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<document id="F4B4FB0DAD551048A747315896F42CCC" ID-DOI="10.11646/zootaxa.3750.5.9" ID-GBIF-Dataset="3afa7c4f-322c-4e3c-a48e-4b571785e695" ID-ISSN="1175-5326" ID-Zenodo-Dep="219169" ID-ZooBank="EC9A05CD-5A99-43B9-9EF5-49DD43C66B84" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460711199886" checkinUser="plazi" docAuthor="Huang, Jia, Li, Tong &amp; Chen, Hongwei" docDate="2013" docId="285487E7FF82FFB9FF1CFE5CA74AFA99" docLanguage="en" docName="zt03750p600.pdf" docOrigin="Zootaxa 3750 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucophenga quadrifurcata Huang, Li &amp; Chen, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="596" masterDocId="D46DFF9FFF8BFFB0FF8BFFD5A56BFFE3" masterDocTitle="The genus Leucophenga (Diptera, Drosophilidae), part III: the interrupta species group from the Oriental region, with morphological and molecular evidence" masterLastPageNumber="600" masterPageNumber="587" pageNumber="596" updateTime="1698358013533" updateUser="plazi">
<mods:mods id="5694EAB92D81E03F19712ABECE53BCF4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A650C342C799B95FCAB530D2C0BBC1C0">
<mods:title id="768FB987496D7234E5A0538D4AAF0CBD">The genus Leucophenga (Diptera, Drosophilidae), part III: the interrupta species group from the Oriental region, with morphological and molecular evidence</mods:title>
</mods:titleInfo>
<mods:name id="AD600E7CDBC60B9FE32D24AEE39A06A5" type="personal">
<mods:role id="46F95325E4894A5255472D5BB920B6C3">
<mods:roleTerm id="2E192214DA64080B46179FBB2FD274B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1943CDED0539E776642D3EFEE623F11B">Huang, Jia</mods:namePart>
</mods:name>
<mods:name id="6725ABBD111FBDBF1F529D987214D636" type="personal">
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<mods:roleTerm id="03AD61B524D9C1B5263130CFAD9BB5A3">Author</mods:roleTerm>
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<mods:namePart id="FDC2EA815815A8E813E43923DF132ED5">Li, Tong</mods:namePart>
</mods:name>
<mods:name id="3521A848B133208F745554E5C809EDB0" type="personal">
<mods:role id="AD5F2F806E5E36EEE1D1353BDCC6BAEE">
<mods:roleTerm id="1C21DAD11F4B897D08DE98212BBC609C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6B0E8594D33977D5ED2022FA27B3FB9">Chen, Hongwei</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D6D2149E8B04348B4EE1625E59477EC">text</mods:typeOfResource>
<mods:relatedItem id="36B758C6AC8185A057147458912F56AE" type="host">
<mods:titleInfo id="7FF1FB51BB84BC846E7C7C43032C4D93">
<mods:title id="989F1C117F65ED4651F282E142F84194">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="30E4E9C634DCC7E64683109336909F70">
<mods:date id="F1C33949A1793800F4B181A7174A439A">2013</mods:date>
<mods:detail id="4AF8DFD3264AD5B67F80F4D9045AEE3D" type="volume">
<mods:number id="9F4AEB7AC1E59771BC05CB58A436AE51">3750</mods:number>
</mods:detail>
<mods:detail id="5DA164909BA2BB6F81421E23014B68D5" type="issue">
<mods:number id="625626E76112FE5C697174D6319F638C">5</mods:number>
</mods:detail>
<mods:extent id="316E8E1915EFDA81A715BA2FE45A84ED" unit="page">
<mods:start id="37C9244EF622B94FC172E2E4A357CD20">587</mods:start>
<mods:end id="46763F35B09BB7312539FB98722B6625">600</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E89F7B23910FD804778C5B0C77438B91">journal article</mods:classification>
<mods:identifier id="09DD507A25D550A382B485C4870E747F" type="DOI">10.11646/zootaxa.3750.5.9</mods:identifier>
<mods:identifier id="9FD493E28E87D851651CE8D6BC5D5C96" type="GBIF-Dataset">3afa7c4f-322c-4e3c-a48e-4b571785e695</mods:identifier>
<mods:identifier id="AF678DF4E718CB046472F195C45AC341" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="12EA7B51327643CD6806303A27C0FA8C" type="Zenodo-Dep">219169</mods:identifier>
<mods:identifier id="DC21B256BEFD3CE0A6BE23E1A6512932" type="ZooBank">EC9A05CD-5A99-43B9-9EF5-49DD43C66B84</mods:identifier>
</mods:mods>
<treatment id="285487E7FF82FFB9FF1CFE5CA74AFA99" ID-DOI="http://doi.org/10.5281/zenodo.6156292" ID-GBIF-Taxon="119585103" ID-Zenodo-Dep="6156292" LSID="urn:lsid:plazi:treatment:285487E7FF82FFB9FF1CFE5CA74AFA99" httpUri="http://treatment.plazi.org/id/285487E7FF82FFB9FF1CFE5CA74AFA99" lastPageNumber="596" pageId="9" pageNumber="596">
<subSubSection id="E8E7657AFF82FFB9FF1CFE5CA4C1FE27" pageId="9" pageNumber="596" type="nomenclature">
<paragraph id="A04236F1FF82FFB9FF1CFE5CA724FE40" blockId="9.[151,591,393,453]" box="[151,591,393,419]" pageId="9" pageNumber="596">
<heading id="FB0A819DFF82FFB9FF1CFE5CA724FE40" bold="true" box="[151,591,393,419]" fontSize="11" level="1" pageId="9" pageNumber="596" reason="1">
<emphasis id="9289EAE3FF82FFB9FF1CFE5CA724FE40" bold="true" box="[151,591,393,419]" pageId="9" pageNumber="596">
<taxonomicName id="67FD4D72FF82FFB9FF1CFE5CA482FE40" box="[151,489,393,419]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="quadrifurcata" status="sp. nov.">
<emphasis id="9289EAE3FF82FFB9FF1CFE5CA482FE40" bold="true" box="[151,489,393,419]" italics="true" pageId="9" pageNumber="596">Leucophenga quadrifurcata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89BA5798FF82FFB9FE7AFE5CA724FE40" box="[497,591,393,419]" pageId="9" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A04236F1FF82FFB9FF1CFE79A4C1FE27" blockId="9.[151,591,393,453]" box="[151,426,428,453]" pageId="9" pageNumber="596">
(
<figureCitation id="38C62A74FF82FFB9FF14FE79A586FE27" box="[159,237,428,453]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Figs. 2</figureCitation>
J, 2K, 2L, 3H, 7)
</paragraph>
</subSubSection>
<subSubSection id="E8E7657AFF82FFB9FF1CFE3BA70EFDAE" pageId="9" pageNumber="596" type="diagnosis">
<paragraph id="A04236F1FF82FFB9FF1CFE3BA70EFDAE" blockId="9.[151,1437,494,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF1CFE3BA47AFDE4" bold="true" box="[151,273,494,519]" pageId="9" pageNumber="596">Diagnosis.</emphasis>
This species differs from the other species of the
<taxonomicName id="67FD4D72FF82FFB9FCC7FE3AA6D5FDE5" box="[844,958,495,518]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="9289EAE3FF82FFB9FCC7FE3AA6D5FDE5" box="[844,958,495,518]" italics="true" pageId="9" pageNumber="596">interrupta</emphasis>
</taxonomicName>
group in having the following characters: wing clouded on r-m and dm-cu crossveins (
<figureCitation id="38C62A74FF82FFB9FD02FDC4A7A6FDC9" box="[649,717,529,554]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Fig. 2</figureCitation>
J); aedeagus not bifurcated; aedeagal basolateral process slender basally, quadrifurcated distally (
<figureCitation id="38C62A74FF82FFB9FE75FDE1A729FDAE" box="[510,578,564,589]" captionStart="FIGURE 7" captionStartId="10.[151,250,1929,1951]" captionTargetBox="[151,1436,193,1907]" captionTargetId="figure@10.[151,1436,193,1908]" captionTargetPageId="10" captionText="FIGURE 7. Leucophenga quandrifurcata sp. nov., male terminalia in lateral view: A. epandrium, surstylus and cercus; B. hypandrium, paramere, gonopods, aedeagus and aedeagal apodeme; C. paramere; D. aedeagus. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/219176/files/figure.png" pageId="9" pageNumber="596">Fig. 7</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="E8E7657AFF82FFB9FF4CFD8DA75BFB44" pageId="9" pageNumber="596" type="description">
<paragraph id="A04236F1FF82FFB9FF4CFD8DA5A3FC6D" blockId="9.[151,1437,494,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF4CFD8DA432FD92" bold="true" box="[199,345,600,625]" pageId="9" pageNumber="596">Description.</emphasis>
Ocellar triangle yellowish brown to dark brown, with 3–4 setae above ocellar setae. Frons brownish yellow to yellowish brown. Pedicel yellowish brown; first flagellomere yellow. Clypeus brown. Scutum yellowish brown (
<figureCitation id="38C62A74FF82FFB9FEEDFD4BA4C7FD54" box="[358,428,670,695]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Fig. 2</figureCitation>
K). Postpronotal lobe brownish yellow (
<figureCitation id="38C62A74FF82FFB9FCF2FD4BA6D5FD54" box="[889,958,670,695]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Fig. 2</figureCitation>
L). Scutellum yellowish brown (
<figureCitation id="38C62A74FF82FFB9FAB8FD4BA013FD54" box="[1331,1400,670,695]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Fig. 2</figureCitation>
K). Wing (
<figureCitation id="38C62A74FF82FFB9FF6DFD14A446FD39" box="[230,301,705,730]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1876]" captionTargetBox="[151,1436,438,1831]" captionTargetId="figure@4.[151,1436,438,1833]" captionTargetPageId="4" captionText="FIGURE 2. Wing, mesonotum, scutellum and pleura of male: A, B, C. Leucophenga interrupta Duda, 1924; D, E, F. Leucophenga neointerrupta Fartyal &amp; Toda, 2005; G, H, I. Leucophenga bifurcata sp. nov.; J, K, L. Leucophenga quandrifurcata sp. nov.; M, N, O. Leucophenga retifoliacea sp. nov." httpUri="https://zenodo.org/record/219171/files/figure.png" pageId="9" pageNumber="596">Fig. 2</figureCitation>
J): Costal vein between R2+3 and R4+5 distally with 6–8 peg-like spinules on ventral surface. Halter mostly brownish. Abdominal tergites black (
<figureCitation id="38C62A74FF82FFB9FD2BFD3DA78CFCE2" box="[672,743,744,769]" captionStart="FIGURE 3" captionStartId="5.[151,250,1432,1454]" captionTargetBox="[157,1434,198,1406]" captionTargetId="figure@5.[151,1436,193,1410]" captionTargetPageId="5" captionText="FIGURE 3. Patterns of abdominal tergites of male and female: A, B. Leucophenga interrupta; C, D, E, F. L. neointerrupta; G. L. bifurcata sp. nov.; H. L. quandrifurcata sp. nov.; I, J. L. retifoliacea sp. nov.; K. Color legend." httpUri="https://zenodo.org/record/219172/files/figure.png" pageId="9" pageNumber="596">Fig. 3</figureCitation>
H); second mostly yellow, but brown posteriorly; third and fourth each with a pair of narrow, yellow patches submedially on anterior margin, yellow laterally; fifth to sixth mostly black. Male terminalia: Epandrium with ca. 10 setae near posterior margin and ventral corner per side (
<figureCitation id="38C62A74FF82FFB9FAE5FCFBA5CDFC89" captionStart="FIGURE 7" captionStartId="10.[151,250,1929,1951]" captionTargetBox="[151,1436,193,1907]" captionTargetId="figure@10.[151,1436,193,1908]" captionTargetPageId="10" captionText="FIGURE 7. Leucophenga quandrifurcata sp. nov., male terminalia in lateral view: A. epandrium, surstylus and cercus; B. hypandrium, paramere, gonopods, aedeagus and aedeagal apodeme; C. paramere; D. aedeagus. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/219176/files/figure.png" pageId="9" pageNumber="596">Fig. 7</figureCitation>
A). Paramere nearly rectangular distally, with several weak wrinkles, 2 sensilla and pubescence subdorsally (
<figureCitation id="38C62A74FF82FFB9FAE5FC87A5CDFC6D" captionStart="FIGURE 7" captionStartId="10.[151,250,1929,1951]" captionTargetBox="[151,1436,193,1907]" captionTargetId="figure@10.[151,1436,193,1908]" captionTargetPageId="10" captionText="FIGURE 7. Leucophenga quandrifurcata sp. nov., male terminalia in lateral view: A. epandrium, surstylus and cercus; B. hypandrium, paramere, gonopods, aedeagus and aedeagal apodeme; C. paramere; D. aedeagus. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/219176/files/figure.png" pageId="9" pageNumber="596">Fig. 7</figureCitation>
C).
</paragraph>
<paragraph id="A04236F1FF82FFB9FF4CFC4DA75BFB44" blockId="9.[151,1437,494,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF4CFC4DA416FC52" bold="true" box="[199,381,920,945]" pageId="9" pageNumber="596">Measurements.</emphasis>
BL =
<quantity id="67059B14FF82FFB9FE42FC4CA745FC53" box="[457,558,921,945]" metricMagnitude="-3" metricUnit="m" metricValue="2.93" pageId="9" pageNumber="596" unit="mm" value="2.93">2.93 mm</quantity>
in
<typeStatus id="7F468853FF82FFB9FDD8FC4DA7DEFC52" box="[595,693,920,945]" pageId="9" pageNumber="596" type="holotype">holotype</typeStatus>
(range in
<specimenCount id="B6FBFD78FF82FFB9FCA0FC4CA626FC52" box="[811,845,921,945]" pageId="9" pageNumber="596" type="generic">1♂</specimenCount>
and 2♀
<typeStatus id="7F468853FF82FFB9FC3AFC4CA14AFC52" box="[945,1057,921,945]" pageId="9" pageNumber="596" type="paratype">paratypes</typeStatus>
:
<quantity id="67059B14FF82FFB9FBA7FC4DA1FAFC53" box="[1068,1169,920,945]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="596" unit="mm" value="3.6">3.60 mm</quantity>
in ♂,
<quantity id="67059B14FF82FFB9FB50FC4DA014FC53" box="[1243,1407,920,945]" metricMagnitude="-3" metricUnit="m" metricValue="3.615" metricValueMax="3.73" metricValueMin="3.5" pageId="9" pageNumber="596" unit="mm" value="3.615" valueMax="3.73" valueMin="3.5">3.50–3.73 mm</quantity>
in ♀), THL =
<quantity id="67059B14FF82FFB9FE90FC6EA4EAFC30" box="[283,385,955,980]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="9" pageNumber="596" unit="mm" value="1.53">1.53 mm</quantity>
(
<quantity id="67059B14FF82FFB9FE1AFC69A49CFC30" box="[401,503,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="9" pageNumber="596" unit="mm" value="1.77">1.77 mm</quantity>
in ♂,
<quantity id="67059B14FF82FFB9FDC9FC69A78DFC30" box="[578,742,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="1.93" metricValueMin="1.7" pageId="9" pageNumber="596" unit="mm" value="1.815" valueMax="1.93" valueMin="1.7">1.70–1.93 mm</quantity>
in ♀), WL =
<quantity id="67059B14FF82FFB9FC0DFC6EA680FC30" box="[902,1003,955,980]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="596" unit="mm" value="2.6">2.60 mm</quantity>
(3.00 mm in ♂,
<quantity id="67059B14FF82FFB9FB27FC69A03AFC30" box="[1196,1361,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.27" metricValueMin="3.1" pageId="9" pageNumber="596" unit="mm" value="3.185" valueMax="3.27" valueMin="3.1">3.10–3.27 mm</quantity>
in ♀), WW =
<quantity id="67059B14FF82FFB9FF65FC0AA43EFC14" box="[238,341,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="596" unit="mm" value="1.2">1.20 mm</quantity>
(
<quantity id="67059B14FF82FFB9FEEDFC0AA4A5FC14" box="[358,462,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="596" unit="mm" value="1.33">1.33 mm</quantity>
in ♂,
<quantity id="67059B14FF82FFB9FD97FC0AA7AAFC14" box="[540,705,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.4" metricValueMin="1.24" pageId="9" pageNumber="596" unit="mm" value="1.3199999999999998" valueMax="1.4" valueMin="1.24">1.24–1.40 mm</quantity>
in ♀), arb = 5/3 (7–8/3–4), avd = 0.91 (0.81–0.90), adf = 1.94 (2.06–2.13), flw = 1.56 (2.25), FW/HW = 0.38 (0.34–0.37), ch/o = 0.03 (0.03–0.04), prorb = 0.56 (0.63–0.63), rcorb = 0.72 (0.65–0.75), vb = 0.29 (0.39–0.46), dcl = 0.39 (0.45–0.55), presctl = 0.61 (0.48–0.64), sctl = 1.29 (1.03–1.13), sterno = 0.75 (0.78–0.86), orbito = 1.89 (2.00), dcp = 0.21 (0.19–0.20), sctlp = 1.22 (1.11–1.13), C = 2.35 (2.69–2.71), 4c = 1.08 (0.93–0.95),
<date id="D4431031FF82FFB9FDE8FBB9A714FB67" box="[611,639,1132,1156]" pageId="9" pageNumber="596">4v</date>
= 1.81 (1.68–1.82),
<date id="D4431031FF82FFB9FCECFBBEA6E8FB67" box="[871,899,1131,1156]" pageId="9" pageNumber="596">5x</date>
= 0.75 (0.82–1.00), ac = 2.67 (2.10–2.18), M = 0.41 (0.36–0.39), C3F = 0.85 (0.88).
</paragraph>
</subSubSection>
<subSubSection id="E8E7657AFF82FFB9FF4CFB67A730FAF2" pageId="9" pageNumber="596" type="materials_examined">
<paragraph id="A04236F1FF82FFB9FF4CFB67A730FAF2" blockId="9.[151,1437,494,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF4CFB67A4E1FB28" bold="true" box="[199,394,1202,1227]" pageId="9" pageNumber="596">
<typeStatus id="7F468853FF82FFB9FF4CFB67A469FB28" box="[199,258,1202,1227]" pageId="9" pageNumber="596">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="7F468853FF82FFB9FE19FB67A491FB28" box="[402,506,1202,1227]" pageId="9" pageNumber="596" type="holotype">Holotype</typeStatus>
♂ (SCAU, No.
<date id="D4431031FF82FFB9FD38FB67A660FB29" box="[691,779,1202,1226]" pageId="9" pageNumber="596" value="83-12-18">121883</date>
),
<collectingCountry id="D8EA7661FF82FFB9FC94FB67A617FB29" box="[799,892,1202,1226]" name="China" pageId="9" pageNumber="596">CHINA</collectingCountry>
: Muyiji, Ximeng, Yunnan,
<quantity id="67059B14FF82FFB9FB3BFB67A06BFB29" box="[1200,1280,1202,1226]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="596" unit="m" value="1100.0">1100m</quantity>
,
<date id="D4431031FF82FFB9FA87FB67A01EFB28" box="[1292,1397,1202,1227]" pageId="9" pageNumber="596" value="2011-04-02">2.iv.2011</date>
, ex tussock, SJ Yan.
<typeStatus id="7F468853FF82FFB9FED8FB03A4AFFB0D" box="[339,452,1238,1262]" pageId="9" pageNumber="596" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="B6FBFD78FF82FFB9FE45FB03A498FB0D" box="[462,499,1238,1262]" pageId="9" pageNumber="596" type="generic">1♂</specimenCount>
, 2♀ (SCAU, Nos 123296–98),
<collectingCountry id="D8EA7661FF82FFB9FCD8FB03A6DBFB0D" box="[851,944,1238,1262]" name="China" pageId="9" pageNumber="596">CHINA</collectingCountry>
: Yixiang, Puer, Yunnan,
<geoCoordinate id="C5C95036FF82FFB9FB4FFB03A049FB0D" box="[1220,1314,1238,1262]" direction="north" orientation="latitude" pageId="9" pageNumber="596" precision="925" value="27.2">27°12'N</geoCoordinate>
,
<geoCoordinate id="C5C95036FF82FFB9FAA5FB03A0FCFB0D" box="[1326,1431,1238,1262]" direction="east" orientation="longitude" pageId="9" pageNumber="596" precision="925" value="100.05">100°03'E</geoCoordinate>
,
<quantity id="67059B14FF82FFB9FF1CFB2CA58CFAF2" box="[151,231,1273,1297]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="596" unit="m" value="1400.0">1400m</quantity>
,
<date id="D4431031FF82FFB9FF78FB2DA405FAF2" box="[243,366,1272,1297]" pageId="9" pageNumber="596" value="2013-03-29">29.iii.2013</date>
, ex tussocks, JJ Gao.
</paragraph>
</subSubSection>
<subSubSection id="E8E7657AFF82FFB9FF4CFACEA7B6FAB4" pageId="9" pageNumber="596" type="etymology">
<paragraph id="A04236F1FF82FFB9FF4CFACEA7B6FAB4" blockId="9.[151,1437,494,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF4CFACEA425FAD7" bold="true" box="[199,334,1307,1332]" pageId="9" pageNumber="596">Etymology.</emphasis>
A combination of the Latin word “
<emphasis id="9289EAE3FF82FFB9FD66FAC9A622FAD7" box="[749,841,1308,1332]" italics="true" pageId="9" pageNumber="596">quadrus</emphasis>
” (= four) + “foliaceus” (= folium), referring to the aedeagal basolateral process quadrifurcated distally.
</paragraph>
</subSubSection>
<subSubSection id="E8E7657AFF82FFB9FF4CFAB7A74AFA99" box="[199,545,1378,1403]" pageId="9" pageNumber="596" type="distribution">
<paragraph id="A04236F1FF82FFB9FF4CFAB7A74AFA99" blockId="9.[151,1437,494,1403]" box="[199,545,1378,1403]" pageId="9" pageNumber="596">
<emphasis id="9289EAE3FF82FFB9FF4CFAB7A40BFA98" bold="true" box="[199,352,1378,1403]" pageId="9" pageNumber="596">Distribution.</emphasis>
<collectingCountry id="D8EA7661FF82FFB9FEECFAB7A4C1FA98" box="[359,426,1378,1403]" name="China" pageId="9" pageNumber="596">China</collectingCountry>
(Yunnan).
</paragraph>
</subSubSection>
</treatment>
</document>
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