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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.60072" ID-GBIF-Dataset="a8375122-971c-4804-92a6-fe16e10e3224" ID-PMC="PMC8222295" ID-Pensoft-Pub="1313-2970-1044-427" ID-Pensoft-UUID="8320077CEFA5571D971EFEA66B0C6268" ID-PubMed="34183882" ID-ZooBank="9AA72654B6CC4081B557CD0BAE3939A3" ModsDocID="1313-2970-1044-427" checkinTime="1623875398087" checkinUser="pensoft" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2021" docId="382C4F214C4D5D4F801E642E5B814422" docLanguage="en" docName="ZooKeys 1044: 427-448" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.60072" docTitle="Diplocheila (Diplocheila) laevigata" docType="treatment" docVersion="4" id="8320077CEFA5571D971EFEA66B0C6268" lastPageNumber="427" masterDocId="8320077CEFA5571D971EFEA66B0C6268" masterDocTitle="The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="448" masterPageNumber="427" pageNumber="427" updateTime="1668150474377" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Allegro, Gianni</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5079-6671</mods:nameIdentifier>
<mods:affiliation>Strada Patro 11, I- 14036 Moncalvo (AT), Italy</mods:affiliation>
<mods:nameIdentifier type="email">gianni.allegro54@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giachino, Pier Mauro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1167-5447</mods:nameIdentifier>
<mods:affiliation>Via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1044</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>427</mods:start>
<mods:end>448</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1044.60072</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1044-427</mods:identifier>
<mods:identifier type="ZooBank">9AA72654B6CC4081B557CD0BAE3939A3</mods:identifier>
<mods:identifier type="Pensoft-UUID">8320077CEFA5571D971EFEA66B0C6268</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182754013" LSID="urn:lsid:plazi:treatment:382C4F214C4D5D4F801E642E5B814422" httpUri="http://treatment.plazi.org/id/382C4F214C4D5D4F801E642E5B814422" lastPageNumber="427" pageId="0" pageNumber="427">
<subSubSection pageId="0" pageNumber="427" type="nomenclature">
<paragraph pageId="0" pageNumber="427">
<taxonomicName LSID="382C4F21-4C4D-5D4F-801E-642E5B814422" authority="(Bates, 1892)" baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila) laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata" subGenus="Diplocheila">Diplocheila (Diplocheila) laevigata (Bates, 1892)</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diplocheila laevigata (Bates, 1892): habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure2" httpUri="https://binary.pensoft.net/fig/554093" pageId="0" pageNumber="427">Figures 2</figureCitation>
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">, 6</figureCitation>
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">, 8</figureCitation>
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">, 12</figureCitation>
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">, 14</figureCitation>
<figureCitation captionStart="Figures 18–19" captionStartId="F8" captionText="Figures 18 - 19. Male protarsi of Diplocheila species 18 D. laevigata from Cambodia 19 D. walterrossii sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures18-19" httpUri="https://binary.pensoft.net/fig/554099" pageId="0" pageNumber="427">, 18</figureCitation>
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">, 20</figureCitation>
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 26</figureCitation>
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="427" type="reference_group">
<paragraph pageId="0" pageNumber="427">
<taxonomicName class="Insecta" family="Carabidae" genus="Rembus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rembus politus" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="politus">Rembus politus</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.65151" author="MacLeay, WS" journalOrPublisher="Kingsbury, Parbury, and Allen, London" pageId="0" pageNumber="427" refId="B25" refString="MacLeay, WS, 1825. Annulosa Javanica, or an Attempt to Illustrate the Natural Affinities and Analogies of the Insects Collected in Java by Thomas Horsfield, M.D.F.L. &amp; G.S. and Deposited by Him in the Museum of the Honourable East-India Company. Number I. Kingsbury, Parbury, and Allen, London, DOI: https://doi.org/10.5962/bhl.title.65151" title="Annulosa Javanica, or an Attempt to Illustrate the Natural Affinities and Analogies of the Insects Collected in Java by Thomas Horsfield, M. D. F. L. &amp; G. S. and Deposited by Him in the Museum of the Honourable East-India Company. Number I." url="https://doi.org/10.5962/bhl.title.65151" year="1825">MacLeay 1825</bibRefCitation>
: 16
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName class="Insecta" family="Carabidae" genus="Rembus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rembus politus" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="politus">Rembus politus</taxonomicName>
MacLeay, 1825:
<bibRefCitation author="Redtenbacher, L" journalOrPublisher="Karl Gerold's Sohn, Wien" pageId="0" pageNumber="427" refId="B26" refString="Redtenbacher, L, 1867. Reise des oesterreichischen Fregatte Novara um die Erde in der Jahren 1857, 1858, 1859 unter der Befehlen des Commodore B. von Wuellerstorf-Urbair. Zoologischer Theil, Band 2. I. Coleoptera. Karl Gerold's Sohn, Wien" title="Reise des oesterreichischen Fregatte Novara um die Erde in der Jahren 1857, 1858, 1859 unter der Befehlen des Commodore B. von Wuellerstorf-Urbair. Zoologischer Theil, Band 2. I. Coleoptera." year="1867">Redtenbacher 1867</bibRefCitation>
: 10
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName class="Insecta" family="Carabidae" genus="Rhembus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhembus laevigatus" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigatus">Rhembus laevigatus</taxonomicName>
<bibRefCitation author="Bates, HW" journalOrPublisher="Annali del Museo civico di Storia naturale di Genova" pageId="0" pageNumber="427" pagination="265 - 248" refId="B10" refString="Bates, HW, 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo civico di Storia naturale di Genova 32: 265 - 248" title="Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae." volume="32" year="1892">Bates 1892</bibRefCitation>
: 326
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Eccoptogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eccoptogenius moestus" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="moestus">Eccoptogenius moestus</taxonomicName>
<bibRefCitation author="Bates, HW" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="427" pagination="261 - 286" refId="B9" refString="Bates, HW, 1889. Contributions a la faune Indo-Chinoise, 3e Memoire (1), Carabidae. Annales de la Societe Entomologique de France 9: 261 - 286" title="Contributions a la faune Indo-Chinoise, 3 e Memoire (1), Carabidae." volume="9" year="1889">Bates 1889</bibRefCitation>
: 267
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName authorityName="Chaudoir" authorityYear="1852" class="Insecta" family="Carabidae" genus="Eccoptogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eccoptogenius moestus" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="moestus">Eccoptogenius moestus</taxonomicName>
Bates, 1889:
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B1" refString="Andrewes, HE, 1921. III. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Royal Entomological Society of London: 145-190. https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" title="III. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Royal Entomological Society of London: 145 - 190." url="https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" year="1921">Andrewes 1921</bibRefCitation>
: 176
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName class="Insecta" family="Carabidae" genus="Diplochila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplochila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplochila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B1" refString="Andrewes, HE, 1921. III. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Royal Entomological Society of London: 145-190. https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" title="III. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Royal Entomological Society of London: 145 - 190." url="https://doi.org/10.1111/j.1365-2311.1921.tb02805.x" year="1921">Andrewes 1921</bibRefCitation>
: 177
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName class="Insecta" family="Carabidae" genus="Diplochila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplochila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplochila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation DOI="https://doi.org/10.1080/00222932208632677" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B2" refString="Andrewes, HE, 1922. Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281-295. https://doi.org/10.1080/00222932208632677" title="Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281 - 295." url="https://doi.org/10.1080/00222932208632677" year="1922">Andrewes 1922</bibRefCitation>
: 282
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Government of India Central Publication Branch, Calcutta" pageId="0" pageNumber="427" refId="B3" refString="Andrewes, HE, 1930. Catalogue of Indian insects. Part 18 - Carabidae. Government of India Central Publication Branch, Calcutta" title="Catalogue of Indian insects. Part 18 - Carabidae." year="1930">Andrewes 1930</bibRefCitation>
: 149
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Habu, A" journalOrPublisher="Bulletin of the National Institute of Agricultural Sciences (Series C)" pageId="0" pageNumber="427" pagination="49 - 73" refId="B15" refString="Habu, A, 1956. On the species of Diplocheila (Coleoptera, Carabidae) and its allied genera of Japan. Bulletin of the National Institute of Agricultural Sciences (Series C) 6: 49 - 73" title="On the species of Diplocheila (Coleoptera, Carabidae) and its allied genera of Japan." volume="6" year="1956">Habu 1956</bibRefCitation>
: 55
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
: 35
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Lorenz, W" journalOrPublisher="Published by the author, Tutzing" pageId="0" pageNumber="427" refId="B22" refString="Lorenz, W, 2005a. Systematic List of Extant Ground Beetles of the World (InsectaColeoptera Geadephaga: Trachypachidae and Carabidae Incl. Paussinae, Cicindelinae, Rhysodinae). Published by the author, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeoptera Geadephaga: Trachypachidae and Carabidae Incl. Paussinae, Cicindelinae, Rhysodinae)." year="2005 a">Lorenz 2005a</bibRefCitation>
: 342
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Lorenz, W" journalOrPublisher="Published by the author, Tutzing" pageId="0" pageNumber="427" refId="B23" refString="Lorenz, W, 2005b. Nomina Carabidarum. A Directory of the Scientific Names of Ground Beetles (InsectaColeoptera 'Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). (2nd Edn.). Published by the author, Tutzing" title="Nomina Carabidarum. A Directory of the Scientific Names of Ground Beetles (InsectaColeoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). (2 nd Edn.)." year="2005 b">Lorenz 2005b</bibRefCitation>
: 571
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev 2008</bibRefCitation>
: 682
</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">Diplocheila laevigata</taxonomicName>
(Bates, 1892):
<bibRefCitation author="Huber, C" editor="Loebl, I" journalOrPublisher="Archostemata - Myxophaga - Adephaga (Vol. 1). Revised and Updated Edition. Koninklijke Brill NV, Leiden" pageId="0" pageNumber="427" pagination="621 - 577" refId="B16" refString="Huber, C, Marggi, W, 2017. Licinini. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Archostemata - Myxophaga - Adephaga (Vol. 1). Revised and Updated Edition. Koninklijke Brill NV, Leiden: 621 - 577" title="Licinini." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Huber and Marggi 2017</bibRefCitation>
: 626
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="427" type="type locality">
<paragraph pageId="0" pageNumber="427">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="427">Kawkareet in Tenasserim (Myanmar).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="427" type="materials_examined">
<paragraph pageId="0" pageNumber="427">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="427">
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
: HT
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Fea" country="Myanmar" location="Gen. Febbr." municipality="Kawkareet in Tenasserim" specimenCount="♂">
<specimenCount>♂</specimenCount>
,
<collectingMunicipality>Kawkareet in Tenasserim</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:382C4F214C4D5D4F801E642E5B814422:4921B926D29FD7537265D5E742494C96" country="Myanmar" municipality="Kawkareet in Tenasserim" name="Gen. Febbr.">Gen. Febbr.</location>
1887,
<collectorName>Fea</collectorName>
legit (MCSNG) (figs 2, 6, 12, 26)
</materialsCitation>
.
<materialsCitation collectingDate="2019-05-17" collectorName="W. Rossi, V. Kong" country="Cambodia" location="Tonle Sap Lake" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Kampong Chhnang">
<collectingCountry name="Cambodia">Cambodia</collectingCountry>
:
<specimenCount type="male">3 ♂♂</specimenCount>
<specimenCount type="female">2 ♀♀</specimenCount>
<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
, banks of
<location LSID="urn:lsid:plazi:treatment:382C4F214C4D5D4F801E642E5B814422:26EEBA4CAEB0E0EA51671D266E5C9990" country="Cambodia" name="Tonle Sap Lake" stateProvince="Kampong Chhnang">Tonle Sap Lake</location>
,
<collectingDate value="2019-05-17">17.V.2019</collectingDate>
,
<collectorName>W. Rossi</collectorName>
and
<collectorName>V. Kong</collectorName>
leg. (CAl, CGi)
</materialsCitation>
;
<materialsCitation collectingDate="2019-05-18" collectorName="W. Rossi, V. Kong" country="Cambodia" county="Rolea" location="Toekchenh Village" municipality="District" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang">
<specimenCount type="male">1 ♂</specimenCount>
<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
,
<collectingCounty>Rolea</collectingCounty>
<normalizedToken originalValue="B’ier">B'ier</normalizedToken>
<collectingMunicipality>District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:382C4F214C4D5D4F801E642E5B814422:9E8F31BB94E319ED4A0D75CE08E3A78D" country="Cambodia" county="Rolea" municipality="District" name="Toekchenh Village" stateProvince="Kampong Chhnang">Toekchenh Village</location>
,
<collectingDate value="2019-05-18">18.V.2019</collectingDate>
,
<collectorName>W. Rossi</collectorName>
and
<collectorName>V. Kong</collectorName>
leg. (CAl)
</materialsCitation>
;
<materialsCitation collectingDate="2019-11-29" collectingDateMax="2019-12-03" collectingDateMin="2019-11-29" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" latitude="12.279722" location="Khsam" longLatPrecision="15" longitude="104.65794" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang">
<specimenCount type="male">1 ♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:382C4F214C4D5D4F801E642E5B814422:0EA301C0729F840D9C1BCEC949B95588" country="Cambodia" latitude="12.279722" longLatPrecision="15" longitude="104.65794" name="Khsam" stateProvince="Kampong Chhnang">Khsam</location>
,
<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
,
<geoCoordinate degrees="12" direction="north" minutes="16" orientation="latitude" precision="15" seconds="47" value="12.279722">12°16'47&quot;N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="39" orientation="longitude" precision="1" seconds="28.6" value="104.65794">104°39'28.6&quot;E</geoCoordinate>
,
<collectingDate value="2019-11-29" valueMax="2019-12-03" valueMin="2019-11-29">29.XI-3.XII.2019</collectingDate>
,
<collectingMethod>light trap</collectingMethod>
,
<collectorName>W. Rossi</collectorName>
and
<collectorName>V. Kong</collectorName>
leg. (CAl)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1044.60072.figure2" httpUri="https://binary.pensoft.net/fig/554093" pageId="0" pageNumber="427" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="427">
<emphasis bold="true" pageId="0" pageNumber="427">Figure 2.</emphasis>
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila laevigata</emphasis>
</taxonomicName>
(Bates, 1892): habitus of
<typeStatus>holotype</typeStatus>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="427" type="diagnosis">
<paragraph pageId="0" pageNumber="427">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila laevigata</emphasis>
</taxonomicName>
may be distinguished by the combination of the following characters. ABL = 14-16 mm; head with 1 supraorbital setiferous pore on each side; labrum with 6 setae (4 medial + 2 lateral), symmetrical and moderately emarginate (LR = 0.63-0.65) (Figs
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">12</figureCitation>
,
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">14</figureCitation>
); anterior margin of clypeus hardly concave (Fig.
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">12</figureCitation>
); pronotum transverse (PW/PL = 1.29-1.31), with sides delicately sinuate backwards (Fig.
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">6</figureCitation>
); elytral striae distinctly punctate; apical lamella of the median lobe of aedeagus in dorsal view shortly triangular, with blunt apex (Figs
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">20</figureCitation>
,
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="427" type="remarks">
<paragraph pageId="0" pageNumber="427">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="427">
<taxonomicName baseAuthorityName="Bates" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevigata" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila laevigata</emphasis>
</taxonomicName>
is recorded from southern China, Myanmar, Vietnam, Laos, and Cambodia (
<bibRefCitation DOI="https://doi.org/10.1080/00222932208632677" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B2" refString="Andrewes, HE, 1922. Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281-295. https://doi.org/10.1080/00222932208632677" title="Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281 - 295." url="https://doi.org/10.1080/00222932208632677" year="1922">Andrewes 1922</bibRefCitation>
;
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev 2008</bibRefCitation>
). Unfortunately, the male HT in the Fea Collection at MCSNG is an immature specimen (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diplocheila laevigata (Bates, 1892): habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure2" httpUri="https://binary.pensoft.net/fig/554093" pageId="0" pageNumber="427">2</figureCitation>
) with a scarcely chitinized aedeagus (Fig.
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">26</figureCitation>
); however, its examination, made possible thanks to the courtesy of the Honorary Curator Roberto Poggi, revealed that it was conspecific with the specimens from Cambodia (Figs
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">8</figureCitation>
,
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">14</figureCitation>
,
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">27</figureCitation>
). An aedeagus of
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
</taxonomicName>
, substantially corresponding to the HT, is also illustrated by
<bibRefCitation DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev (2008)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="427">
Two specimens from Thailand deposited in CGi (2 ♂♂, Chiang Mai, 6.V.1988, R. Sciaky det.; Figs
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">9</figureCitation>
,
<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">15</figureCitation>
) show a similar aedeagal morphology but pronotum and labrum are differently shaped compared to the HT and to the specimens from Cambodia; they could belong to a new species, but it is not described here, awaiting more abundant material. A further specimen from Thailand deposited at BMNH (1 ♀, Bangkok, Larnaudie leg., H.E. Andrewes Coll.) probably belongs to a different, new species.
</paragraph>
<paragraph pageId="0" pageNumber="427">
A male specimen from Indonesia deposited at BMNH (1 ♂, Indes Neerl., Boucard leg., H.E. Andrewes Coll.) differs from the
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
</taxonomicName>
HT not only in external morphology, but also in the shape of the aedeagus (although this is damaged, the apical lamella appears to be clearly different); its external morphology does not match
<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
</taxonomicName>
HT either, in spite of the opinion of
<bibRefCitation DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev (2008)</bibRefCitation>
that the records of
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
</taxonomicName>
from Indonesia, together with those from Japan and the Philippines, are probably instead
<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
</taxonomicName>
. Therefore, this specimen also likely belongs to a new species, awaiting description when more abundant material becomes available.
</paragraph>
</subSubSection>
</treatment>
</document>
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