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<document id="7905BD65132A44ECBBD3EA772CF1BD80" ID="10.11646/zootaxa.4235.1.1SLASH10244" ID-CLB-Dataset="36483" ID-DOI="10.5281/zenodo.322062" ID-GBIF-Dataset="29139646-2af5-4d4f-a7a4-b03c54cb4af8" ID-ISSN="1175-5326" ID-Zenodo-Dep="322062" ID-ZooBank="AEA565CC-1A14-4987-977A-1C269B0FE2A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487671437888" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2017" docId="A15040439A24FFD0FF39FBDCFBE9FA1B" docLanguage="en" docName="zootaxa.4235.1.1SLASH10244.pdf" docOrigin="Zootaxa 4235 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia rudolphi Garrison &amp; von Ellenrieder, n. sp." docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="5D69383B9A32FFC8FFAEFF9EFF8AFFA4" masterDocTitle="New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="23" updateTime="1698437136683" updateUser="ExternalLinkService">
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<mods:titleInfo id="15287FE0B9BD0A0439E2EEFA4CF9339F">
<mods:title id="097C8731E4128F25900E8BDCA416F350">New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="D5EBB7FDFD5EBA62A4839074914C8905" type="personal">
<mods:role id="2024EB7F8ABC552CB9F25DCAA087C452">
<mods:roleTerm id="4321427CA10050FB858119C648218607">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A7060424E2C94E00008DBCCC44EFED3">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:name id="11B6AFD68E7B72ECACF496B3CF7C2EA8" type="personal">
<mods:role id="71600E7C8C5680A05CED81BA6914B7A5">
<mods:roleTerm id="CDA2CDC2D09E16AD627D8D7DE62EF475">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91573EB6772B2DAEE75F64F39A288772">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A53DEF37162A69F913D95F87F3B44E7">text</mods:typeOfResource>
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<mods:title id="E54402D8340E209E750BFB55B04E8B59">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="917028962CB5CEAD23A89391D68CADBD">
<mods:date id="37A8734AEAF0DA71C539579FFD52AB81">2017</mods:date>
<mods:detail id="EB568FB2B4D2F88B05A39FFE63D96277" type="volume">
<mods:number id="0E2A4CBEBDA851C0BFAA9F88AFE56245">4235</mods:number>
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<mods:classification id="324A2754FCAF6E8014B0C28ECD229B37">journal article</mods:classification>
<mods:identifier id="114023FFEE2C8546AF09286E9493405B" type="CLB-Dataset">36483</mods:identifier>
<mods:identifier id="3A38F6E2A6A081C8AA7757F646271FC8" type="DOI">10.5281/zenodo.322062</mods:identifier>
<mods:identifier id="6FF21079B90FB7E2F01FAF3EFE5A8222" type="GBIF-Dataset">29139646-2af5-4d4f-a7a4-b03c54cb4af8</mods:identifier>
<mods:identifier id="2D73420F3256403F6C61449F9019DCEC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="70454D02A6D819BE74C9274FA90256B0" type="Zenodo-Dep">322062</mods:identifier>
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<treatment id="A15040439A24FFD0FF39FBDCFBE9FA1B" ID-DOI="http://doi.org/10.5281/zenodo.6030015" ID-GBIF-Taxon="127697287" ID-Zenodo-Dep="6030015" LSID="urn:lsid:plazi:treatment:A15040439A24FFD0FF39FBDCFBE9FA1B" httpUri="http://treatment.plazi.org/id/A15040439A24FFD0FF39FBDCFBE9FA1B" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="61E3A2DE9A24FFDEFF39FBDCFCC6FB06" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="2946F1559A24FFDEFF39FBDCFD76FBF9" blockId="22.[151,1437,1090,1186]" box="[151,764,1090,1117]" pageId="22" pageNumber="23">
<heading id="720E46399A24FFDEFF39FBDCFD76FBF9" bold="true" box="[151,764,1090,1117]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="1B8D2D479A24FFDEFF39FBDCFD76FBF9" bold="true" box="[151,764,1090,1117]" pageId="22" pageNumber="23">
<taxonomicName id="EEF98AD69A24FFDEFF39FBDCFD27FBF9" ID-CoL="9X233" authority="Garrison &amp; von Ellenrieder" authorityName="Garrison &amp; von Ellenrieder" box="[151,685,1090,1117]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rudolphi" status="sp. nov.">
<emphasis id="1B8D2D479A24FFDEFF39FBDCFEC1FBF8" bold="true" box="[151,331,1090,1116]" italics="true" pageId="22" pageNumber="23">Argia rudolphi</emphasis>
Garrison &amp; von Ellenrieder
</taxonomicName>
,
<taxonomicNameLabel id="00BE903C9A24FFDEFD17FBDDFD76FBF9" box="[697,764,1091,1117]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2946F1559A24FFDEFF39FBFBFCC6FB06" blockId="22.[151,1437,1090,1186]" pageId="22" pageNumber="23">
<figureCitation id="B1C2EDD09A24FFDEFF39FBFBFF7BFBDA" box="[151,241,1125,1150]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="22" pageNumber="23">Figs. 19</figureCitation>
(head, thorax, S1
<specimenCount id="3FFF3ADC9A24FFDEFE6AFBF8FE78FBDA" box="[452,498,1126,1150]" pageId="22" pageNumber="23" type="male">4 ♂</specimenCount>
); 43 (head, thorax, S1
<specimenCount id="3FFF3ADC9A24FFDEFD56FBF8FCA9FBDA" box="[760,803,1126,1150]" pageId="22" pageNumber="23" type="female">4 ♀</specimenCount>
); 68 (S7
<specimenCount id="3FFF3ADC9A24FFDEFC3AFBF8FC44FBDA" box="[916,974,1126,1150]" pageId="22" pageNumber="23" type="male">10 ♂</specimenCount>
); 92 (S7
<specimenCount id="3FFF3ADC9A24FFDEFB92FBF8FBF9FBDA" box="[1084,1139,1126,1150]" pageId="22" pageNumber="23" type="female">10 ♀</specimenCount>
); 113 (mesostigmal plates
<specimenCount id="3FFF3ADC9A24FFDEFF39FB14FF21FB06" box="[151,171,1162,1186]" pageId="22" pageNumber="23" type="female"></specimenCount>
); 154 (appendages
<specimenCount id="3FFF3ADC9A24FFDEFE26FB14FE15FB06" box="[392,415,1162,1186]" pageId="22" pageNumber="23" type="male"></specimenCount>
); 168 (map);
<tableCitation id="647BC4EE9A24FFDEFD96FB17FD01FB06" box="[568,651,1161,1186]" captionStart="TABLE 2" captionStartId="10.[119,200,559,580]" captionTargetBox="[119,1846,707,1059]" captionTargetPageId="10" captionText="TABLE 2. Wing counts and measurements (average and standard deviation followed by range in square brackets) for metallic Argia species treated in this study." httpUri="http://table.plazi.org/id/7D86A1DD9A38FFC2FFD9FDB1F909FDE0" pageId="22" pageNumber="23" tableUuid="7D86A1DD9A38FFC2FFD9FDB1F909FDE0">Table 2</tableCitation>
(measurements).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A24FFDEFF39FB4FFDF2F946" pageId="22" pageNumber="23" type="etymology">
<paragraph id="2946F1559A24FFDEFF39FB4FFDC8FA96" blockId="22.[151,1437,1233,2014]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF39FB4FFE9CFB4E" bold="true" box="[151,278,1233,1258]" pageId="22" pageNumber="23">Etymology</emphasis>
. Named
<taxonomicName id="EEF98AD69A24FFDEFE2EFB4FFE6EFB4E" box="[384,484,1233,1258]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A24FFDEFE2EFB4FFE6EFB4E" bold="true" box="[384,484,1233,1258]" italics="true" pageId="22" pageNumber="23">rudolphi</emphasis>
</taxonomicName>
(Latinized name) in honor of our colleague Rainer Rudolph, in recognition for his generous support of the International Dragonfly Fund, which has helped further the knowledge of
<taxonomicName id="EEF98AD69A24FFDEFA93FB6BFA16FAAA" box="[1341,1436,1269,1294]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
worldwide over the past twenty years.
</paragraph>
<paragraph id="2946F1559A24FFDEFF69FAA3FF7BF9D2" blockId="22.[151,1437,1233,2014]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FAA3FE4CFAF2" bold="true" box="[199,454,1341,1366]" pageId="22" pageNumber="23">Specimens examined</emphasis>
. 18 ♂, 5 ♀.
<emphasis id="1B8D2D479A24FFDEFDDAFAA3FD33FAF2" bold="true" box="[628,697,1341,1366]" pageId="22" pageNumber="23">Types</emphasis>
.
<materialsCitation id="9991FB089A24FFDEFD60FAA3FC6EFAF2" ID-GBIF-Occurrence="1434659027" box="[718,996,1341,1366]" country="Mexico" location="MEXICO" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="1B8D2D479A24FFDEFD60FAA3FCB1FAF2" bold="true" box="[718,827,1341,1366]" pageId="22" pageNumber="23">
<typeStatus id="F6424FF79A24FFDEFD60FAA3FCB1FAF2" box="[718,827,1341,1366]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="3FFF3ADC9A24FFDEFCE7FAA0FCEBFAF1" box="[841,865,1342,1365]" pageId="22" pageNumber="23" type="male"></specimenCount>
:
<collectingCountry id="51EEB1C59A24FFDEFCDDFAA0FC6EFAF2" box="[883,996,1342,1366]" name="Mexico" pageId="22" pageNumber="23">MEXICO</collectingCountry>
</materialsCitation>
,
<materialsCitation id="9991FB089A24FFDEFC58FAA3FEB7FADE" ID-GBIF-Occurrence="1434659025" collectorName="Sierra de Huauchinango" country="Mexico" location="Puebla" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Puebla" typeStatus="holotype">
<collectingRegion id="EB3D3FB79A24FFDEFC58FAA3FBC8FAF2" box="[1014,1090,1341,1366]" country="Mexico" name="Puebla" pageId="22" pageNumber="23">Puebla</collectingRegion>
State, Zihuateutla,
<collectorName id="840C94839A24FFDEFA9FFAA3FEB7FADE" pageId="22" pageNumber="23">Sierra de Huauchinango</collectorName>
</materialsCitation>
,
<materialsCitation id="9991FB089A24FFDEFEE6FAFCFEACFA3A" ID-GBIF-Occurrence="1434659017" collectingDate="1987-05-21" collectionCode="CSCA" collectorName="R. Novelo &amp; A. Gomez" country="Mexico" elevation="596" latitude="20.240278" location="La Union" longLatPrecision="21" longitude="-97.89389" pageId="22" pageNumber="23" specimenCount="1" stateProvince="La Union" typeStatus="holotype">
<collectingRegion id="EB3D3FB79A24FFDEFEE6FAFCFE3CFADE" box="[328,438,1377,1402]" country="Philippines" name="La Union" pageId="22" pageNumber="23">La Unión</collectingRegion>
, in drainage area (
<geoCoordinate id="4CCD97929A24FFDEFD2DFAFFFC9AFADD" box="[643,784,1377,1402]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="20.240278">20°14'25'' N</geoCoordinate>
,
<geoCoordinate id="4CCD97929A24FFDEFCB5FAFFFC26FADD" box="[795,940,1377,1402]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-97.89389">97°53'38'' W</geoCoordinate>
,
<quantity id="EE015CB09A24FFDEFC18FAFFFB8AFADE" box="[950,1024,1377,1402]" metricMagnitude="2" metricUnit="m" metricValue="5.96" pageId="22" pageNumber="23" unit="m" value="596.0">
<elevation id="A2D416669A24FFDEFC18FAFFFB8AFADE" box="[950,1024,1377,1402]" metricMagnitude="2" metricUnit="m" metricValue="5.96" pageId="22" pageNumber="23" unit="m" value="596.0">596 m</elevation>
</quantity>
),
<date id="5D47D7959A24FFDEFBBAFAFCFB0DFADE" box="[1044,1159,1378,1402]" pageId="22" pageNumber="23" value="1987-05-21">
<collectingDate id="4D032E7D9A24FFDEFBBAFAFCFB0DFADE" box="[1044,1159,1378,1402]" pageId="22" pageNumber="23" value="1987-05-21">21 v 1987</collectingDate>
</date>
,
<collectorName id="840C94839A24FFDEFB3CFAFCFA8CFADE" box="[1170,1286,1377,1402]" pageId="22" pageNumber="23">R. Novelo</collectorName>
&amp;
<collectorName id="840C94839A24FFDEFA87FAFCFA16FADE" box="[1321,1436,1378,1402]" pageId="22" pageNumber="23">A. Gomez</collectorName>
leg. [
<collectionCode id="4FE869909A24FFDEFF60FA18FEAAFA3A" box="[206,288,1414,1438]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23">CSCA</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="9991FB089A24FFDEFE9EFA18FD97FA3A" ID-GBIF-Occurrence="1434659023" box="[304,541,1414,1438]" country="Mexico" location="MEXICO" pageId="22" pageNumber="23" specimenCount="1" typeStatus="paratype">
<typeStatus id="F6424FF79A24FFDEFE9EFA18FE2BFA3A" box="[304,417,1414,1438]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="51EEB1C59A24FFDEFE02FA18FD97FA3A" box="[428,541,1414,1438]" name="Mexico" pageId="22" pageNumber="23">MEXICO</collectingCountry>
</materialsCitation>
,
<materialsCitation id="9991FB089A24FFDEFD86FA1BFC91FA66" ID-GBIF-Occurrence="1434659041" collectingDate="1992-07-21" collectorName="G. Harp" country="Mexico" elevation="1300" latitude="21.051388" location="Tlanchinol" longLatPrecision="21" longitude="-98.66778" pageId="22" pageNumber="23" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Hidalgo" typeStatus="paratype">
<collectingRegion id="EB3D3FB79A24FFDEFD86FA1BFD08FA3A" box="[552,642,1413,1438]" country="Mexico" name="Hidalgo" pageId="22" pageNumber="23">Hidalgo</collectingRegion>
State:
<specimenCount id="3FFF3ADC9A24FFDEFD61FA1BFD71FA39" box="[719,763,1413,1438]" pageId="22" pageNumber="23" type="male">6 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A24FFDEFCA8FA18FCBAFA39" box="[774,816,1414,1438]" pageId="22" pageNumber="23" type="female">3 ♀</specimenCount>
, stream with cascade
<quantity id="EE015CB09A24FFDEFB88FA18FBD5FA3A" box="[1062,1119,1413,1438]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="km" value="7.0">7 km</quantity>
N of
<location id="2C26A78E9A24FFDEFB30FA1BFA9DFA3A" LSID="urn:lsid:plazi:treatment:A15040439A24FFD0FF39FBDCFBE9FA1B:2C26A78E9A24FFDEFB30FA1BFA9DFA3A" box="[1182,1303,1413,1438]" country="Mexico" latitude="21.051388" longLatPrecision="21" longitude="-98.66778" name="Tlanchinol" pageId="22" pageNumber="23" stateProvince="Hidalgo">Tlanchinol</location>
(
<geoCoordinate id="4CCD97929A24FFDEFA86FA1BFA12FA39" box="[1320,1432,1413,1438]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="21.051388">21°3'5'' N</geoCoordinate>
,
<geoCoordinate id="4CCD97929A24FFDEFF39FA34FE91FA65" box="[151,283,1450,1474]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-98.66778">98°40'4'' W</geoCoordinate>
,
<quantity id="EE015CB09A24FFDEFE86FA34FE02FA66" box="[296,392,1450,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="23" unit="m" value="1300.0">
<elevation id="A2D416669A24FFDEFE86FA34FE02FA66" box="[296,392,1450,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="23" unit="m" value="1300.0">1,300 m</elevation>
</quantity>
),
<date id="5D47D7959A24FFDEFE30FA34FDAFFA66" box="[414,549,1449,1474]" pageId="22" pageNumber="23" value="1992-07-21">
<collectingDate id="4D032E7D9A24FFDEFE30FA34FDAFFA66" box="[414,549,1449,1474]" pageId="22" pageNumber="23" value="1992-07-21">21 vii 1992</collectingDate>
</date>
,
<collectorName id="840C94839A24FFDEFD9FFA34FD00FA65" box="[561,650,1450,1474]" pageId="22" pageNumber="23">G. Harp</collectorName>
leg. [RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A24FFDEFC86FA34FEADFA42" ID-GBIF-Occurrence="1434659033" collectingDate="1992-07-22" collectingDateMax="1992-07-23" collectingDateMin="1992-07-22" collectorName="G. Harp" country="Mexico" elevation="1300" latitude="21.051388" location="Tlanchinol" longLatPrecision="21" longitude="-98.66778" pageId="22" pageNumber="23" specimenCount="2" specimenCount-male="2" stateProvince="Puebla" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A24FFDEFC86FA34FCDCFA65" box="[808,854,1450,1474]" pageId="22" pageNumber="23" type="male">2 ♂</specimenCount>
, same data but
<date id="5D47D7959A24FFDEFBA4FA34FB30FA66" box="[1034,1210,1449,1474]" pageId="22" pageNumber="23" value="1992-07-22" valueMax="1992-07-23" valueMin="1992-07-22">
<collectingDate id="4D032E7D9A24FFDEFBA4FA34FB30FA66" box="[1034,1210,1449,1474]" pageId="22" pageNumber="23" value="1992-07-22" valueMax="1992-07-23" valueMin="1992-07-22">2223 vii 1992</collectingDate>
</date>
, TWD leg. [RWG];
<collectingRegion id="EB3D3FB79A24FFDEFF39FA53FF69FA42" box="[151,227,1485,1510]" country="Mexico" name="Puebla" pageId="22" pageNumber="23">Puebla</collectingRegion>
State
</materialsCitation>
:
<materialsCitation id="9991FB089A24FFDEFE9FFA50FC31F9AE" ID-GBIF-Occurrence="1434659039" collectingDate="1987-07-23" collectionCode="CSCA" collectorName="Sierra de Huauchinango &amp; R. Novelo" country="Mexico" county="Zihuateutla" elevation="570" latitude="20.258612" location="Zona Cafetalera" longLatPrecision="21" longitude="-97.86806" municipality="Sierra de Huauchinango" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="La Union" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A24FFDEFE9FFA50FED7FA41" box="[305,349,1486,1510]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
,
<collectingCounty id="C02789D99A24FFDEFEC4FA53FE66FA42" box="[362,492,1485,1510]" pageId="22" pageNumber="23">Zihuateutla</collectingCounty>
,
<collectingMunicipality id="C9226B2F9A24FFDEFE56FA53FC80FA42" box="[504,778,1485,1510]" pageId="22" pageNumber="23">
<collectorName id="840C94839A24FFDEFE56FA53FC80FA42" box="[504,778,1485,1510]" pageId="22" pageNumber="23">Sierra de Huauchinango</collectorName>
</collectingMunicipality>
, km 3 road to.
<collectingRegion id="EB3D3FB79A24FFDEFC1AFA50FBA8FA42" box="[948,1058,1485,1510]" country="Philippines" name="La Union" pageId="22" pageNumber="23">La Unión</collectingRegion>
,
<location id="2C26A78E9A24FFDEFB83FA50FB6BFA42" LSID="urn:lsid:plazi:treatment:A15040439A24FFD0FF39FBDCFBE9FA1B:2C26A78E9A24FFDEFB83FA50FB6BFA42" box="[1069,1249,1485,1510]" country="Mexico" county="Zihuateutla" latitude="20.258612" longLatPrecision="21" longitude="-97.86806" municipality="Sierra de Huauchinango" name="Zona Cafetalera" pageId="22" pageNumber="23" stateProvince="La Union">Zona Cafetalera</location>
in drainage area (
<geoCoordinate id="4CCD97929A24FFDEFF31FA6FFEA6F9AD" box="[159,300,1521,1546]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="20.258612">20°15'31'' N</geoCoordinate>
,
<geoCoordinate id="4CCD97929A24FFDEFE96FA6FFE30F9AD" box="[312,442,1521,1546]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-97.86806">97°52'5'' W</geoCoordinate>
,
<quantity id="EE015CB09A24FFDEFE6BFA6FFD85F9AE" box="[453,527,1521,1546]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="22" pageNumber="23" unit="m" value="570.0">
<elevation id="A2D416669A24FFDEFE6BFA6FFD85F9AE" box="[453,527,1521,1546]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="22" pageNumber="23" unit="m" value="570.0">570 m</elevation>
</quantity>
),
<date id="5D47D7959A24FFDEFD8DFA6CFD2CF9AE" box="[547,678,1521,1546]" pageId="22" pageNumber="23" value="1987-07-23">
<collectingDate id="4D032E7D9A24FFDEFD8DFA6CFD2CF9AE" box="[547,678,1521,1546]" pageId="22" pageNumber="23" value="1987-07-23">23 vii 1987</collectingDate>
</date>
,
<collectorName id="840C94839A24FFDEFD1FFA6CFCAEF9AE" box="[689,804,1521,1546]" pageId="22" pageNumber="23">R. Novelo</collectorName>
leg. [
<collectionCode id="4FE869909A24FFDEFCCCFA6CFC3EF9AE" box="[866,948,1522,1546]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="22" pageNumber="23">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A24FFDEFC69FA6CFB7CF9AE" ID-GBIF-Occurrence="1434659035" box="[967,1270,1521,1546]" collectingDate="1987-07-23" collectionCode="CSCA" collectorName="Sierra de Huauchinango &amp; R. Novelo" country="Mexico" county="Zihuateutla" elevation="570" latitude="20.258612" location="Zona Cafetalera" longLatPrecision="21" longitude="-97.86806" municipality="Sierra de Huauchinango" pageId="22" pageNumber="23" specimenCount="4" specimenCount-male="4" stateProvince="La Union" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A24FFDEFC69FA6CFC78F9AD" box="[967,1010,1522,1546]" pageId="22" pageNumber="23" type="male">4 ♂</specimenCount>
, same data but [RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A24FFDEFAAFFA6CFCDFF98A" ID-GBIF-Occurrence="1434659040" country="Mexico" location="MEXICO" pageId="22" pageNumber="23" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Veracruz" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A24FFDEFAAFFA6CFAA7F9AD" box="[1281,1325,1522,1546]" pageId="22" pageNumber="23" type="male">3 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A24FFDEFA97FA6CFAE9F9AD" box="[1337,1379,1522,1546]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(one pair in tandem), same data as holotype [RWG];
<collectingRegion id="EB3D3FB79A24FFDEFD01F988FC99F989" box="[687,787,1558,1581]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="22" pageNumber="23">Veracruz</collectingRegion>
State
</materialsCitation>
:
<materialsCitation id="9991FB089A24FFDEFCF1F988FF7BF9D2" ID-GBIF-Occurrence="1434659021" collectingDate="2000-07-15" collectorName="El Muro &amp; R. Novelo" country="Mexico" elevation="1100" latitude="19.878056" location="Tlapacoyan-Altotonga" longLatPrecision="21" longitude="-97.22806" municipality="El Muro" pageId="22" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A24FFDEFCF1F988FC01F989" box="[863,907,1558,1582]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
,
<specimenCount id="3FFF3ADC9A24FFDEFC39F988FC4BF989" box="[919,961,1558,1582]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
(in tandem, female ovipositing),
<collectingMunicipality id="C9226B2F9A24FFDEFA99F98BFA12F989" box="[1335,1432,1557,1582]" pageId="22" pageNumber="23">
<collectorName id="840C94839A24FFDEFA99F98BFA12F989" box="[1335,1432,1557,1582]" pageId="22" pageNumber="23">El Muro</collectorName>
</collectingMunicipality>
, km 14 on road 131
<location id="2C26A78E9A24FFDEFE28F9A7FD08F9F6" LSID="urn:lsid:plazi:treatment:A15040439A24FFD0FF39FBDCFBE9FA1B:2C26A78E9A24FFDEFE28F9A7FD08F9F6" box="[390,642,1593,1618]" country="Mexico" latitude="19.878056" longLatPrecision="21" longitude="-97.22806" municipality="El Muro" name="Tlapacoyan-Altotonga" pageId="22" pageNumber="23" stateProvince="Veracruz">Tlapacoyan-Altotonga</location>
(
<geoCoordinate id="4CCD97929A24FFDEFD38F9A7FCADF9F5" box="[662,807,1593,1618]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="19.878056">19°52'41'' N</geoCoordinate>
,
<geoCoordinate id="4CCD97929A24FFDEFC99F9A4FC41F9F5" box="[823,971,1594,1618]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-97.22806">97°13'41'' W</geoCoordinate>
,
<quantity id="EE015CB09A24FFDEFC74F9A4FBB4F9F6" box="[986,1086,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">
<elevation id="A2D416669A24FFDEFC74F9A4FBB4F9F6" box="[986,1086,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
),
<date id="5D47D7959A24FFDEFBF8F9A7FB6BF9F6" box="[1110,1249,1593,1618]" pageId="22" pageNumber="23" value="2000-07-15">
<collectingDate id="4D032E7D9A24FFDEFBF8F9A7FB6BF9F6" box="[1110,1249,1593,1618]" pageId="22" pageNumber="23" value="2000-07-15">15 vii 2000</collectingDate>
</date>
,
<collectorName id="840C94839A24FFDEFB5EF9A4FAEDF9F6" box="[1264,1383,1593,1618]" pageId="22" pageNumber="23">R. Novelo</collectorName>
leg. [RWG].
</materialsCitation>
</paragraph>
<paragraph id="2946F1559A24FFDEFF69F91CFDF2F946" blockId="22.[151,1437,1233,2014]" pageId="22" pageNumber="23">
A medium-sized largely violaceous-blue species (
<figureCitation id="B1C2EDD09A24FFDEFCB2F91FFC0BF93E" box="[796,897,1665,1690]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="22" pageNumber="23">Figs. 19</figureCitation>
,
<figureCitation id="B1C2EDD09A24FFDEFC3BF91FFC3EF93E" box="[917,948,1665,1690]" captionStart="FIGURES 63 72" captionStartId="54.[151,264,1847,1870]" captionTargetBox="[162,1430,216,1811]" captionTargetId="figure@54.[151,1436,193,1826]" captionTargetPageId="54" captionText="FIGURES 63 72. Color pattern of male S 7 10 of extranea - group species, lateral view." httpUri="https://zenodo.org/record/322077/files/figure.png" pageId="22" pageNumber="23">68</figureCitation>
, based on well-preserved material) with approximate cerci (
<figureCitation id="B1C2EDD09A24FFDEFEDDF93BFE59F91A" box="[371,467,1701,1726]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="22" pageNumber="23">Fig. 154</figureCitation>
) the male of which is similar to
<taxonomicName id="EEF98AD69A24FFDEFCE5F938FC33F919" box="[843,953,1702,1725]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A24FFDEFCE5F938FC33F919" box="[843,953,1702,1725]" italics="true" pageId="22" pageNumber="23">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFC64F93BFBA0F91A" box="[970,1066,1701,1726]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="22" pageNumber="23">Figs. 20</figureCitation>
; 69; 181),
<taxonomicName id="EEF98AD69A24FFDEFB08F938FB71F91A" box="[1190,1275,1702,1726]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A24FFDEFB08F938FB71F91A" box="[1190,1275,1702,1726]" italics="true" pageId="22" pageNumber="23">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFAA5F93BFAE1F91A" box="[1291,1387,1701,1726]" captionStart="FIGURES 21 23" captionStartId="44.[151,264,1842,1865]" captionTargetBox="[179,1400,210,1687]" captionTargetId="figure@44.[151,1436,193,1821]" captionTargetPageId="44" captionText="FIGURES 21 23. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322067/files/figure.png" pageId="22" pageNumber="23">Figs. 21</figureCitation>
; 70; 183) and
<taxonomicName id="EEF98AD69A24FFDEFEAEF954FEF4F945" box="[256,382,1737,1761]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A24FFDEFEAEF954FEF4F945" box="[256,382,1737,1761]" italics="true" pageId="22" pageNumber="23">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFE20F957FD9FF946" box="[398,533,1737,1762]" captionStart="FIGURES 21 23" captionStartId="44.[151,264,1842,1865]" captionTargetBox="[179,1400,210,1687]" captionTargetId="figure@44.[151,1436,193,1821]" captionTargetPageId="44" captionText="FIGURES 21 23. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322067/files/figure.png" pageId="22" pageNumber="23">Figs. 22, 23</figureCitation>
,
<figureCitation id="B1C2EDD09A24FFDEFD8FF954FDE1F946" box="[545,619,1738,1762]" captionStart="FIGURES 63 72" captionStartId="54.[151,264,1847,1870]" captionTargetBox="[162,1430,216,1811]" captionTargetId="figure@54.[151,1436,193,1826]" captionTargetPageId="54" captionText="FIGURES 63 72. Color pattern of male S 7 10 of extranea - group species, lateral view." httpUri="https://zenodo.org/record/322077/files/figure.png" pageId="22" pageNumber="23">71, 72</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A24FFD0FF69F973FF52FF70" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="2946F1559A24FFDEFF69F973FD03F832" blockId="22.[151,1437,1233,2014]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69F973FDAAF8A2" bold="true" box="[199,544,1773,1798]" pageId="22" pageNumber="23">
Description of male
<typeStatus id="F6424FF79A24FFDEFE14F973FDAAF8A2" box="[442,544,1773,1798]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
</emphasis>
. Head (
<figureCitation id="B1C2EDD09A24FFDEFDD4F973FD44F8A2" box="[634,718,1773,1798]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="22" pageNumber="23">Fig. 19</figureCitation>
): Labium pale except for small black medial spot at base; ante- and postclypeus, antefrons, base of mandibles, anterior half of scape and genae pale, remainder of head black with violaceous postocular spots confluent with eyes, incomplete pale occipital bar and small pale spot anterolateral to lateral ocellus; posterior half of scape and remainder of antennae black, rear of head washed with black, especially medially, margin bordering eye margin pale.
</paragraph>
<paragraph id="2946F1559A24FFDFFF69F83FFECCFE77" blockId="22.[151,1437,1233,2014]" lastBlockId="23.[151,1437,151,2018]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Prothorax black with following areas pale: obscure lateral spot on anterior lobe, dorsolateral spot on middle lobe, lateral 0.30 of posterior margin of pronotum, propleuron except for notopleural suture. Pale areas of pterothorax violaceous blue, with black middorsal stripe about as wide as pale antehumeral stripe, the latter slightly narrowing dorsally; anterior half of mesinfraepisternum black, continuous with broad humeral stripe at base, then narrowing at basal fifth along mesopleural suture, then abruptly forming a thin hairline at 0.50 before expanding again at mesopleural fossa and connecting narrowly below antealar crest with middorsal stripe above; metapleural suture with a narrow black stripe separating paler areas on side of thorax (
<figureCitation id="B1C2EDD09A25FFDFFC44FEB9FBB5FEE4" box="[1002,1087,295,320]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
). Wings hyaline with venation black; pterostigma dark brown, surmounting 2 cells in both Fw and 2 cells in left Hw [distal 0.80 of right Hw missing]; postnodals Fw 18/19, left Hw 16; postquadrangular cells Fw 4/4, left Hw 3; RP2 at Fw 8/8, left Hw 6. Coxae and trochanters pale medially washed with brown; extensor surfaces of tibiae pale, remainder of legs and armature black.
</paragraph>
<paragraph id="2946F1559A25FFDFFF69FE40FA17FCB3" blockId="23.[151,1437,151,2018]" pageId="23" pageNumber="24">
Abdomen (
<figureCitation id="B1C2EDD09A25FFDFFEE8FE40FE2FFE53" box="[326,421,478,503]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="23" pageNumber="24">Figs. 19</figureCitation>
,
<figureCitation id="B1C2EDD09A25FFDFFE1DFE40FE5AFE53" box="[435,464,478,503]" captionStart="FIGURES 63 72" captionStartId="54.[151,264,1847,1870]" captionTargetBox="[162,1430,216,1811]" captionTargetId="figure@54.[151,1436,193,1826]" captionTargetPageId="54" captionText="FIGURES 63 72. Color pattern of male S 7 10 of extranea - group species, lateral view." httpUri="https://zenodo.org/record/322077/files/figure.png" pageId="23" pageNumber="24">68</figureCitation>
) mostly violaceous blue; S1 pale with narrow black basal ring expanding dorsally to form a connected black spot occupying basal 0.50; S2 pale above with an incomplete black lateral stripe widening posteriorly and ending at distal 0.30, this stripe connected to obscure black spot below thus isolating pale areas anteriorly and posteriorly, genital lobe not strongly produced, apical annulus black; S3 pale with an narrow lateral black stripe, apical 0.20 including annulus black except for small isolated transverse spot before annulus; S4 similar to S3 but with remnant of postbasal streak; S5 similar but postbasal streak more extensive with vestigial line connecting to apical black; S6 with postbasal black confluent with apical black thus forming a broad ventral stripe; S7 black except for narrow pale ring with a broad triangular extension middorsally ending at basal 0.20; S8 10 pale dorsolaterally and black ventrally; torus pale, cerci black; dorsal portion of paraprocts pale, ventrally black.
</paragraph>
<paragraph id="2946F1559A25FFDFFF69FCBCFE73FCC4" blockId="23.[151,1437,151,2018]" pageId="23" pageNumber="24">
Genital ligula (
<figureCitation id="B1C2EDD09A25FFDFFEDBFCBCFE5CFC9F" box="[373,470,802,827]" captionStart="FIGURES 134 137" captionStartId="71.[151,264,1936,1959]" captionTargetBox="[151,1417,205,1896]" captionTargetId="figure@71.[151,1436,193,1916]" captionTargetPageId="71" captionText="FIGURES 134 137. Genital ligula of extranea - group species; complete view (134, 137) and partial view of apex (135, 136). (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322094/files/figure.png" pageId="23" pageNumber="24">Fig. 137</figureCitation>
) similar to
<taxonomicName id="EEF98AD69A25FFDFFDF1FCBDFD68FC9F" box="[607,738,803,827]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="1B8D2D479A25FFDFFDF1FCBDFD68FC9F" box="[607,738,803,827]" italics="true" pageId="23" pageNumber="24">A. elongata</emphasis>
</taxonomicName>
(as in
<figureCitation id="B1C2EDD09A25FFDFFC9BFCBCFC1CFC9F" box="[821,918,802,827]" captionStart="FIGURES 132 133" captionStartId="70.[151,264,1765,1788]" captionTargetBox="[152,1423,219,1645]" captionTargetId="figure@70.[151,1436,193,1744]" captionTargetPageId="70" captionText="FIGURES 132 133. Genital ligula of extranea - group species. (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322093/files/figure.png" pageId="23" pageNumber="24">Fig. 132</figureCitation>
) but tip narrower. Lateral lobes appressed to lateral margin of distal segment
</paragraph>
<paragraph id="2946F1559A25FFDFFF69FCF5FF75FBFF" blockId="23.[151,1437,151,2018]" pageId="23" pageNumber="24">
Torus large, transversely triangular, swollen ventrally, occupying ventral half of torifer and overlapping recessed bilobed epiproct (
<figureCitation id="B1C2EDD09A25FFDFFE6CFC10FDAEFC03" box="[450,548,910,935]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="23" pageNumber="24">Fig. 154</figureCitation>
c); narrow area around epiproct and extreme base of same black; cercus roughly quadrate in dorsal view with outer margin about 1.5 times as long as inner margin (
<figureCitation id="B1C2EDD09A25FFDFFB98FC2CFB12FC6F" box="[1078,1176,946,971]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="23" pageNumber="24">Fig. 154</figureCitation>
c); in mediodorsal view upper surface slightly concave with a broadly based tooth at basal 0.20 (
<figureCitation id="B1C2EDD09A25FFDFFC73FC48FBCAFC4B" box="[989,1088,982,1007]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="23" pageNumber="24">Fig. 154</figureCitation>
a, d); in lateral view, narrowly triangular, subequal to length of paraproct (
<figureCitation id="B1C2EDD09A25FFDFFD2FFC64FD6EFBB7" box="[641,740,1018,1043]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="23" pageNumber="24">Fig. 154</figureCitation>
b); paraproct slightly bilobed, its ventral branch much smaller than longer, anteriorly directed, bluntly triangular dorsal branch, its tip, viewed mediodorsally (
<figureCitation id="B1C2EDD09A25FFDFFB6EFB80FAABFB93" box="[1216,1313,1054,1079]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="23" pageNumber="24">Fig. 154</figureCitation>
a), directed medially.
</paragraph>
<paragraph id="2946F1559A25FFDFFF69FBF8FCAEFBDB" blockId="23.[151,1437,151,2018]" box="[199,804,1126,1151]" pageId="23" pageNumber="24">
<emphasis id="1B8D2D479A25FFDFFF69FBF8FED8FBDB" bold="true" box="[199,338,1126,1151]" pageId="23" pageNumber="24">Dimensions</emphasis>
. Hw 21.5, abdomen 30, total length 37.5.
</paragraph>
<paragraph id="2946F1559A25FFDFFF69FB14FC54F98B" blockId="23.[151,1437,151,2018]" pageId="23" pageNumber="24">
<emphasis id="1B8D2D479A25FFDFFF69FB14FDC5FB07" bold="true" box="[199,591,1162,1187]" pageId="23" pageNumber="24">
Description of female
<typeStatus id="F6424FF79A25FFDFFE4AFB14FDC5FB07" box="[484,591,1162,1187]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
</emphasis>
(
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<collectingRegion id="EB3D3FB79A25FFDFFDC8FB14FD4BFB07" box="[614,705,1162,1187]" country="Mexico" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingRegion>
</collectingCountry>
:
<collectingRegion id="EB3D3FB79A25FFDFFD7AFB15FCB2FB06" box="[724,824,1163,1186]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="24">Veracruz</collectingRegion>
State, El Muro, km 14 on road 131 Tlapacoyan- Altotonga). Similar to male but pale color light blue. Head (
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) as in male but with a black basal spot mediolaterally on labrum and postocular spots larger and confluent with eye, prothorax as in male but black areas less extensive, pterothorax with humeral stripe narrower and more deeply forked; S1 as in male; S2 with dorsal oval spot at basal 0.60, apically and laterally with black interrupted below by a pale lateral stripe narrowed at apical 0.30, pale lateral stripe bordered below by a black stripe; S3 dorsally pale with apical 0.30 black, the latter with a ventral offshoot of black along ventral margin and expanding at basal 0.10 and almost connecting with an elongate postbasal streak, which is at first broad but narrows to a thin line just connecting with apical black; S45 similar to S3 but black more extensive, isolating a pale midlateral stripe; dorsal blue also reduced, especially apically, but with an incomplete pale basal ring; S67 black with an incomplete pale basal ring; S8 blue dorsally, black at basal 0.20 and laterally; S9 blue dorsally with a black dorsolateral spot at basal 0.50 and connected to lateral stripe below; S10 blue dorsally, black laterally; appendages and ovipositor black.
</paragraph>
<paragraph id="2946F1559A25FFDFFF69F9A4FAE7F91B" blockId="23.[151,1437,151,2018]" pageId="23" pageNumber="24">
Mesostigmal lobe forming a broad oval (
<figureCitation id="B1C2EDD09A25FFDFFD3BF9A4FD7FF9F7" box="[661,757,1594,1619]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
a, b), about twice as wide as high, occupying medial 0.60 of posterior margin of mesostigmal plate (
<figureCitation id="B1C2EDD09A25FFDFFDF8F9C0FD32F9D3" box="[598,696,1630,1655]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
c), medial margin thickened with a slightly recurved margin (
<figureCitation id="B1C2EDD09A25FFDFFAC1F9C0FF48F93F" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
d, e), and slightly overlying fork of middorsal carina; distance between mesostigmal lobes small, barely separated (
<figureCitation id="B1C2EDD09A25FFDFFEBEF938FEE5F91B" box="[272,367,1702,1727]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="23" pageNumber="24">Fig. 113</figureCitation>
a, c); in lateral view, lobe erect with posterior margin curving medioposteriorly (
<figureCitation id="B1C2EDD09A25FFDFFB5CF938FAD8F91B" box="[1266,1362,1702,1727]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="23" pageNumber="24">
Fig.
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</figureCitation>
g).
</paragraph>
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<emphasis id="1B8D2D479A25FFDFFF69F954FE62F947" bold="true" box="[199,488,1738,1763]" pageId="23" pageNumber="24">
Variation in
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</emphasis>
. Little variation was observed in the
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series. One male (Sierra de Huauchinango) had a small indication of a forked humeral stripe; another male (same locality) had reduced black abdominal markings thus isolating black postbasal streaks on S45 similar to S
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<figureCitation id="B1C2EDD09A25FFDFFBEEF88CFB18F88F" box="[1088,1170,1810,1835]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
. The small lateral lobes on the genital ligula of the
<typeStatus id="F6424FF79A25FFDFFE7CF8A8FDBEF8EB" box="[466,564,1846,1871]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(
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b) are adjacent to the sides of the ligula, but in
<typeStatus id="F6424FF79A25FFDFFB62F8A6FABDF8F4" box="[1228,1335,1848,1872]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
they are usually separated basally as in
<taxonomicName id="EEF98AD69A25FFDFFDA8F8C5FD06F8D7" box="[518,652,1883,1907]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="1B8D2D479A25FFDFFDA8F8C5FD06F8D7" box="[518,652,1883,1907]" italics="true" pageId="23" pageNumber="24">A. elongata</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A25FFDFFD31F8C4FC8FF8D7" box="[671,773,1882,1907]" captionStart="FIGURES 132 133" captionStartId="70.[151,264,1765,1788]" captionTargetBox="[152,1423,219,1645]" captionTargetId="figure@70.[151,1436,193,1744]" captionTargetPageId="70" captionText="FIGURES 132 133. Genital ligula of extranea - group species. (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322093/files/figure.png" pageId="23" pageNumber="24">Fig. 132</figureCitation>
b). Females can be blue (
<figureCitation id="B1C2EDD09A25FFDFFB9EF8C4FB0CF8D7" box="[1072,1158,1882,1907]" captionStart="FIGURES 43 - 46" captionStartId="50.[151,264,1915,1938]" captionTargetBox="[172,1436,193,1880]" captionTargetId="figure@50.[151,1436,193,1894]" captionTargetPageId="50" captionText="FIGURES 43 - 46. Color pattern of female head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322073/files/figure.png" pageId="23" pageNumber="24">Fig. 43</figureCitation>
) or brown. Pterostigma surmounting 12 cells in males and females; postnodals: Fw
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males and females, Hw
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males,
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females; postquadrangular cells Fw 35, Hw
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males, Fw 4, Hw
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females; RP2 at Fw 78, Hw
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males and females.
<emphasis id="1B8D2D479A25FFDFFED9F857FD88F846" bold="true" box="[375,514,1993,2018]" pageId="23" pageNumber="24">Dimensions</emphasis>
.
<specimenCount id="3FFF3ADC9A25FFDFFDBEF854FDA2F845" box="[528,552,1994,2017]" pageId="23" pageNumber="24" type="male"></specimenCount>
: Hw 23 ± 1.95 [2024], abdomen 30.3 ± 1.69 [27.633.1], total length 38.4 ± 2.17 [35.441.9];
<specimenCount id="3FFF3ADC9A2AFFD0FEF1FF06FEFCFF0B" box="[351,374,152,175]" pageId="24" pageNumber="25" type="female"></specimenCount>
: Hw 24.8 ± 1.74 [22.326.5], abdomen 29.4 ± 2.11 [28.631.3], total length 37.7 ± 2.73 [33.5 40.1].
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A2AFFD0FF69FF41FB90FADC" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="2946F1559A2AFFD0FF69FF41FBB7FCF8" blockId="24.[151,1437,151,1472]" pageId="24" pageNumber="25">
<emphasis id="1B8D2D479A2AFFD0FF69FF41FEB0FF5C" bold="true" box="[199,314,223,248]" pageId="24" pageNumber="25">Diagnosis</emphasis>
. This species may be confused with three other largely blue species, two of which,
<taxonomicName id="EEF98AD69A2AFFD0FB4FFF7EFAC4FF5C" box="[1249,1358,224,248]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FB4FFF7EFAC4FF5C" box="[1249,1358,224,248]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A2AFFD0FA2AFF7EFF7CFEB8" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FA2AFF7EFF7CFEB8" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
, are sympatric with
<taxonomicName id="EEF98AD69A2AFFD0FE74FE9AFDD3FEB8" box="[474,601,260,284]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FE74FE9AFDD3FEB8" box="[474,601,260,284]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FDC6FE9AFD40FEB9" box="[616,714,260,285]" captionStart="FIGURES 167 168" captionStartId="83.[151,264,1723,1746]" captionTargetBox="[209,1344,219,1653]" captionTargetId="figure@83.[188,1398,193,1702]" captionTargetPageId="83" captionText="FIGURES 167 168. Distribution of extranea-group species." httpUri="https://zenodo.org/record/322106/files/figure.png" pageId="24" pageNumber="25">Fig. 168</figureCitation>
). The cerci in male
<taxonomicName id="EEF98AD69A2AFFD0FC09FE9AFBACFEB8" box="[935,1062,260,284]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FC09FE9AFBACFEB8" box="[935,1062,260,284]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
are more approximate, with their inner surfaces nearly touching in normal position (
<figureCitation id="B1C2EDD09A2AFFD0FD7BFEB9FCB2FEE4" box="[725,824,295,320]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 154</figureCitation>
c); in both
<taxonomicName id="EEF98AD69A2AFFD0FC1CFEB6FBABFEE4" box="[946,1057,296,320]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FC1CFEB6FBABFEE4" box="[946,1057,296,320]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FB9EFEB9FB1BFEE4" box="[1072,1169,295,320]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 155</figureCitation>
c) and
<taxonomicName id="EEF98AD69A2AFFD0FB71FEB6FAD4FEE4" box="[1247,1374,296,320]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FB71FEB6FAD4FEE4" box="[1247,1374,296,320]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FAC0FEB9FF4EFEC1" captionStart="FIGURES 156 157" captionStartId="79.[151,264,1901,1924]" captionTargetBox="[158,1414,203,1867]" captionTargetId="figure@79.[151,1436,193,1880]" captionTargetPageId="79" captionText="FIGURES 156 157. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322102/files/figure.png" pageId="24" pageNumber="25">Fig. 157</figureCitation>
c), the cerci are widely separated by a gap subequal to the basal width of each cercus. In dorsomedial view, the cercus of
<taxonomicName id="EEF98AD69A2AFFD0FEA9FEEEFE02FE2C" box="[263,392,368,392]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FEA9FEEEFE02FE2C" box="[263,392,368,392]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FE37FEF1FE70FE2C" box="[409,506,367,392]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 154</figureCitation>
d) is less elongate than in either
<taxonomicName id="EEF98AD69A2AFFD0FCC0FEEEFC56FE2C" box="[878,988,368,392]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FCC0FEEEFC56FE2C" box="[878,988,368,392]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FC43FEF1FBC7FE2C" box="[1005,1101,367,392]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 155</figureCitation>
d) or
<taxonomicName id="EEF98AD69A2AFFD0FB23FEEEFA87FE2C" box="[1165,1293,368,392]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FB23FEEEFA87FE2C" box="[1165,1293,368,392]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FAB3FEF1FA0AFE2C" box="[1309,1408,367,392]" captionStart="FIGURES 156 157" captionStartId="79.[151,264,1901,1924]" captionTargetBox="[158,1414,203,1867]" captionTargetId="figure@79.[151,1436,193,1880]" captionTargetPageId="79" captionText="FIGURES 156 157. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322102/files/figure.png" pageId="24" pageNumber="25">Fig. 157</figureCitation>
d). The nearly approximate condition of the cerci in
<taxonomicName id="EEF98AD69A2AFFD0FD62FE0AFCC4FE08" box="[716,846,404,428]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FD62FE0AFCC4FE08" box="[716,846,404,428]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FCF1FE0AFC4BFE09" box="[863,961,404,429]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 154</figureCitation>
d) is similar to that of the more southerly (
<collectingCountry id="51EEB1C59A2AFFD0FF30FE26FE91FE74" box="[158,283,439,464]" name="Costa Rica" pageId="24" pageNumber="25">
<collectingRegion id="EB3D3FB79A2AFFD0FF30FE26FF6AFE74" box="[158,224,440,464]" country="New Zealand" name="West Coast" pageId="24" pageNumber="25">Costa</collectingRegion>
Rica
</collectingCountry>
to
<collectingCountry id="51EEB1C59A2AFFD0FEEFFE29FE3BFE74" box="[321,433,439,464]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
)
<taxonomicName id="EEF98AD69A2AFFD0FE6EFE26FDD2FE74" authority="Selys" authorityName="Selys" box="[448,600,439,464]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FE6EFE26FD9EFE74" box="[448,532,440,464]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
Selys
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FDC6FE29FD40FE74" box="[616,714,439,464]" captionStart="FIGURES 156 157" captionStartId="79.[151,264,1901,1924]" captionTargetBox="[158,1414,203,1867]" captionTargetId="figure@79.[151,1436,193,1880]" captionTargetPageId="79" captionText="FIGURES 156 157. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322102/files/figure.png" pageId="24" pageNumber="25">Fig. 156</figureCitation>
c), but the transverse width of the torus in
<taxonomicName id="EEF98AD69A2AFFD0FB07FE26FB77FE74" box="[1193,1277,440,464]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FB07FE26FB77FE74" box="[1193,1277,440,464]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FAA3FE29FAE7FE74" box="[1293,1389,439,464]" captionStart="FIGURES 156 157" captionStartId="79.[151,264,1901,1924]" captionTargetBox="[158,1414,203,1867]" captionTargetId="figure@79.[151,1436,193,1880]" captionTargetPageId="79" captionText="FIGURES 156 157. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322102/files/figure.png" pageId="24" pageNumber="25">Fig. 156</figureCitation>
c) is subequal to the transverse width of each cercus; in
<taxonomicName id="EEF98AD69A2AFFD0FD77FE42FCD3FE50" box="[729,857,476,500]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FD77FE42FCD3FE50" box="[729,857,476,500]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
the transverse width of each torus is about half the transverse width of each cercus (
<figureCitation id="B1C2EDD09A2AFFD0FDA5FE61FDE7FDBC" box="[523,621,511,536]" captionStart="FIGURES 154 155" captionStartId="78.[151,264,1933,1956]" captionTargetBox="[152,1420,208,1899]" captionTargetId="figure@78.[151,1436,193,1912]" captionTargetPageId="78" captionText="FIGURES 154 155. Male caudal appendages of extranea-group species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, mediodorsal view." httpUri="https://zenodo.org/record/322101/files/figure.png" pageId="24" pageNumber="25">Fig. 154</figureCitation>
c). The tip of the terminal segment of the genital ligula in
<taxonomicName id="EEF98AD69A2AFFD0FAAEFD9EFA0AFDBC" box="[1280,1408,512,536]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FAAEFD9EFA0AFDBC" box="[1280,1408,512,536]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
is short and narrow, ending in a more or less ridged trough-shaped tip (
<figureCitation id="B1C2EDD09A2AFFD0FC0EFDBAFB8EFD98" box="[928,1028,548,573]" captionStart="FIGURES 134 137" captionStartId="71.[151,264,1936,1959]" captionTargetBox="[151,1417,205,1896]" captionTargetId="figure@71.[151,1436,193,1916]" captionTargetPageId="71" captionText="FIGURES 134 137. Genital ligula of extranea - group species; complete view (134, 137) and partial view of apex (135, 136). (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322094/files/figure.png" pageId="24" pageNumber="25">Fig. 137</figureCitation>
); the tip in the other three species is longer and flexible and can assume the shapes illustrated here (
<taxonomicName id="EEF98AD69A2AFFD0FCF8FDD6FC48FDC4" box="[854,962,584,608]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FCF8FDD6FC48FDC4" box="[854,962,584,608]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
<figureCitation id="B1C2EDD09A2AFFD0FC67FDD9FBE6FDC4" box="[969,1132,583,608]" captionStart="FIGURES 134 137" captionStartId="71.[151,264,1936,1959]" captionTargetBox="[151,1417,205,1896]" captionTargetId="figure@71.[151,1436,193,1916]" captionTargetPageId="71" captionText="FIGURES 134 137. Genital ligula of extranea - group species; complete view (134, 137) and partial view of apex (135, 136). (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322094/files/figure.png" pageId="24" pageNumber="25">Figs. 134136</figureCitation>
; essentially the same for
<taxonomicName id="EEF98AD69A2AFFD0FA2AFDD6FF41FD20" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FA2AFDD6FF41FD20" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A2AFFD0FEACFDF2FE0BFD20" box="[258,385,620,644]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FEACFDF2FE0BFD20" box="[258,385,620,644]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
). The ectobasal portion of the terminal segment of the genital ligula in
<taxonomicName id="EEF98AD69A2AFFD0FB05FDF2FA93FD20" box="[1195,1305,620,644]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FB05FDF2FA93FD20" box="[1195,1305,620,644]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FA87FDF2FA00FD20" box="[1321,1418,620,645]" captionStart="FIGURES 134 137" captionStartId="71.[151,264,1936,1959]" captionTargetBox="[151,1417,205,1896]" captionTargetId="figure@71.[151,1436,193,1916]" captionTargetPageId="71" captionText="FIGURES 134 137. Genital ligula of extranea - group species; complete view (134, 137) and partial view of apex (135, 136). (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322094/files/figure.png" pageId="24" pageNumber="25">Fig. 134</figureCitation>
a, b),
<taxonomicName id="EEF98AD69A2AFFD0FF10FD0EFE9BFD0C" box="[190,273,656,680]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FF10FD0EFE9BFD0C" box="[190,273,656,680]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
, and
<taxonomicName id="EEF98AD69A2AFFD0FEFCFD0EFE58FD0C" box="[338,466,656,680]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FEFCFD0EFE58FD0C" box="[338,466,656,680]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
has a prominent narrow mound-like lobe that is absent in
<taxonomicName id="EEF98AD69A2AFFD0FBDEFD0EFB7AFD0C" box="[1136,1264,656,680]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FBDEFD0EFB7AFD0C" box="[1136,1264,656,680]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FAACFD11FAEEFD0C" box="[1282,1380,655,680]" captionStart="FIGURES 134 137" captionStartId="71.[151,264,1936,1959]" captionTargetBox="[151,1417,205,1896]" captionTargetId="figure@71.[151,1436,193,1916]" captionTargetPageId="71" captionText="FIGURES 134 137. Genital ligula of extranea - group species; complete view (134, 137) and partial view of apex (135, 136). (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322094/files/figure.png" pageId="24" pageNumber="25">Fig. 137</figureCitation>
a, b). The following field markings will also distinguish male
<taxonomicName id="EEF98AD69A2AFFD0FCBAFD2AFC1EFD68" box="[788,916,692,716]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FCBAFD2AFC1EFD68" box="[788,916,692,716]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
from
<taxonomicName id="EEF98AD69A2AFFD0FC74FD2AFBC2FD68" box="[986,1096,692,716]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FC74FD2AFBC2FD68" box="[986,1096,692,716]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF98AD69A2AFFD0FBF8FD2AFB20FD68" box="[1110,1194,692,716]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FBF8FD2AFB20FD68" box="[1110,1194,692,716]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A2AFFD0FB4CFD2AFA16FD69" authority="The" authorityName="The" box="[1250,1436,692,717]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FB4CFD2AFAEBFD68" box="[1250,1377,692,716]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
: The
</taxonomicName>
black postbasal streaks (sometimes extensive) present on S
<quantity id="EE015CB09A2AFFD0FC90FD49FC00FD54" box="[830,906,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="24" pageNumber="25" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">46 in</quantity>
<taxonomicName id="EEF98AD69A2AFFD0FC3DFD46FB9FFD54" box="[915,1045,728,752]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FC3DFD46FB9FFD54" box="[915,1045,728,752]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FB86FD49FBF6FD54" box="[1064,1148,727,752]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="24" pageNumber="25">Fig. 19</figureCitation>
) are absent in
<taxonomicName id="EEF98AD69A2AFFD0FA82FD46FA16FD54" box="[1324,1436,728,752]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FA82FD46FA16FD54" box="[1324,1436,728,752]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FF31FD62FE8AFCB0" box="[159,256,764,789]" captionStart="FIGURES 17 20" captionStartId="43.[151,264,1898,1921]" captionTargetBox="[161,1389,208,1860]" captionTargetId="figure@43.[151,1436,193,1877]" captionTargetPageId="43" captionText="FIGURES 17 20. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322066/files/figure.png" pageId="24" pageNumber="25">Figs. 20</figureCitation>
; 181),
<taxonomicName id="EEF98AD69A2AFFD0FEFFFD62FE2DFCB0" box="[337,423,764,788]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FEFFFD62FE2DFCB0" box="[337,423,764,788]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FE16FD62FD92FCB0" box="[440,536,764,789]" captionStart="FIGURES 21 23" captionStartId="44.[151,264,1842,1865]" captionTargetBox="[179,1400,210,1687]" captionTargetId="figure@44.[151,1436,193,1821]" captionTargetPageId="44" captionText="FIGURES 21 23. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322067/files/figure.png" pageId="24" pageNumber="25">Figs. 21</figureCitation>
; 183) and
<taxonomicName id="EEF98AD69A2AFFD0FD3AFD62FC9FFCB0" box="[660,789,764,788]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FD3AFD62FC9FFCB0" box="[660,789,764,788]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FC88FD62FC39FCB0" box="[806,947,764,789]" captionStart="FIGURES 21 23" captionStartId="44.[151,264,1842,1865]" captionTargetBox="[179,1400,210,1687]" captionTargetId="figure@44.[151,1436,193,1821]" captionTargetPageId="44" captionText="FIGURES 21 23. Color pattern of male head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322067/files/figure.png" pageId="24" pageNumber="25">Figs. 22, 23</figureCitation>
); the wings are hyaline in
<taxonomicName id="EEF98AD69A2AFFD0FB46FD62FAE0FCB0" box="[1256,1386,764,788]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FB46FD62FAE0FCB0" box="[1256,1386,764,788]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
and strongly tinged with yellow in
<taxonomicName id="EEF98AD69A2AFFD0FDAAFCBEFDFFFC9C" box="[516,629,800,824]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FDAAFCBEFDFFFC9C" box="[516,629,800,824]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF98AD69A2AFFD0FD28FCBEFD54FC9C" box="[646,734,800,824]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FD28FCBEFD54FC9C" box="[646,734,800,824]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FD5EFC81FC1CFC9C" box="[752,918,799,824]" captionStart="FIGURES 181 183" captionStartId="89.[151,264,1733,1756]" captionTargetBox="[204,1317,225,1680]" captionTargetId="figure@89.[188,1398,193,1712]" captionTargetPageId="89" captionText="FIGURES 181 183. Field photographs of extranea-group species. (181) A. anceps, male from Upper Aravaipa Canyon, Arizona State, USA, 1 x 2015; (182) A. anceps, teneral female at Village of Campo de Oro, 4.5 km SW of El Dos, Guanacaste Prov., Costa Rica, 31 iii 2007; (183) pair in copula of A. fissa from E of Cuesta de Piedras, Chiriquí Prov., Panama, 22 viii 2012. Photograph (181) by D. Danforth; (182) by WAH; (183) by N. Smith." httpUri="https://zenodo.org/record/322112/files/figure.png" pageId="24" pageNumber="25">Figs. 181183</figureCitation>
) and
<taxonomicName id="EEF98AD69A2AFFD0FC73FCBEFBEAFC9C" box="[989,1120,800,824]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FC73FCBEFBEAFC9C" box="[989,1120,800,824]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
; and the pale coloration is violaceous blue in
<taxonomicName id="EEF98AD69A2AFFD0FEC4FCDAFE63FCF8" box="[362,489,836,860]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FEC4FCDAFE63FCF8" box="[362,489,836,860]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
and vivid blue in
<taxonomicName id="EEF98AD69A2AFFD0FD1AFCDAFCABFCF8" box="[692,801,836,860]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FD1AFCDAFCABFCF8" box="[692,801,836,860]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF98AD69A2AFFD0FC81FCDAFC0BFCF8" box="[815,897,836,860]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FC81FCDAFC0BFCF8" box="[815,897,836,860]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A2AFFD0FC16FCDAFBBCFCF8" box="[952,1078,836,860]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FC16FCDAFBBCFCF8" box="[952,1078,836,860]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2946F1559A2AFFD0FF69FCF9FB8CFAA9" blockId="24.[151,1437,151,1472]" pageId="24" pageNumber="25">
The large erect oval mesostigmal plates in
<taxonomicName id="EEF98AD69A2AFFD0FD0CFCF6FC90FC24" box="[674,794,872,896]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FD0CFCF6FC90FC24" box="[674,794,872,896]" italics="true" pageId="24" pageNumber="25">A rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FC87FCF9FC0DFC24" box="[809,903,871,896]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="24" pageNumber="25">Fig. 113</figureCitation>
) are almost approximate and are similar only to those of
<taxonomicName id="EEF98AD69A2AFFD0FF54FC12FEC5FC00" box="[250,335,908,932]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FF54FC12FEC5FC00" box="[250,335,908,932]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FEF1FC12FE35FC01" box="[351,447,908,933]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Fig. 116</figureCitation>
) and, to a lesser extent,
<taxonomicName id="EEF98AD69A2AFFD0FD76FC12FCDDFC00" box="[728,855,908,932]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FD76FC12FCDDFC00" box="[728,855,908,932]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FCC6FC12FB81FC00" box="[872,1035,908,933]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Figs. 117, 118</figureCitation>
) and
<taxonomicName id="EEF98AD69A2AFFD0FBE3FC12FB36FC00" box="[1101,1212,908,932]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FBE3FC12FB36FC00" box="[1101,1212,908,932]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FB62FC12FAF9FC01" box="[1228,1395,908,933]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="24" pageNumber="25">Figs. 114, 115</figureCitation>
). A transverse swollen tubercle is present at the base of the mesostigmal lobe in posterior view in
<taxonomicName id="EEF98AD69A2AFFD0FB09FC2EFA99FC6C" box="[1191,1299,944,968]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FB09FC2EFA99FC6C" box="[1191,1299,944,968]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FA8CFC31FA0BFC6C" box="[1314,1409,943,968]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="24" pageNumber="25">Fig. 115</figureCitation>
e); no tubercle is present in
<taxonomicName id="EEF98AD69A2AFFD0FE6EFC4AFDCEFC48" box="[448,580,980,1004]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FE6EFC4AFDCEFC48" box="[448,580,980,1004]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FDF7FC4AFD37FC48" box="[601,701,980,1005]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="24" pageNumber="25">Fig. 113</figureCitation>
e),
<taxonomicName id="EEF98AD69A2AFFD0FD4DFC4AFCB9FC48" box="[739,819,980,1004]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FD4DFC4AFCB9FC48" box="[739,819,980,1004]" italics="true" pageId="24" pageNumber="25">A fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FCE9FC4AFC20FC49" box="[839,938,980,1005]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Fig. 116</figureCitation>
e) or
<taxonomicName id="EEF98AD69A2AFFD0FC43FC4AFBFAFC48" box="[1005,1136,980,1004]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FC43FC4AFBFAFC48" box="[1005,1136,980,1004]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FB2AFC4AFB78FC48" box="[1156,1266,980,1005]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Figs. 117</figureCitation>
e, 118e). In
<taxonomicName id="EEF98AD69A2AFFD0FA2AFC4AFF7DFBB4" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FA2AFC4AFF7DFBB4" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
, the medial margin of the mesostigmal lobe is swollen and slightly recurved (
<figureCitation id="B1C2EDD09A2AFFD0FB38FC69FB70FBB4" box="[1174,1274,1015,1040]" captionStart="FIGURES 113 115" captionStartId="63.[151,264,1857,1880]" captionTargetBox="[157,1433,216,1820]" captionTargetId="figure@63.[151,1436,193,1836]" captionTargetPageId="63" captionText="FIGURES 113 115. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view; (g) lateral view." httpUri="https://zenodo.org/record/322086/files/figure.png" pageId="24" pageNumber="25">Fig. 113</figureCitation>
d, e); no such swollen area is present in
<taxonomicName id="EEF98AD69A2AFFD0FE15FB82FD84FB90" box="[443,526,1052,1076]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="1B8D2D479A2AFFD0FE15FB82FD84FB90" box="[443,526,1052,1076]" italics="true" pageId="24" pageNumber="25">A. fissa</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FDB3FB82FDF6FB91" box="[541,636,1052,1077]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Fig. 116</figureCitation>
e) or
<taxonomicName id="EEF98AD69A2AFFD0FD18FB82FCBFFB90" box="[694,821,1052,1076]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FD18FB82FCBFFB90" box="[694,821,1052,1076]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FCEAFB82FC25FB90" box="[836,943,1052,1077]" captionStart="FIGURES 116 118" captionStartId="64.[151,264,1910,1933]" captionTargetBox="[159,1417,218,1866]" captionTargetId="figure@64.[151,1436,193,1890]" captionTargetPageId="64" captionText="FIGURES 116 118. Female mesostigmal plates of extranea - group species. (a) frontal (anterior) view; (b) detail of left mesostigmal lobe, frontal view; (c) dorsal view; (d) detail of left mesostigmal lobe, dorsal view; (e) detail of right mesostigmal lobe, posterior view." httpUri="https://zenodo.org/record/322087/files/figure.png" pageId="24" pageNumber="25">Figs. 117</figureCitation>
e, 118e). In the field, females of
<taxonomicName id="EEF98AD69A2AFFD0FAB2FB82FA16FB90" box="[1308,1436,1052,1076]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FAB2FB82FA16FB90" box="[1308,1436,1052,1076]" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A2AFFD0FF31FBA1FF7BFBFC" box="[159,241,1087,1112]" captionStart="FIGURES 43 - 46" captionStartId="50.[151,264,1915,1938]" captionTargetBox="[172,1436,193,1880]" captionTargetId="figure@50.[151,1436,193,1894]" captionTargetPageId="50" captionText="FIGURES 43 - 46. Color pattern of female head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322073/files/figure.png" pageId="24" pageNumber="25">Fig. 43</figureCitation>
) are easily distinguished from those of the other three species (
<figureCitation id="B1C2EDD09A2AFFD0FC17FBA1FBC8FBFC" box="[953,1090,1087,1112]" captionStart="FIGURES 43 - 46" captionStartId="50.[151,264,1915,1938]" captionTargetBox="[172,1436,193,1880]" captionTargetId="figure@50.[151,1436,193,1894]" captionTargetPageId="50" captionText="FIGURES 43 - 46. Color pattern of female head, thorax, and S 1 4 of extranea - group species. Head, dorsal view, thorax and S 1 4, lateral view." httpUri="https://zenodo.org/record/322073/files/figure.png" pageId="24" pageNumber="25">Figs. 4446</figureCitation>
; 181183) by the overall more extensive dark markings on the head (mostly black epicranium), thorax (prominent middorsal and humeral stripes), and abdomen (postbasal streaks coupled with distal and ventral black); the other three species are almost immaculate, there being only vestiges of black on the epicranium, a thin middorsal thoracic stripe, a narrow to absent humeral stripe, and vestiges of a distal black spot laterally on abdominal segments. The ovipositor in
<taxonomicName id="EEF98AD69A2AFFD0FA2AFB4EFF7DFAA8" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rudolphi">
<emphasis id="1B8D2D479A2AFFD0FA2AFB4EFF7DFAA8" italics="true" pageId="24" pageNumber="25">A. rudolphi</emphasis>
</taxonomicName>
is black (
<figureCitation id="B1C2EDD09A2AFFD0FECAFB6AFE3DFAA8" box="[356,439,1268,1293]" captionStart="FIGURES 89 95" captionStartId="57.[151,264,1887,1910]" captionTargetBox="[164,1421,232,1839]" captionTargetId="figure@57.[151,1436,193,1866]" captionTargetPageId="57" captionText="FIGURES 89 95. Color pattern of female S 7 10 of extranea - group species, lateral view." httpUri="https://zenodo.org/record/322080/files/figure.png" pageId="24" pageNumber="25">Fig. 92</figureCitation>
), not pale as in the other three species (
<figureCitation id="B1C2EDD09A2AFFD0FCDEFB6AFC73FAA9" box="[880,1017,1268,1293]" captionStart="FIGURES 89 95" captionStartId="57.[151,264,1887,1910]" captionTargetBox="[164,1421,232,1839]" captionTargetId="figure@57.[151,1436,193,1866]" captionTargetPageId="57" captionText="FIGURES 89 95. Color pattern of female S 7 10 of extranea - group species, lateral view." httpUri="https://zenodo.org/record/322080/files/figure.png" pageId="24" pageNumber="25">Figs. 9395</figureCitation>
).
</paragraph>
<paragraph id="2946F1559A2AFFD0FF69FA89FB90FADC" blockId="24.[151,1437,151,1472]" pageId="24" pageNumber="25">
<emphasis id="1B8D2D479A2AFFD0FF69FA89FEA9FA94" bold="true" box="[199,291,1303,1328]" pageId="24" pageNumber="25">Habitat</emphasis>
. Streams and drainage areas. Specimens were taken at elevations ranging from about
<quantity id="EE015CB09A2AFFD0FB4FFA89FAA2FA8B" box="[1249,1320,1303,1328]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="24" pageNumber="25" unit="m" value="570.0">570 m</quantity>
(Sierra de Huauchinango) to
<quantity id="EE015CB09A2AFFD0FEC5FAA2FE43FAF0" box="[363,457,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="25" unit="m" value="1300.0">1,300 m</quantity>
(stream with cascade
<quantity id="EE015CB09A2AFFD0FD64FAA3FC8FFAF1" box="[714,773,1340,1365]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="km" value="7.0">7 km</quantity>
N of Tlanchinol). Specimens have been collected in May (Sierra de Huauchinango) and July (stream with cascade
<quantity id="EE015CB09A2AFFD0FCB8FAFEFCC5FADC" box="[790,847,1375,1400]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="km" value="7.0">7 km</quantity>
N of Tlanchinol).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A2AFFD0FF69FA1AFBE9FA1B" pageId="24" pageNumber="25" type="distribution">
<paragraph id="2946F1559A2AFFD0FF69FA1AFBE9FA1B" blockId="24.[151,1437,151,1472]" pageId="24" pageNumber="25">
<emphasis id="1B8D2D479A2AFFD0FF69FA1AFED3FA39" bold="true" box="[199,345,1412,1437]" pageId="24" pageNumber="25">Distribution</emphasis>
. Limited to a small area in the Atlantic slope of central
<collectingCountry id="51EEB1C59A2AFFD0FC56FA1AFBC7FA39" box="[1016,1101,1412,1437]" name="Mexico" pageId="24" pageNumber="25">
<collectingRegion id="EB3D3FB79A2AFFD0FC56FA1AFBC7FA39" box="[1016,1101,1412,1437]" country="Mexico" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingRegion>
</collectingCountry>
within
<collectingRegion id="EB3D3FB79A2AFFD0FB04FA1AFB73FA39" box="[1194,1273,1412,1437]" country="Mexico" name="Puebla" pageId="24" pageNumber="25">Puebla</collectingRegion>
,
<collectingRegion id="EB3D3FB79A2AFFD0FAA6FA1AFAEFFA39" box="[1288,1381,1412,1437]" country="Mexico" name="Hidalgo" pageId="24" pageNumber="25">Hidalgo</collectingRegion>
, and
<collectingRegion id="EB3D3FB79A2AFFD0FF39FA36FF71FA1B" box="[151,251,1448,1471]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="24" pageNumber="25">Veracruz</collectingRegion>
states (
<figureCitation id="B1C2EDD09A2AFFD0FEE1FA39FE3AFA64" box="[335,432,1447,1472]" captionStart="FIGURES 167 168" captionStartId="83.[151,264,1723,1746]" captionTargetBox="[209,1344,219,1653]" captionTargetId="figure@83.[188,1398,193,1702]" captionTargetPageId="83" captionText="FIGURES 167 168. Distribution of extranea-group species." httpUri="https://zenodo.org/record/322106/files/figure.png" pageId="24" pageNumber="25">Fig. 168</figureCitation>
), where it occurs in sympatry with
<taxonomicName id="EEF98AD69A2AFFD0FC95FA36FC22FA64" box="[827,936,1448,1472]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="1B8D2D479A2AFFD0FC95FA36FC22FA64" box="[827,936,1448,1472]" italics="true" pageId="24" pageNumber="25">A. anceps</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A2AFFD0FC70FA36FBD7FA64" box="[990,1117,1448,1472]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="1B8D2D479A2AFFD0FC70FA36FBD7FA64" box="[990,1117,1448,1472]" italics="true" pageId="24" pageNumber="25">A. westfalli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>