C2457A" pageId="2">
8. Apical process of tenth abdominal tergite tapering to an acute apex, with ventral process short and forked apically (in lateral view) (Fig. 1C); forked into two triangular lobes; excision between lobes subacute (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFB07FEB5A485455C" box="[1185,1243,327,348]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">Fig. 2</figureCitation>
C)................................................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB13FE97A5C2457A" authority="Karny, 1926" authorityName="Karny" authorityYear="1926" box="[1205,1436,357,378]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="828D6228B74FFFCBFB13FE97A57E457A" box="[1205,1312,357,378]" italics="true" pageId="2">T. javanica</emphasis>
Karny, 1926
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FE70A5C245B4" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FE70A5C245B4" pageId="2">
- Apical process of tenth abdominal tergite blunt, with ventral process long and acute apically (in lateral view) (Fig. 1D); forked into two truncated lobes; excision between lobes rounded (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFC14FE6DA3B245B4" box="[946,1004,415,436]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">Fig. 2</figureCitation>
D)..........
<taxonomicName id="77F9C5B9B74FFFCBFBD8FE6DA5C245B4" authority="Liu &amp; Kang, 2010" authorityName="Liu &amp; Kang" authorityYear="2010" box="[1150,1436,415,436]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="bilobata">
<emphasis id="828D6228B74FFFCBFBD8FE6DA4B845B4" box="[1150,1254,415,436]" italics="true" pageId="2">T. bilobata</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFB4AFE6DA5C245B4" author="Liu" box="[1260,1436,415,436]" pageId="2" refString="Liu, C. - X. &amp; Kang, L. (2010) The genus Tapiena (Orthoptera: Tettigoniidae: Phaneropterinae; Holochlorini), with description of three new species from China. Zootaxa, 2328, 49 - 64." type="journal article" year="2010">Liu &amp; Kang, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FE4FA5C54647" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FE4FA5C5460C" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FE4FA5C5460C" pageId="2">
9. Apical process of tenth abdominal tergite with dorsal surface spiniferous in the apical third, with numerous spines along ventral margin (Figs. 1F, 1G). Cercus with two processes at the apex, with a large distal spine and small subapical spine on exterior margin (
<figureCitation id="28C2A2BFB74FFFCBFE97FE05A12A460C" box="[305,372,503,524]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Figs. 4</figureCitation>
C, 4D)................................................................................. 10
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FDE7A5C54647" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FDE7A5C54647" pageId="2">- Apical process of tenth abdominal tergite with dorsal surface not spiniferous; with few or no spines along ventral margin. Cercus not as above...................................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FDBDA5C246BC" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FDBDA5C54682" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FDBDA5C54682" pageId="2">
10. Apical process of tenth abdominal tergite with 4–5 spines on ventral margin, apex more blunt (in lateral view) (Fig. 1F)................................................................................
<taxonomicName id="77F9C5B9B74FFFCBFBC2FD9FA5C54682" authority="Liu &amp; Xia, 1996" authorityName="Liu &amp; Xia" authorityYear="1996" box="[1124,1435,621,642]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="spinicaudata">
<emphasis id="828D6228B74FFFCBFBC2FD9FA4A94682" box="[1124,1271,621,642]" italics="true" pageId="2">T. spinicaudata</emphasis>
Liu &amp; Xia, 1996
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FD78A5C246BC" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FD78A5C246BC" pageId="2">
- Apical process of tenth abdominal tergite with 2–3 spines on ventral margin, apex more acute (in lateral view) (Fig. 1G)............................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB9AFD55A5C246BC" authority="Liu &amp; Kang, 2010" authorityName="Liu &amp; Kang" authorityYear="2010" box="[1084,1436,679,700]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="parapentagona">
<emphasis id="828D6228B74FFFCBFB9AFD55A4B846BC" box="[1084,1254,679,700]" italics="true" pageId="2">T. parapentagona</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFB4AFD55A5C246BC" author="Liu" box="[1260,1436,679,700]" pageId="2" refString="Liu, C. - X. &amp; Kang, L. (2010) The genus Tapiena (Orthoptera: Tettigoniidae: Phaneropterinae; Holochlorini), with description of three new species from China. Zootaxa, 2328, 49 - 64." type="journal article" year="2010">Liu &amp; Kang, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FD37A5C24715" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FD37A5C546DA" blockId="2.[151,1436,151,1522]" box="[151,1435,709,730]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FD37A5C546DA" box="[151,1435,709,730]" pageId="2">
11. Apical process of tenth abdominal tergite forming a triangular ventral process (in lateral view) (Figs. 1H,
<date id="C44798FAB74FFFCBFB73FD37A4B446DA" box="[1237,1258,709,730]" pageId="2">1I</date>
, 2G, 2H)...... 12
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FD10A5C24715" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FD10A5C24715" pageId="2">
- Apical process of tenth abdominal tergite produced into two long processes, not forming a triangular ventral process (in lateral view) (
<figureCitation id="28C2A2BFB74FFFCBFE83FD0DA1034714" box="[293,349,767,788]" captionStart="FIGURE 5" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[152,1435,361,1897]" captionTargetId="figure@7.[152,1435,361,1901]" captionTargetPageId="7" captionText="FIGURE 5. Tapiena sakaerat sp. n.: habitus of male holotype in lateral view (A); head and pronotum in lateral view (B); male tegmen in dorsal view (C); stridulatory file of left tegmen in ventral view (D); male abdominal apex in posterior (E) and lateral (F) views. Scale bars: 10 mm (A), 1 mm (B, C, E, F), 0.5 mm (D). d = dorsal apical process of tenth abdominal tergite; v = ventral apical process of tenth abdominal tergite." httpUri="https://zenodo.org/record/236807/files/figure.png" pageId="2">Fig. 5</figureCitation>
F)...........................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB53FD0DA53E4714" box="[1269,1376,767,788]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="sakaerat" status="sp. nov.">
<emphasis id="828D6228B74FFFCBFB53FD0DA53E4714" box="[1269,1376,767,788]" italics="true" pageId="2">T. sakaerat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BEDF53B74FFFCBFAC0FD0DA5C24715" box="[1382,1436,767,789]" pageId="2" rank="species">
<emphasis id="828D6228B74FFFCBFAC0FD0DA5C24715" bold="true" box="[1382,1436,767,789]" pageId="2">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FCEFA5C2478A" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FCEFA5C2474F" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FCEFA5C2474F" pageId="2">
12. Apical process of tenth abdominal tergite with apex acute (in lateral view and dorsally) (Figs. 1H, 2G)....................................................................................
<taxonomicName id="77F9C5B9B74FFFCBFC55FCC8A43A474F" box="[1011,1124,826,847]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="cucullata">
<emphasis id="828D6228B74FFFCBFC55FCC8A43A474F" box="[1011,1124,826,847]" italics="true" pageId="2">T. cucullata</emphasis>
</taxonomicName>
(Brunner von Wattenwyl, 1891)
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FCA5A5C2478A" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FCA5A5C2478A" pageId="2">
- Apical process of tenth abdominal tergite with apex truncated (in lateral view and dorsally) (Fig.
<date id="C44798FAB74FFFCBFB22FCA5A4C7476C" box="[1156,1177,855,876]" pageId="2">1I</date>
, 2H)...............................................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB2BFC87A5C2478A" authority="Xia &amp; Liu, 1990" authorityName="Xia &amp; Liu" authorityYear="1990" box="[1165,1436,885,906]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="828D6228B74FFFCBFB2BFC87A4A6478A" box="[1165,1272,885,906]" italics="true" pageId="2">T. yunnana</emphasis>
Xia &amp; Liu, 1990
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FC60A5C547C4" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FC60A5C547A7" blockId="2.[151,1436,151,1522]" box="[151,1435,914,935]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FC60A5C547A7" box="[151,1435,914,935]" pageId="2">
13. Tenth abdominal tergite with teeth or long process apically (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFC19FC60A45C47A7" box="[959,1026,914,935]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">
Figs.
<date id="C44798FAB74FFFCBFC53FC60A45C47A7" box="[1013,1026,914,935]" pageId="2">2</date>
</figureCitation>
I, 2J)............................ 14
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FC5DA5C547C4" blockId="2.[151,1436,151,1522]" box="[151,1435,943,964]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FC5DA5C547C4" box="[151,1435,943,964]" pageId="2">- Tenth abdominal tergite without tooth or long process apically (in dorsal view)................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FC3FA5C5403A" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FC3FA5C247FF" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FC3FA5C247FF" pageId="2">
14. Tenth abdominal tergite with a long process that tapers into an acute apex (Figs. 1J,
<date id="C44798FAB74FFFCBFBB5FC3FA47747E2" box="[1043,1065,973,994]" pageId="2">2I</date>
). Cercus simple with two small apical spines (Figs. 1J,
<date id="C44798FAB74FFFCBFE27FC18A1C847FF" box="[385,406,1002,1023]" pageId="2">2I</date>
)..................................................................
<taxonomicName id="77F9C5B9B74FFFCBFB17FC18A5C247FF" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[1201,1436,1002,1023]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="ensigera">
<emphasis id="828D6228B74FFFCBFB17FC18A57E47FF" box="[1201,1312,1002,1023]" italics="true" pageId="2">T. ensigera</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFA83FC18A5C247FF" author="Karny" box="[1317,1436,1002,1023]" pageId="2" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FBF5A5C5403A" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FBF5A5C5403A" pageId="2">
- Tenth abdominal tergite with four teeth apically, two lateral and two medial teeth (in dorsal view) (
<figureCitation id="28C2A2BFB74FFFCBFB38FBF5A486401C" box="[1182,1240,1031,1052]" captionStart="FIGURE 2" captionStartId="4.[151,250,1221,1243]" captionTargetBox="[162,1431,201,1191]" captionTargetId="figure@4.[151,1436,193,1200]" captionTargetPageId="4" captionText="FIGURE 2. Male tenth abdominal tergites in dorsal views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena javanica (C), Tapiena bilobata (D), Tapiena longzhouensis (E), Tapiena parapentagona (F), Tapiena cucullata (G), Tapiena yunnana (with cerci, H), Tapiena ensigera (with cerci, I), and Tapiena quadridens (J). Line drawings redrawn from figures from Karny (1923, 1926 a, 1926 b), Liu &amp; Kang (2010). c = cercus." httpUri="https://zenodo.org/record/236804/files/figure.png" pageId="2">Fig. 2</figureCitation>
J). Cercus with interior margin inflated from middle with apex forming a wide compressed tooth (
<figureCitation id="28C2A2BFB74FFFCBFC77FBD7A457403A" box="[977,1033,1061,1082]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
E)......
<taxonomicName id="77F9C5B9B74FFFCBFBD2FBD7A5C5403A" authority="Liu &amp; Xia, 1996" authorityName="Liu &amp; Xia" authorityYear="1996" box="[1140,1435,1061,1082]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="828D6228B74FFFCBFBD2FBD7A4A9403A" box="[1140,1271,1061,1082]" italics="true" pageId="2">T. quadridens</emphasis>
Liu &amp; Xia, 1996
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FBB0A5C54074" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FBB0A5C54057" blockId="2.[151,1436,151,1522]" box="[151,1435,1090,1111]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FBB0A5C54057" box="[151,1435,1090,1111]" pageId="2">
15. Cercus simple, bent inwards apically into an acute apex (
<figureCitation id="28C2A2BFB74FFFCBFCA5FBB0A3624057" box="[771,828,1090,1111]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
F)..........................
<taxonomicName id="77F9C5B9B74FFFCBFB29FBB0A5C54057" authority="Xia &amp; Liu, 1992" authorityName="Xia &amp; Liu" authorityYear="1992" box="[1167,1435,1090,1111]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="bivittata">
<emphasis id="828D6228B74FFFCBFB29FBB0A4A94057" box="[1167,1271,1090,1111]" italics="true" pageId="2">T. bivittata</emphasis>
Xia &amp; Liu, 1992
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FBADA5C54074" blockId="2.[151,1436,151,1522]" box="[151,1435,1119,1140]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FBADA5C54074" box="[151,1435,1119,1140]" pageId="2">
- Cercus modified (Figs. 1K, 1L, 4G–
<date id="C44798FAB74FFFCBFD98FBADA20D4074" box="[574,595,1119,1140]" pageId="2">4I</date>
, 6E–6G)............................................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FB8FA5C540AF" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FB8FA5C54092" blockId="2.[151,1436,151,1522]" box="[151,1435,1149,1170]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FB8FA5C54092" box="[151,1435,1149,1170]" pageId="2">16. Tenth abdominal tergite excised in the middle, to form two lobes............................................... 17</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FB68A5C540AF" blockId="2.[151,1436,151,1522]" box="[151,1435,1178,1199]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FB68A5C540AF" box="[151,1435,1178,1199]" pageId="2">- Tenth abdominal tergite not excised in the middle........................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FB45A5C240EA" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FB45A5C240CC" blockId="2.[151,1436,151,1522]" box="[151,1436,1207,1228]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FB45A5C240CC" box="[151,1436,1207,1228]" pageId="2">
17. Cercus with apex forming a broad, ovate node (Fig. 1K)......................................
<taxonomicName id="77F9C5B9B74FFFCBFB65FB45A5C240CC" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[1219,1436,1207,1228]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="bullata">
<emphasis id="828D6228B74FFFCBFB65FB45A54140CC" box="[1219,1311,1207,1228]" italics="true" pageId="2">T. bullata</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFA83FB45A5C240CC" author="Karny" box="[1317,1436,1207,1228]" pageId="2" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FB27A5C240EA" blockId="2.[151,1436,151,1522]" box="[151,1436,1237,1258]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FB27A5C240EA" box="[151,1436,1237,1258]" pageId="2">
- Cercus distinctively bilobate apically with obtuse inner tooth (Fig. 1L)...........................
<taxonomicName id="77F9C5B9B74FFFCBFB69FB27A5C240EA" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[1231,1436,1237,1258]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="828D6228B74FFFCBFB69FB27A54040EA" box="[1231,1310,1237,1258]" italics="true" pageId="2">T. incisa</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFA83FB27A5C240EA" author="Karny" box="[1317,1436,1237,1258]" pageId="2" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FB00A5C54142" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FB00A5C24125" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FB00A5C24125" pageId="2">
18. Cercus elongated; with apex acute and sclerotized, with dorsal margin denticulate (
<figureCitation id="28C2A2BFB74FFFCBFB86FB00A43A4107" box="[1056,1124,1266,1287]" captionStart="FIGURE 6" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[178,1408,185,1925]" captionTargetId="figure@9.[178,1409,185,1925]" captionTargetPageId="9" captionText="FIGURE 6. Tapiena denticulata sp. n.: habitus of male holotype in lateral view (A); head and pronotum in lateral view (B); male tegmen in dorsal view (C); stridulatory file of left tegmen in ventral view (D); male abdominal apex in dorso-posterior (E), lateral (F) and ventral (G) views. Scale bars: 10 mm (A), 1 mm (B, C, E – G), 0.5 mm (D)." httpUri="https://zenodo.org/record/236808/files/figure.png" pageId="2">Figs. 6</figureCitation>
F, 6G). Subgenital plate deeply excised between apical lobes (
<figureCitation id="28C2A2BFB74FFFCBFDA5FAFDA2624124" box="[515,572,1295,1316]" captionStart="FIGURE 6" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[178,1408,185,1925]" captionTargetId="figure@9.[178,1409,185,1925]" captionTargetPageId="9" captionText="FIGURE 6. Tapiena denticulata sp. n.: habitus of male holotype in lateral view (A); head and pronotum in lateral view (B); male tegmen in dorsal view (C); stridulatory file of left tegmen in ventral view (D); male abdominal apex in dorso-posterior (E), lateral (F) and ventral (G) views. Scale bars: 10 mm (A), 1 mm (B, C, E – G), 0.5 mm (D)." httpUri="https://zenodo.org/record/236808/files/figure.png" pageId="2">Fig. 6</figureCitation>
G)......................................................
<taxonomicName id="77F9C5B9B74FFFCBFB7DFAFDA53E4124" box="[1243,1376,1295,1316]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="828D6228B74FFFCBFB7DFAFDA53E4124" box="[1243,1376,1295,1316]" italics="true" pageId="2">T. denticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BEDF53B74FFFCBFAC0FAFDA5C24125" box="[1382,1436,1295,1317]" pageId="2" rank="species">
<emphasis id="828D6228B74FFFCBFAC0FAFDA5C24125" bold="true" box="[1382,1436,1295,1317]" pageId="2">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FADFA5C54142" blockId="2.[151,1436,151,1522]" box="[151,1435,1325,1346]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FADFA5C54142" box="[151,1435,1325,1346]" pageId="2">
- Cercus not as above. Subgenital plate not deeply excised between apical lobes (
<figureCitation id="28C2A2BFB74FFFCBFC7DFADFA4424142" box="[987,1052,1325,1346]" captionStart="FIGURE 3" captionStartId="5.[151,250,1365,1387]" captionTargetBox="[158,1427,201,1333]" captionTargetId="figure@5.[152,1435,193,1344]" captionTargetPageId="5" captionText="FIGURE 3. Male subgenital plates in ventral views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena longzhouensis (C), Tapiena javanica (D), Tapiena bilobata (E), Tapiena stylata (F), Tapiena cerciata (G), and Tapiena simplicis (H). Line drawings redrawn from figures from Hebard (1922), Karny (1923), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236805/files/figure.png" pageId="2">Figs. 3</figureCitation>
F–3H)........................ 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FAB8A5C5417C" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FAB8A5C5415F" blockId="2.[151,1436,151,1522]" box="[151,1435,1354,1375]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FAB8A5C5415F" box="[151,1435,1354,1375]" pageId="2">
19. Cercus more with more than apical half lamellate, narrowing into an apex that recurved dorsally (
<figureCitation id="28C2A2BFB74FFFCBFB21FAB8A494415F" box="[1159,1226,1354,1375]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Figs. 4</figureCitation>
G, 4H).......... 20
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FA95A5C5417C" blockId="2.[151,1436,151,1522]" box="[151,1435,1383,1404]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FA95A5C5417C" box="[151,1435,1383,1404]" pageId="2">
- Cercus apical part enlarged apically, with an upcurved hook (
<figureCitation id="28C2A2BFB74FFFCBFC8DFA95A33B417C" box="[811,869,1383,1404]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">
Fig.
<date id="C44798FAB74FFFCBFCFEFA95A33B417C" box="[856,869,1383,1404]" pageId="2">4</date>
</figureCitation>
I).......................
<taxonomicName id="77F9C5B9B74FFFCBFB2CFA95A5C5417C" authority="Liu &amp; Xia, 1996" authorityName="Liu &amp; Xia" authorityYear="1996" box="[1162,1435,1383,1404]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="simplicis">
<emphasis id="828D6228B74FFFCBFB2CFA95A4A8417C" box="[1162,1270,1383,1404]" italics="true" pageId="2">T. simplicis</emphasis>
Liu &amp; Xia, 1996
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0B0DA69FB74FFFCBFF31FA77A5C241F2" pageId="2">
<paragraph id="B046BE3AB74FFFCBFF31FA77A5C241B7" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FA77A5C241B7" pageId="2">
20. Cercus with a lamellate appendage pointing dorsally in the middle (
<figureCitation id="28C2A2BFB74FFFCBFCD2FA77A3F1419A" box="[884,943,1413,1434]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
G). Subgenital plate with posterior margin triangularly excised between the lobes (
<figureCitation id="28C2A2BFB74FFFCBFDBFFA50A20F41B7" box="[537,593,1442,1463]" captionStart="FIGURE 3" captionStartId="5.[151,250,1365,1387]" captionTargetBox="[158,1427,201,1333]" captionTargetId="figure@5.[152,1435,193,1344]" captionTargetPageId="5" captionText="FIGURE 3. Male subgenital plates in ventral views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena longzhouensis (C), Tapiena javanica (D), Tapiena bilobata (E), Tapiena stylata (F), Tapiena cerciata (G), and Tapiena simplicis (H). Line drawings redrawn from figures from Hebard (1922), Karny (1923), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236805/files/figure.png" pageId="2">Fig. 3</figureCitation>
F).............................................
<taxonomicName id="77F9C5B9B74FFFCBFB28FA50A5C241B7" authority="Bey-Bienko, 1935" authorityName="Bey-Bienko" authorityYear="1935" box="[1166,1436,1442,1463]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="stylata">
<emphasis id="828D6228B74FFFCBFB28FA50A4BB41B7" box="[1166,1253,1442,1463]" italics="true" pageId="2">T. stylata</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFB4AFA50A5C241B7" author="Bey-Bienko" box="[1260,1436,1442,1463]" pageId="2" refString="Bey-Bienko, G. (1935) Acrididae and Tettigoniidae from Luzon Philippine islands. Philippine Journal of Science, 57, 377 - 407" type="journal article" year="1935">Bey-Bienko, 1935</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B046BE3AB74FFFCBFF31FA4DA5C241F2" blockId="2.[151,1436,151,1522]" pageId="2">
<keyLead id="0B081D0FB74FFFCBFF31FA4DA5C241F2" pageId="2">
- Cercus without a lamellate appendage pointing dorsally in the middle (
<figureCitation id="28C2A2BFB74FFFCBFC15FA4DA3AF41D4" box="[947,1009,1471,1492]" captionStart="FIGURE 4" captionStartId="6.[151,250,1075,1097]" captionTargetBox="[154,1424,204,1042]" captionTargetId="figure@6.[151,1435,193,1054]" captionTargetPageId="6" captionText="FIGURE 4. Male cercus in dorsal views (all except G, H) of Tapiena bilobata (A), Tapiena longzhouensis (B), Tapiena spinicaudata (C), Tapiena parapentagona (D), Tapiena quadridens (E), Tapiena bivittata (F), Tapiena stylata (in lateral view, G), Tapiena cerciata (in lateral view, H), and Tapiena simplicis (I). Line drawings redrawn from figures from Karny (1923, 1926 b), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236806/files/figure.png" pageId="2">Fig. 4</figureCitation>
H). Subgenital plate with posterior margin roundly excised between the lobes (
<figureCitation id="28C2A2BFB74FFFCBFD9EFA2FA22E41F2" box="[568,624,1501,1522]" captionStart="FIGURE 3" captionStartId="5.[151,250,1365,1387]" captionTargetBox="[158,1427,201,1333]" captionTargetId="figure@5.[152,1435,193,1344]" captionTargetPageId="5" captionText="FIGURE 3. Male subgenital plates in ventral views of Tapiena stridulous (A), Tapiena hainanensis (B), Tapiena longzhouensis (C), Tapiena javanica (D), Tapiena bilobata (E), Tapiena stylata (F), Tapiena cerciata (G), and Tapiena simplicis (H). Line drawings redrawn from figures from Hebard (1922), Karny (1923), Bey-Bienko (1935), Liu &amp; Kang (2010)." httpUri="https://zenodo.org/record/236805/files/figure.png" pageId="2">Fig. 3</figureCitation>
G).............................................
<taxonomicName id="77F9C5B9B74FFFCBFB0BFA2FA5C241F2" authority="Hebard, 1922" authorityName="Hebard" authorityYear="1922" box="[1197,1436,1501,1522]" class="Insecta" family="Phaneropteridae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="2" phylum="Arthropoda" rank="species" species="cerciata">
<emphasis id="828D6228B74FFFCBFB0BFA2FA54D41F2" box="[1197,1299,1501,1522]" italics="true" pageId="2">T. cerciata</emphasis>
<bibRefCitation id="D468C3CBB74FFFCBFABFFA2FA5C241F2" author="Hebard" box="[1305,1436,1501,1522]" pageId="2" refString="Hebard, M. (1922) Studies in Malayan, Melanesian and Australian Tettigoniidae (Orthoptera). Proceedings of the Academy of Natural Sciences, Philadelphia, 74, 121 - 299, plates 11 - 22." type="journal article" year="1922">Hebard, 1922</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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