175 lines
17 KiB
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175 lines
17 KiB
XML
<document ID-CLB-Dataset="21607" ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="7142DF34C24055CDB980925206686C0D" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis stolonifera L., Sp. Pl. 1: 62. 1753" docType="treatment" docVersion="6" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1732688126400" updateUser="ExternalLinkService">
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<mods:titleInfo id="7A2B80FFD18062ACC711792688A55BE1">
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<mods:title id="44C7C28FF3E2F4BAA3F9CCD8CBB48BBB">Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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<mods:name id="E1F814470B51E8D060824EE724EBDA65" type="personal">
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<mods:role id="1BF33F06EB269841659672D5BD913F0E">
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<mods:namePart id="433B3A4C6C1D975CF4D2020A86EE0319">Sylvester, Steven P.</mods:namePart>
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<mods:affiliation id="682C81967FF14FBAABDF7D1948BCE214">College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier id="D59DC44E24558E15AF15A223323C0068" type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier id="42752128724BE645E35D177010084606" type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="37CC8C6E19ED1DF9626176F9A312BE5B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FF908D0DDF2FB6EBF149FAEB810C8B15">Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation id="462E25CB18964BA96D0AF7C16E3DF2B2">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name id="3AFF02E4794076391068D9AB69934DE2" type="personal">
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<mods:role id="C4857A362AB82CDAF8C0CD66B1F0BBEB">
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<mods:roleTerm id="BED5ACC82F6754CBE2C5D40EE25D6A01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C76A887A5AFF148F4C0991F9954883DF">Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation id="27809B9AB5B2FBC32435F35DA61201C2">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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<mods:roleTerm id="4470B29141A51ABEA1210A3D4D8CC72B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2B6D852030BDE71E90B2ECFD5565A86A">Soreng, Robert J.</mods:namePart>
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<mods:affiliation id="AA0DBF344BEDCB0C2DDC78928A48F49B">Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier id="50922A32E3A7722CAE0DF266DDDEB0F7" type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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<mods:typeOfResource id="41A6185095D40BC83E0DBCA6DA219B19">text</mods:typeOfResource>
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<mods:title id="0BC9644F0AC255E9C90D3EBDF11771E2">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="9DDA8B21E822F77239D3C3DE684F46A2">
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<mods:date id="E1D07BBD6FC74B3173E54E347E5EB684">2020</mods:date>
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<mods:detail id="870FEC638AA1C94626C8D6DBF43E0D01" type="volume">
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<mods:number id="839E7301AF6B53D64E3B1124A8034732">151</mods:number>
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</mods:detail>
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<mods:start id="566765F46979F2A6E8C3620900C5C7D2">107</mods:start>
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<mods:end id="66C77F9D12311E985CEB2D45BB46A711">160</mods:end>
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</mods:extent>
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<mods:location id="C3BAD33465417F38FEC88953D9F1A7EE">
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<mods:url id="80FA34ADE9B73FBFDCA520318D03867D">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification id="7EA0C620683D1CE7D5D9636D7FDBD35F">journal article</mods:classification>
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<mods:identifier id="42E93918AAA909557D7E39348D261DFF" type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier id="9DDAB25139E4A8F22FD54C7EC2CB285B" type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier id="80A90B5555CBF8FA29E7C0E3DFC8D43A" type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment id="7142DF34C24055CDB980925206686C0D" ID-GBIF-Taxon="164730851" LSID="urn:lsid:plazi:treatment:7142DF34C24055CDB980925206686C0D" httpUri="http://treatment.plazi.org/id/7142DF34C24055CDB980925206686C0D" lastPageNumber="107" pageId="0" pageNumber="107" scope_family="Poaceae">
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<subSubSection id="1133CC5FF98B225C5C7545055254E69B" pageId="0" pageNumber="107" type="nomenclature">
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<paragraph id="F10EC73445E7A044B76D25EDC1B56437" pageId="0" pageNumber="107">
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<taxonomicName id="D37B92B09BD9C37EEE793D7BE0C5A2D4" ID-CoL="665HP" LSID="7142DF34-C240-55CD-B980-925206686C0D" authority="L., Sp. Pl. 1: 62. 1753" authorityName="L., Sp. Pl. 1: 62." authorityYear="1753" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis stolonifera" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="stolonifera">Agrostis stolonifera L., Sp. Pl. 1: 62. 1753</taxonomicName>
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<figureCitation id="14743E01F66EED68ED43A6F6AE2BDA45" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Agrostis stolonifera A whole plant B inflorescence, close-up C spikelet, close-up, lateral view. Image A Cuatrecasas 452 (US 1772826) B Garcia-Barriga 6 (US 2115003)." figureDoi="10.3897/phytokeys.151.50538.figure9" httpUri="https://binary.pensoft.net/fig/424847" pageId="0" pageNumber="107">Fig. 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0E23201474FFC39652C431925334203D" pageId="0" pageNumber="107" type="materials_examined">
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<paragraph id="CB36B430F4990AC5C4F764625053A14A" pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph id="E58037F378AF3DEECD2D32C9C831BF0A" pageId="0" pageNumber="107">
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[Habitat in Europa], Herb. A. van Royen s.n. (lectotype, designated by
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<bibRefCitation id="8A0E7A9F09C066C8229CA4AFACAFA577" author="Widen, K-G" journalOrPublisher="Flora Fennica" pageId="0" pageNumber="107" pagination="1 - 209" refId="B41" refString="Widen, K-G, 1971. The Genus Agrostis in Eastern Fennoscandia. Taxonomy and Distribution. Flora Fennica 5: 1 - 209" title="The Genus Agrostis in Eastern Fennoscandia. Taxonomy and Distribution." volume="5" year="1971">
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<normalizedToken id="F7411E9AEA1BD8309D5ACAB35D8C12DB" originalValue="Widén">Widen</normalizedToken>
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1971
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</bibRefCitation>
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: 77: L (L0059234 [image!])).
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</paragraph>
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<paragraph id="DFF7B6A3DE1185BBB2CC94FF2C451F64" pageId="0" pageNumber="107">Many heterotypic synonyms.</paragraph>
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</subSubSection>
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<subSubSection id="0DB3CF9B7B22F1EE17671068BC78EC1A" pageId="0" pageNumber="107" type="description">
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<paragraph id="E7C13C3473FE1FE51F55686E45843244" pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph id="1BA7A69D0AD4D26D1D63C61679E15575" pageId="0" pageNumber="107">
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<emphasis id="3AB83972A671B6C402DC66B7724E04E6" bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
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, generally creeping, usually extensively stoloniferous with long stolons to 200 cm long, less often with short stolons, rhizomes usually absent.
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<emphasis id="6011A245BE7CD244E8CCB4BB3D6BD34F" bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
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extravaginal, with cataphylls present.
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<emphasis id="4DB90E3A9B9DF50C8ED81D58302308E4" bold="true" pageId="0" pageNumber="107">Culms</emphasis>
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15-100 cm tall, erect or decumbent at their base, delicate to fairly firm, nodes usually held within sheaths with 0(-3) exerted at flowering, usually smooth to rarely scaberulous.
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<emphasis id="C4E6C84F0AEF9A4F82EDDB8C6A6BA9E0" bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
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mainly cauline, glabrous, scaberulous;
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<emphasis id="8DC6AC4AEE593C58191EEFE43211DD2B" bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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1-6.5(-8) mm long, of basal leaves and tillers 1-3 mm long, of upper culm 2-6.5(-8) mm long, rounded to truncate, not or slightly to moderately decurrent with the sheath, abaxial surface scaberulous;
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<emphasis id="1B50547365B82774178123DC7BAAF806" bold="true" pageId="0" pageNumber="107">blades</emphasis>
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2-26 cm long, 1-8 mm wide, flat or sometimes folded, soft and lax, surfaces and margins scaberulous to scabrous, apices acute.
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<emphasis id="F7F7C37A26F7443AFDE446E037CC74BD" bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
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2-20(-32) cm long, 1-16 cm wide, open and lax or contracted after flowering, ovoid to pyramidal, slightly to usually greatly exerted from the basal foliage, lateral branches naked in the lower 1/3-1/4 or with spikelets present to the base, long, ascending, spreading, to somewhat divergent and not held close to the central inflorescence axis at or before flowering, held close to the central rachis at maturity, central axis and panicle branches scabrous or very rarely smooth;
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<emphasis id="30594F437D7A26EA1D99CA36C13003D8" bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
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0.5-2.5 mm long, usually shorter than their spikelets, dilated or not at their apex, scabrous.
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<emphasis id="C125BC74537BEEEB49CA805F1667B849" bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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(not including awn, if present) 1.8-3 mm long;
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<emphasis id="A5E1CDF89A3E268BC5038BB4602F0887" bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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subequal or unequal, the lower slightly longer than the upper by up to c. 0.5 mm, 1-veined, lower glume keel usually scabrous in the distal half, upper glume often smooth throughout, apices acute;
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<emphasis id="865A06DB603AE08D34E7774FA3B1FF9C" bold="true" pageId="0" pageNumber="107">floret</emphasis>
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usually 2/3-3/4 the length of the glumes;
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<emphasis id="28CB002FE6A1C63CD13D0C8366128295" bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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with 2 sparse lateral tufts of short hairs;
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<emphasis id="BF107FE211DCA5F7B7D4B6777539FFDD" bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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1.2-2.2(-2.5) mm long, (3-) 5-veined, glabrous or rarely pubescent at the base with hairs 0.1-0.2 mm long, smooth, apex obtuse or truncate, erose, muticous or rarely with an awn 0.5-3 mm long, usually inserted above the middle of the dorsal keel, exerted or not from the glumes, straight, flexuose or geniculate, twisted or not, weak and falling easily;
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<emphasis id="DD7C70A5A2EA57714ADF2C9EFFB13761" bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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0.7-1.3(-1.6) mm long, (2/5-)
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<normalizedToken id="CB9DB6EFDCC88DB0C4EE4EEE2557EC54" originalValue="½-¾">1/2-3/4</normalizedToken>
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the length of the lemma;
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<emphasis id="6A21F661C1BE7826C3C5AC2DA3F50BA3" bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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absent;
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<emphasis id="1B941DD882FC97B8B0983FBC25AC7503" bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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0.9-1.6 mm long.
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</paragraph>
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<caption id="D1F04FADB833736590A35D724EE44781" doi="10.3897/phytokeys.151.50538.figure9" httpUri="https://binary.pensoft.net/fig/424847" pageId="0" pageNumber="107" start="Figure 9" startId="F9">
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<paragraph id="2E8F11563DBCE932E69D73BD058AC1E2" pageId="0" pageNumber="107">
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<emphasis id="68FF1DD96C6107E3640B09878C3888A1" bold="true" pageId="0" pageNumber="107">Figure 9.</emphasis>
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<taxonomicName id="DF05898F9673C6CF0CA9FECEBE727090" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis stolonifera" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="stolonifera">
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<emphasis id="2BEE4EFB6827FD47B3C63E89FA1DA056" italics="true" pageId="0" pageNumber="107">Agrostis stolonifera</emphasis>
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</taxonomicName>
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<emphasis id="5743C13B9E70113C8FB4487FDCC3DBAB" bold="true" pageId="0" pageNumber="107">A</emphasis>
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whole plant
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<emphasis id="64C6D57372574EB1A74FD21B5F269734" bold="true" pageId="0" pageNumber="107">B</emphasis>
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inflorescence, close-up
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<emphasis id="C2FD3E142A3B61FE6AE26DD8E7BB44BC" bold="true" pageId="0" pageNumber="107">C</emphasis>
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spikelet, close-up, lateral view. Image
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<emphasis id="9E7B13AF967D864763766D43D28D0C3C" bold="true" pageId="0" pageNumber="107">A</emphasis>
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Cuatrecasas 452 (US1772826)
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<emphasis id="2A830907B36DA1532E8BB6A5A103809F" bold="true" pageId="0" pageNumber="107">B</emphasis>
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<normalizedToken id="0380ADEACD5D437D5C52A15B1EE4B6F6" originalValue="García-Barriga">Garcia-Barriga</normalizedToken>
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6 (US2115003).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="66A3E938439932ACCF6897B71FF70408" pageId="0" pageNumber="107" type="distribution">
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<paragraph id="D908B38E43133618E810C5DFCF161C77" pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
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<paragraph id="D06284819FD48FE287BD07FA55C1FD04" pageId="0" pageNumber="107">
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Widespread and cosmopolitan or Eurasian origin, introduced to South America from Europe. While no specimens of
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<taxonomicName id="1C82D08D362F0BB8CF07D21B0C54E70E" lsidName="A. stolonifera" pageId="0" pageNumber="107" rank="species" species="stolonifera">
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<emphasis id="88713E8645A62EA84928BD03580430E8" italics="true" pageId="0" pageNumber="107">A. stolonifera</emphasis>
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</taxonomicName>
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were verified from Departamento
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<normalizedToken id="83250DDF33F6322C4AB134996036A548" originalValue="Boyacá">Boyaca</normalizedToken>
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at US or encountered during extensive fieldwork in the region, the species is cited for
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<normalizedToken id="78C18F3C85F0F77C778EED63FBBE7943" originalValue="Boyacá">Boyaca</normalizedToken>
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in the checklist (
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<bibRefCitation id="00FBD783093772B2A18DB929BD7970F1" author="Giraldo-Canas, D" editor="Bernal, R" journalOrPublisher="2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D. C." pageId="0" pageNumber="107" pagination="2127 - 2205" refId="B10" refString="Giraldo-Canas, D, Londono, X, Clark, LG, 2016. Poaceae. In: Bernal, R, Gradstein, SR, Celis, M, Eds., Catalogo de plantas y liquenes de Colombia (Vol. 2). Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D.C.: 2127 - 2205" title="Poaceae." volumeTitle="Catalogo de plantas y liquenes de Colombia (Vol." year="2016">
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<normalizedToken id="AD4401FA9617344F2CA26A9494C23BD7" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
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et al. 2016
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</bibRefCitation>
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) and specimens were found at US from the neighboring Departamento Cundinamarca in the Cordillera Oriental meaning it likely that the species occurs in
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<normalizedToken id="A5F767D7187FA46EA3176CE540552F67" originalValue="Boyacá">Boyaca</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7EEC1BC766ACC42F7825B1F785CA65EC" pageId="0" pageNumber="107" type="materials_examined">
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<paragraph id="E9D1260CD26DB02388CFEC7375421A65" pageId="0" pageNumber="107">Other specimens examined.</paragraph>
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<paragraph id="4DA0756158B249FD733C4A9EBC87DB12" pageId="0" pageNumber="107">
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<emphasis id="3989BE51C56AE93C5E528DBC2D5C9772" bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
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.
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<emphasis id="A7A22A08322AA709BC5998F6700E24C5" bold="true" pageId="0" pageNumber="107">Cundinamarca</emphasis>
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: Sabana de Bogota, 2600 m alt., 29 Dec. 1938, J. Cuatrecasas 452 (US1772826); Tunjuelo, granja experimental "La Picota", [
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<geoCoordinate id="895E856732F9ACBCC73DF292250228AA" degrees="4.5331" direction="north" orientation="latitude" precision="5" value="4.5331">4.5331N</geoCoordinate>
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,
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<geoCoordinate id="0F702F24576DA6DE8FD65B0FCD4E5685" degrees="74.1117" direction="west" orientation="longitude" precision="5" value="-74.1117">74.1117W</geoCoordinate>
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], 2600 m alt., 3 Feb. 1933, H.
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<normalizedToken id="6D37C6DAADB15D3896D1FC7C4AF5B309" originalValue="García-Barriga">Garcia-Barriga</normalizedToken>
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6 (US2115003).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |