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<document id="055CC603AE0AE49C9FCD8006970712DC" ID-DOI="10.12782/specdiv.26.249" ID-ISSN="2189-7301" ID-Zenodo-Dep="5739009" ID-ZooBank="2D6B646A-63C4-4BEE-A655-DBEB827DFC10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638231029206" checkinUser="felipe" docAuthor="Uyeno, Daisuke &amp; Rain, Enjoy" docDate="2021" docId="270887B0FFD4465FB5DEF956FC73F949" docLanguage="en" docName="SpeciesDiversity.26.2.249-254.pdf" docOrigin="Species Diversity 26" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Pseudanthessius chuukensis Uyeno &amp; Rain 2021, n. sp." docType="treatment" docUuid="BCFB5B36-76CB-4F98-9088-6AEDFE67ACE7" docUuidSource="ZooBank" docVersion="4" lastPageNumber="250" masterDocId="DB31FFC8FFD4465EB64CFFB6FFA8FFDF" masterDocTitle="A New Species of Pseudanthessius (Copepoda: Cyclopoida: Pseudanthessiidae) on Epinephelus sp. (Actinopterygii: Perciformes: Serranidae) from Chuuk Lagoon, Federated States of Micronesia" masterLastPageNumber="254" masterPageNumber="249" pageNumber="249" updateTime="1699434522783" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="6A7D99DD5D621E528B70C64C4321D46F">
<mods:title id="BF30F9247E77626513371E67DF253E78">A New Species of Pseudanthessius (Copepoda: Cyclopoida: Pseudanthessiidae) on Epinephelus sp. (Actinopterygii: Perciformes: Serranidae) from Chuuk Lagoon, Federated States of Micronesia</mods:title>
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<mods:name id="EB71C4A9B443256EC3013BDA2E7DEEED" type="personal">
<mods:role id="90AB755A67F6446C128627C6505C457E">
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<mods:namePart id="6731EFF8C77D053F19E5C06DE025A93A">Uyeno, Daisuke</mods:namePart>
<mods:affiliation id="7CEB9FC2280DB12CEDDC54E28DADFEDB">Graduate School of Science and Engineering, Kagoshima University, 1 - 21 - 35 Korimoto, Kagoshima 890 - 0065, Japan E-mail: duyeno @ sci. kagoshima-u. ac. jp &amp; Corresponding author</mods:affiliation>
<mods:nameIdentifier id="84400BD6A2EF5E666065AB29C3352741" type="email">duyeno@sci.kagoshima-u.ac.jp</mods:nameIdentifier>
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<mods:roleTerm id="A261899F6027851AE7F2CA16B10C17B9">Author</mods:roleTerm>
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<mods:namePart id="9B7DB882AB55A8F88AFC4FE807CCE06B">Rain, Enjoy</mods:namePart>
<mods:affiliation id="34350921741A064037A99369374302AE">The Chuuk Department of Marine Resources, P. O. Box 207, Weno, Chuuk State 96942, Federated States of Micronesia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BB40B3484B54159978AA96C68ACBE2DB">text</mods:typeOfResource>
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<mods:title id="3E461E77FBFA7D0A956106EAFA02879A">Species Diversity</mods:title>
</mods:titleInfo>
<mods:part id="89129AEFF06D3FE68BE9EABACA45C212">
<mods:date id="98BA5B95D4AF1EA9234C9066D272A1DB">2021</mods:date>
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<mods:number id="33FA5295083BE4F13A693F1F42E360B2">2021-09-10</mods:number>
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<mods:number id="972E15BE1ABC675DAC84CEA1EABE14F8">26</mods:number>
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<mods:identifier id="5BC828BB1D817EE3522D7DB73F4BFA8C" type="ISSN">2189-7301</mods:identifier>
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<treatment id="270887B0FFD4465FB5DEF956FC73F949" LSID="urn:lsid:zoobank.org:act:BCFB5B36-76CB-4F98-9088-6AEDFE67ACE7" httpUri="http://treatment.plazi.org/id/270887B0FFD4465FB5DEF956FC73F949" lastPageId="1" lastPageNumber="250" pageId="0" pageNumber="249">
<subSubSection id="E7BB652DFFD4465EB5DEF956FB31F8C6" pageId="0" pageNumber="249" type="nomenclature">
<paragraph id="AF1E36A6FFD4465EB5DEF956FB31F8C6" blockId="0.[914,1338,1760,1817]" pageId="0" pageNumber="249">
<heading id="F45681CAFFD4465EB5DEF956FB31F8C6" centered="true" fontSize="10" level="2" pageId="0" pageNumber="249" reason="2">
<taxonomicName id="68A14D25FFD4465EB5DEF956FB53F927" authority="Uyeno" authorityName="Uyeno &amp; Rain" authorityYear="2021" box="[914,1275,1760,1784]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="0" pageNumber="249" phylum="Arthropoda" rank="species" species="chuukensis" status="sp. nov.">
<emphasis id="9DD5EAB4FFD4465EB5DEF956FB19F927" bold="true" box="[914,1201,1760,1784]" italics="true" pageId="0" pageNumber="249">Pseudanthessius chuukensis</emphasis>
Uyeno
</taxonomicName>
,
<taxonomicNameLabel id="86E657CFFFD4465EB349F956FA92F928" box="[1285,1338,1760,1783]" pageId="0" pageNumber="249" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="379A2A23FFD4465EB276F8B6FB3AF8C6" box="[1082,1170,1792,1817]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[157,188,1711,1733]" captionStartId-1="3.[156,187,1756,1778]" captionStartId-2="4.[156,187,1551,1573]" captionTargetBox-0="[190,1397,206,1669]" captionTargetBox-1="[189,1397,206,1716]" captionTargetBox-2="[189,1397,207,1511]" captionTargetId-0="figure-160@2.[187,1399,205,1673]" captionTargetId-1="figure-83@3.[187,1399,205,1718]" captionTargetId-2="figure-81@4.[187,1399,205,1513]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." captionText-1="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." captionText-2="Fig. 4. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right leg 2, anterior; B, right leg 3, anterior; C, left leg 4, anterior; D, right leg 5, dorsal. Scale bars: AD, 50µm." figureDoi-0="http://doi.org/10.5281/zenodo.5739013" figureDoi-1="http://doi.org/10.5281/zenodo.5739017" figureDoi-2="http://doi.org/10.5281/zenodo.5739021" httpUri-0="https://zenodo.org/record/5739013/files/figure.png" httpUri-1="https://zenodo.org/record/5739017/files/figure.png" httpUri-2="https://zenodo.org/record/5739021/files/figure.png" pageId="0" pageNumber="249">Figs 24</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E7BB652DFFD4465FB502F8F4FD9CF864" lastPageId="1" lastPageNumber="250" pageId="0" pageNumber="249" type="materials_examined">
<paragraph id="AF1E36A6FFD4465EB502F8F4FB94F803" blockId="0.[821,1433,1857,2077]" pageId="0" pageNumber="249">
<emphasis id="9DD5EAB4FFD4465EB502F8F4FC4EF886" bold="true" box="[846,998,1857,1882]" pageId="0" pageNumber="249">Type material.</emphasis>
<typeStatus id="701A8804FFD4465EB5A1F8F4FBE6F885" box="[1005,1102,1858,1882]" pageId="0" pageNumber="249" type="holotype">Holotype</typeStatus>
:
<specimenCount id="B9A7FD2FFFD4465EB21BF8F4FB35F885" box="[1111,1181,1858,1882]" pageId="0" pageNumber="249" type="adult">1 adult</specimenCount>
female (
<collectionCode id="C9B0AE63FFD4465EB2BFF8F7FAF8F886" box="[1267,1360,1857,1881]" pageId="0" pageNumber="249">BPBM-S</collectionCode>
18653), ex
<taxonomicName id="68A14D25FFD4465EB51FF8D4FC45F8A5" box="[851,1005,1890,1914]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="249" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="9DD5EAB4FFD4465EB51FF8D4FC61F8A5" box="[851,969,1890,1914]" italics="true" pageId="0" pageNumber="249">Epinephelus</emphasis>
sp.
</taxonomicName>
[probably
<taxonomicName id="68A14D25FFD4465EB212F8D5FA39F8A5" authority="(Bleeker, 1855)" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[1118,1425,1890,1914]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="249" phylum="Chordata" rank="species" species="macrospilos">
<emphasis id="9DD5EAB4FFD4465EB212F8D5FB46F8A5" box="[1118,1262,1890,1914]" italics="true" pageId="0" pageNumber="249">E. macrospilos</emphasis>
(Bleeker, 1855)
</taxonomicName>
] (
<taxonomicName id="68A14D25FFD4465EB577F835FC11F844" box="[827,953,1923,1947]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="249" phylum="Chordata" rank="order">Perciformes</taxonomicName>
:
<taxonomicName id="68A14D25FFD4465EB588F835FB9BF844" authorityName="Swainson" authorityYear="1839" box="[964,1075,1923,1947]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="249" phylum="Chordata" rank="family">Serranidae</taxonomicName>
) (
<figureCitation id="379A2A23FFD4465EB204F835FB2AF844" box="[1096,1154,1923,1947]" captionStart="Fig" captionStartId="1.[156,187,691,713]" captionTargetBox="[189,1398,207,650]" captionTargetId="figure-776@1.[187,1400,205,652]" captionTargetPageId="1" captionText="Fig. 1. A grouper, Epinephelus sp. [probably E. macrospilos (Bleeker, 1855)], from which the holotype of Pseudanthessius chuukensis Uyeno, n. sp. was found in Chuuk Lagoon, FSM. Scale bar: 50mm." figureDoi="http://doi.org/10.5281/zenodo.5739011" httpUri="https://zenodo.org/record/5739011/files/figure.png" pageId="0" pageNumber="249">Fig. 1</figureCitation>
), off western coast of Piis- Paneu Island (
<geoCoordinate id="CA955061FFD4465EB588F815FBB8F864" box="[964,1040,1955,1979]" degrees="7" direction="north" minutes="40" orientation="latitude" pageId="0" pageNumber="249" precision="925" value="7.6666665">7°40N</geoCoordinate>
,
<geoCoordinate id="CA955061FFD4465EB256F815FBD1F864" box="[1050,1145,1955,1979]" degrees="151" direction="east" minutes="45" orientation="longitude" pageId="0" pageNumber="249" precision="925" value="151.75">151°45E</geoCoordinate>
),
<collectingRegion id="6D65F844FFD4465EB2C0F815FB7BF864" box="[1164,1235,1955,1979]" country="Micronesia" name="Chuuk" pageId="0" pageNumber="249">Chuuk</collectingRegion>
,
<collectingCountry id="D7B67636FFD4465EB291F815FAA7F864" box="[1245,1295,1955,1979]" name="Micronesia" pageId="0" pageNumber="249">FSM</collectingCountry>
, North Pacific Ocean,
<date id="DB1F1066FFD4465EB5D5F872FB90F803" box="[921,1080,1988,2012]" pageId="0" pageNumber="249" value="2016-08-23">23 August 2016</date>
.
</paragraph>
<paragraph id="AF1E36A6FFD4465FB502F852FDC2FC15" blockId="0.[821,1433,1857,2077]" lastBlockId="1.[156,768,784,2077]" lastPageId="1" lastPageNumber="250" pageId="0" pageNumber="249">
<emphasis id="9DD5EAB4FFD4465EB502F852FC77F823" bold="true" box="[846,991,2020,2044]" pageId="0" pageNumber="249">Adult female.</emphasis>
Body (
<figureCitation id="379A2A23FFD4465EB266F852FBDEF823" box="[1066,1142,2020,2044]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="0" pageNumber="249">Fig. 2A</figureCitation>
) cyclopiform, 1743 long, depressed dorsoventrally with greatest width at cephalosome. Cephalosome, wider than long, 278×403. First to fourth pedigerous somites and urosomites free. Prosome 703 long. Genital double somite (
<figureCitation id="379A2A23FFD5465FB71DFCE7FE1DFCB6" box="[337,437,849,873]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="1" pageNumber="250">Fig. 2A, B</figureCitation>
) longer than wide, 215×176. Abdomen composed of three free somites, 178×162, 164×150, and 131×140, respectively. Caudal ramus (
<figureCitation id="379A2A23FFD5465FB41AFC24FD6DFC75" box="[598,709,914,938]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="1" pageNumber="250">Fig. 2A, C</figureCitation>
) 5.69 times longer than wide, 287×51, with six setae.
</paragraph>
<caption id="FBDE662EFFD5465FB6D0FD05FD19FD3B" ID-DOI="http://doi.org/10.5281/zenodo.5739011" ID-Zenodo-Dep="5739011" httpUri="https://zenodo.org/record/5739011/files/figure.png" pageId="1" pageNumber="250" startId="1.[156,187,691,713]" targetBox="[189,1398,207,650]" targetPageId="1">
<paragraph id="AF1E36A6FFD5465FB6D0FD05FD19FD3B" blockId="1.[156,1432,690,741]" pageId="1" pageNumber="250">
Fig. 1. A grouper,
<taxonomicName id="68A14D25FFD5465FB71AFD05FE77FD17" box="[342,479,691,712]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="9DD5EAB4FFD5465FB71AFD05FE68FD17" box="[342,448,691,712]" italics="true" pageId="1" pageNumber="250">Epinephelus</emphasis>
sp.
</taxonomicName>
[probably
<taxonomicName id="68A14D25FFD5465FB40DFD05FCF9FD16" authority="(Bleeker, 1855)" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[577,849,690,713]" class="Actinopterygii" family="Serranidae" genus="Epinephelus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="250" phylum="Chordata" rank="species" species="macrospilos">
<emphasis id="9DD5EAB4FFD5465FB40DFD05FD69FD17" box="[577,705,691,712]" italics="true" pageId="1" pageNumber="250">E. macrospilos</emphasis>
(Bleeker, 1855)
</taxonomicName>
], from which the holotype of
<taxonomicName id="68A14D25FFD5465FB212FD05FA3CFD17" authority="Uyeno" authorityName="Uyeno &amp; Rain" authorityYear="2021" box="[1118,1428,691,712]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="chuukensis" status="sp. nov.">
<emphasis id="9DD5EAB4FFD5465FB212FD05FAF9FD17" box="[1118,1361,691,712]" italics="true" pageId="1" pageNumber="250">Pseudanthessius chuukensis</emphasis>
Uyeno
</taxonomicName>
,
<taxonomicNameLabel id="86E657CFFFD5465FB6D0FD79FF64FD3B" box="[156,204,719,740]" pageId="1" pageNumber="250" rank="species">n. sp.</taxonomicNameLabel>
was found in Chuuk Lagoon, FSM. Scale bar: 50mm.
</paragraph>
</caption>
<paragraph id="AF1E36A6FFD5465FB6F9FC62FDA0F844" blockId="1.[156,768,784,2077]" pageId="1" pageNumber="250">
Rostrum (
<figureCitation id="379A2A23FFD5465FB752FC65FEC7FC34" box="[286,367,979,1003]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="1" pageNumber="250">Fig. 2D</figureCitation>
) triangular, bearing distinct apex with blunt margin. Antennule (
<figureCitation id="379A2A23FFD5465FB7FEFC42FDA9FBD3" box="[434,513,1012,1036]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="1" pageNumber="250">Fig. 2E</figureCitation>
) 7-segmented; armature formula 5, 14, 4, 3, 4+1 aesthetasc, 2+1 aesthetasc, 7+1 aesthetasc; all setae naked. Antenna (
<figureCitation id="379A2A23FFD5465FB450FB83FDC2FB92" box="[540,618,1077,1101]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3A</figureCitation>
) 4-segmented, composed of coxobasis and 3-segmented endopod; coxobasis large, bearing single naked seta; first endopodal segment bearing simple median seta on inner margin; second endopodal segment bearing three inner setae; third endopodal segment 0.73 times longer than wide, 27×37, bearing one proximal seta and three robust and one elongate claws and two setae on distal part. Labrum (
<figureCitation id="379A2A23FFD5465FB729FAAEFE04FAEE" box="[357,428,1304,1329]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3B</figureCitation>
) bearing pair of elongate posterior lobes. Mandible (
<figureCitation id="379A2A23FFD5465FB707FA8FFE3FFA8E" box="[331,407,1337,1361]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3C</figureCitation>
) bearing serrated blade with single hyaline outer tooth at base of convex margin. Maxillule (
<figureCitation id="379A2A23FFD5465FB495FAEFFF13FA4D" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3D</figureCitation>
) represented by simple lobe armed with one inner and three distal setae. Maxilla (
<figureCitation id="379A2A23FFD5465FB7FEFA2DFE55FA6C" box="[434,509,1435,1459]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3E</figureCitation>
) 2-segmented; basal segment unarmed; distal segment bearing serrated blade and one inner spinulose seta and two naked setae. Maxilliped (
<figureCitation id="379A2A23FFD5465FB4FDFA6AFD50FA2B" box="[689,760,1500,1524]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Fig. 3F</figureCitation>
) 3-segmented: syncoxa unarmed; basis with two simple setae; conical endopodal segment with spinulose tip and one large and one small elements. Legs 1 to 4 (
<figureCitation id="379A2A23FFD5465FB46FF98BFDDEF98A" box="[547,630,1597,1621]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Figs 3G</figureCitation>
,
<figureCitation id="379A2A23FFD5465FB4C8F98BFD17F98A" box="[644,703,1597,1621]" captionStart="Fig" captionStartId="4.[156,187,1551,1573]" captionTargetBox="[189,1397,207,1511]" captionTargetId="figure-81@4.[187,1399,205,1513]" captionTargetPageId="4" captionText="Fig. 4. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right leg 2, anterior; B, right leg 3, anterior; C, left leg 4, anterior; D, right leg 5, dorsal. Scale bars: AD, 50µm." figureDoi="http://doi.org/10.5281/zenodo.5739021" httpUri="https://zenodo.org/record/5739021/files/figure.png" pageId="1" pageNumber="250">4AC</figureCitation>
) biramous, with 3-segmented rami, except unsegmented leg 4 endopod. Leg armature formula as in
<tableCitation id="E223031DFFD5465FB7BBF9C8FD96F949" box="[503,574,1662,1686]" captionStart="Table 1" captionStartId="4.[156,204,1647,1669]" captionText="Table 1. Armature formula of legs 1 to 4 of Pseudanthessius chuukensis Uyeno, n. sp., adult female, BPBM-S18653. Arabic numbers= number of setae, Roman numbers=number of spines." pageId="1" pageNumber="250">Table 1</tableCitation>
. Intercoxal sclerites (
<figureCitation id="379A2A23FFD5465FB6EFF929FF5AF968" box="[163,242,1695,1719]" captionStart="Fig" captionStartId="3.[156,187,1756,1778]" captionTargetBox="[189,1397,206,1716]" captionTargetId="figure-83@3.[187,1399,205,1718]" captionTargetPageId="3" captionText="Fig. 3. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right antenna, anterior; B, labrum, anterior; C, left mandible, posterior; D, right maxillule, anterior; E, right maxilla, posterior; F, right maxilliped, anterior; G, left leg 1, anterior. Scale bars: A, B, G, 40µm; C, D, F, 20µm; E, 30µm." figureDoi="http://doi.org/10.5281/zenodo.5739017" httpUri="https://zenodo.org/record/5739017/files/figure.png" pageId="1" pageNumber="250">Figs 3G</figureCitation>
,
<figureCitation id="379A2A23FFD5465FB6B0F929FE90F968" box="[252,312,1695,1719]" captionStart="Fig" captionStartId="4.[156,187,1551,1573]" captionTargetBox="[189,1397,207,1511]" captionTargetId="figure-81@4.[187,1399,205,1513]" captionTargetPageId="4" captionText="Fig. 4. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right leg 2, anterior; B, right leg 3, anterior; C, left leg 4, anterior; D, right leg 5, dorsal. Scale bars: AD, 50µm." figureDoi="http://doi.org/10.5281/zenodo.5739021" httpUri="https://zenodo.org/record/5739021/files/figure.png" pageId="1" pageNumber="250">4AC</figureCitation>
) of legs 1 to 4 unornamented. All spines spatulate with serrated margins. All setae plumose. Rami of legs 1 to 4 bearing pointed projections on outer margins and rows of hair-like setules on both outer and inner margins. Leg 4 endopod (
<figureCitation id="379A2A23FFD5465FB69CF897FEB2F8E6" box="[208,282,1825,1849]" captionStart="Fig" captionStartId="4.[156,187,1551,1573]" captionTargetBox="[189,1397,207,1511]" captionTargetId="figure-81@4.[187,1399,205,1513]" captionTargetPageId="4" captionText="Fig. 4. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right leg 2, anterior; B, right leg 3, anterior; C, left leg 4, anterior; D, right leg 5, dorsal. Scale bars: AD, 50µm." figureDoi="http://doi.org/10.5281/zenodo.5739021" httpUri="https://zenodo.org/record/5739021/files/figure.png" pageId="1" pageNumber="250">Fig. 4C</figureCitation>
) unsegmented, 2.74 times longer than wide, with small notch on outer margin. Leg 5 (
<figureCitation id="379A2A23FFD5465FB45DF8F4FDF7F885" box="[529,607,1858,1882]" captionStart="Fig" captionStartId="4.[156,187,1551,1573]" captionTargetBox="[189,1397,207,1511]" captionTargetId="figure-81@4.[187,1399,205,1513]" captionTargetPageId="4" captionText="Fig. 4. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Right leg 2, anterior; B, right leg 3, anterior; C, left leg 4, anterior; D, right leg 5, dorsal. Scale bars: AD, 50µm." figureDoi="http://doi.org/10.5281/zenodo.5739021" httpUri="https://zenodo.org/record/5739021/files/figure.png" pageId="1" pageNumber="250">Fig. 4D</figureCitation>
) represented by simple spine and two naked setae. Leg 6 (
<figureCitation id="379A2A23FFD5465FB47AF8D4FDD7F8A5" box="[566,639,1890,1914]" captionStart="Fig" captionStartId="2.[157,188,1711,1733]" captionTargetBox="[190,1397,206,1669]" captionTargetId="figure-160@2.[187,1399,205,1673]" captionTargetPageId="2" captionText="Fig. 2. Pseudanthessius chuukensis Uyeno, n. sp., holotype adult female, BPBM-S18653. A, Habitus, dorsal; B, right genital aperture; C, right caudal ramus, dorsal; D, rostral area, ventral; E, right antennule, anterior. Scale bars: A, 200µm, B, 30µm; C, 100µm; D, 50µm; E, 40µm." figureDoi="http://doi.org/10.5281/zenodo.5739013" httpUri="https://zenodo.org/record/5739013/files/figure.png" pageId="1" pageNumber="250">Fig. 2B</figureCitation>
) represented by two small setae at genital opening.
</paragraph>
<paragraph id="AF1E36A6FFD5465FB6F9F815FD9CF864" blockId="1.[156,768,784,2077]" box="[181,564,1955,1979]" pageId="1" pageNumber="250">
<emphasis id="9DD5EAB4FFD5465FB6F9F815FEF8F864" bold="true" box="[181,336,1955,1979]" pageId="1" pageNumber="250">Infection sites.</emphasis>
Gill to buccal cavities.
</paragraph>
</subSubSection>
<subSubSection id="E7BB652DFFD5465FB6F9F872FC03F98A" pageId="1" pageNumber="250" type="discussion">
<paragraph id="AF1E36A6FFD5465FB6F9F872FC03F98A" blockId="1.[156,768,784,2077]" lastBlockId="1.[820,1432,784,1687]" pageId="1" pageNumber="250">
<emphasis id="9DD5EAB4FFD5465FB6F9F872FEBFF803" bold="true" box="[181,279,1988,2012]" pageId="1" pageNumber="250">Remarks.</emphasis>
Females of nine congeners [
<taxonomicName id="68A14D25FFD5465FB42FF872FDDEF823" authority="Stock, Humes, and Gooding, 1964" authorityName="Stock, Humes, and Gooding" authorityYear="1964" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aestheticus">
<emphasis id="9DD5EAB4FFD5465FB42FF872FEADF823" italics="true" pageId="1" pageNumber="250">Pseudanthessius aestheticus</emphasis>
<bibRefCitation id="CB304B57FFD5465FB743F852FDDEF823" author="Stock, J. H. &amp; Humes, A. G. &amp; Gooding, R. U." box="[271,630,2020,2044]" pageId="1" pageNumber="250" pagination="1 - 74" refId="ref3230" refString="Stock, J. H., Humes, A. G., and Gooding, R. U. 1964. Copepoda associated with West Indian invertebrates. IV. The genera Octopicola, Pseudanthessius and Meomicola (Cyclopoida, Lichomolgidae). Studies on the Fauna of Curacao 18: 1 - 74." type="journal article" year="1964">Stock, Humes, and Gooding, 1964</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB4CFF853FE00F7C2" authority="Thompson and Scott, 1903" authorityName="Thompson and Scott" authorityYear="1903" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="9DD5EAB4FFD5465FB4CFF853FD57F823" box="[643,767,2021,2044]" italics="true" pageId="1" pageNumber="250">P. concinnus</emphasis>
Thompson and Scott, 1903
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB7FDF7B0FD21F7C2" authority="Kim, 2000" authorityName="Kim" authorityYear="2000" box="[433,649,2053,2077]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="9DD5EAB4FFD5465FB7FDF7B0FDB2F7C2" box="[433,538,2053,2077]" italics="true" pageId="1" pageNumber="250">P. dentatus</emphasis>
<bibRefCitation id="CB304B57FFD5465FB46CF7B3FD21F7C2" author="Kim, I. H." box="[544,649,2053,2077]" pageId="1" pageNumber="250" pagination="367 - 432" refId="ref3026" refString="Kim, I. H. 2000. Poecilostomatoid copepods from an intertidal mud flat in the Yellow Sea. Journal of Natural History 34: 367 - 432." type="journal article" year="2000">Kim, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB4DFF7B0FC7DFCF7" authority="Sewell, 1949" authorityName="Sewell" authorityYear="1949" box="[659,981,784,2077]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="gracilioides">
<emphasis id="9DD5EAB4FFD5465FB4DFF7B0FCFCFCF7" box="[659,852,784,2077]" italics="true" pageId="1" pageNumber="250">P. gracilioides</emphasis>
<bibRefCitation id="CB304B57FFD5465FB515FCA6FC7DFCF7" author="Sewell, R. B. S." box="[857,981,784,808]" pageId="1" pageNumber="250" pagination="17 - 199" refId="ref3158" refString="Sewell, R. B. S. 1949. The littoral and semi-parasitic Cyclopoida, the Monstrilloida and Notodelphyoida. The John Murray Expedition, 1933 - 34. Scientific Reports 9: 17 - 199." type="journal article" year="1949">Sewell, 1949</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB592FCA7FB1AFCF7" authority="Claus, 1889" authorityName="Claus" authorityYear="1889" box="[990,1202,784,808]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="9DD5EAB4FFD5465FB592FCA7FB90FCF7" box="[990,1080,784,808]" italics="true" pageId="1" pageNumber="250">P. gracilis</emphasis>
<bibRefCitation id="CB304B57FFD5465FB271FCA6FB1AFCF7" author="Claus, C." box="[1085,1202,784,808]" pageId="1" pageNumber="250" pagination="327 - 370" refId="ref2744" refString="Claus, C. 1889. Uber neue oder wenig bekannte halbparasitische Copepoden insbesondere der Lichomolgiden- und Ascomyzontiden- Gruppe. Arbeiten des Zoologischen Institut der Universitat Wien 8: 327 - 370, pls 21 - 27." type="journal article" year="1889">Claus, 1889</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB2F7FCA7FACCFCF7" authority="Illg, 1950" authorityName="Illg" authorityYear="1950" box="[1211,1380,784,808]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="9DD5EAB4FFD5465FB2F7FCA7FAA8FCF7" box="[1211,1280,784,808]" italics="true" pageId="1" pageNumber="250">P.latus</emphasis>
<bibRefCitation id="CB304B57FFD5465FB34AFCA6FACCFCF7" author="Illg" box="[1286,1380,784,808]" pageId="1" pageNumber="250" pagination="129 - 133" refId="ref2987" refString="Illg, P. L. 1950. A new copepod, Pseudanthessius latus (Cyclopoida: Lichomolgidae), commensal with a marine flatworm. Journal of the Washington Academy of Sciences 40: 129 - 133." type="journal article" year="1950">Illg, 1950</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB321FCA7FC55FC96" authority="Humes, 1978" authorityName="Humes" authorityYear="1978" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="9DD5EAB4FFD5465FB321FCA7FCD9FC97" italics="true" pageId="1" pageNumber="250">P.limatus</emphasis>
<bibRefCitation id="CB304B57FFD5465FB53BFC87FC55FC96" author="Humes, A. G." box="[887,1021,817,841]" pageId="1" pageNumber="250" pagination="242 - 249" refId="ref2820" refString="Humes, A. G. 1978. A new cyclopoid copepod, Pseudanthessius limatus, associated with an ophiuroid in Panama (Atlantic side). Proceedings of the Biological Society of Washington 91: 242 - 249." type="journal article" year="1978">Humes, 1978</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB24BFC87FA3CFC97" authority="(Brady and Robertson, 1875)" baseAuthorityName="Brady and Robertson" baseAuthorityYear="1875" box="[1031,1428,816,840]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="thorellii">
<emphasis id="9DD5EAB4FFD5465FB24BFC87FBC0FC97" box="[1031,1128,816,840]" italics="true" pageId="1" pageNumber="250">P. thorellii</emphasis>
(Brady and Robertson, 1875)
</taxonomicName>
, and
<taxonomicName id="68A14D25FFD5465FB528FCE4FB96FCB6" authority="Scott, 1909" authorityName="Scott" authorityYear="1909" box="[868,1086,849,873]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="9DD5EAB4FFD5465FB528FCE4FC17FCB6" box="[868,959,849,873]" italics="true" pageId="1" pageNumber="250">P. weberi</emphasis>
<bibRefCitation id="CB304B57FFD5465FB585FCE7FB96FCB6" author="Scott, A." box="[969,1086,849,873]" pageId="1" pageNumber="250" pagination="1 - 69" refId="ref3121" refString="Scott, A. 1909. The Copepoda of the Siboga Expedition. Part I. Freeswimming, littoral and semi-parasitic Copepoda. Siboga Expeditie 29 a: 1 - 323, pls 1 - 69." type="book chapter" year="1909">Scott, 1909</bibRefCitation>
</taxonomicName>
] share the 5-segmented urosome and caudal rami which are more than four times longer than wide with
<taxonomicName id="68A14D25FFD5465FB5ECFC25FB8AFC75" authority="Uyeno &amp; Rain, 2021" authorityName="Uyeno &amp; Rain" authorityYear="2021" box="[928,1058,914,938]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="chuukensis" status="sp. nov.">
<emphasis id="9DD5EAB4FFD5465FB5ECFC25FB8AFC75" box="[928,1058,914,938]" italics="true" pageId="1" pageNumber="250">P. chuukensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="86E657CFFFD5465FB266FC25FBC9FC75" box="[1066,1121,915,938]" pageId="1" pageNumber="250" rank="species">n. sp.</taxonomicNameLabel>
Of these congeners,
<taxonomicName id="68A14D25FFD5465FB374FC25FCF0FC15" authorityName="Thompson and Scott" authorityYear="1903" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="9DD5EAB4FFD5465FB374FC25FCF0FC15" italics="true" pageId="1" pageNumber="250">P. concinnus</emphasis>
</taxonomicName>
and
<taxonomicName id="68A14D25FFD5465FB5C5FC05FC42FC14" baseAuthorityName="Brady and Robertson" baseAuthorityYear="1875" box="[905,1002,947,971]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="thorellii">
<emphasis id="9DD5EAB4FFD5465FB5C5FC05FC42FC14" box="[905,1002,947,971]" italics="true" pageId="1" pageNumber="250">P. thorellii</emphasis>
</taxonomicName>
clearly differ from the new species by having caudal rami which are more than ten times longer than wide (vs. 5.69 times) (e.g.,
<bibRefCitation id="CB304B57FFD5465FB21EFC42FAABFBD3" author="Stock, J. H. &amp; Humes, A. G. &amp; Gooding, R. U." box="[1106,1283,1012,1036]" pageId="1" pageNumber="250" pagination="1 - 74" refId="ref3230" refString="Stock, J. H., Humes, A. G., and Gooding, R. U. 1964. Copepoda associated with West Indian invertebrates. IV. The genera Octopicola, Pseudanthessius and Meomicola (Cyclopoida, Lichomolgidae). Studies on the Fauna of Curacao 18: 1 - 74." type="journal article" year="1964">Stock et al. 1964</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB35CFC42FA3CFBD3" author="Humes, A. G." box="[1296,1428,1012,1036]" pageId="1" pageNumber="250" pagination="242 - 249" refId="ref2820" refString="Humes, A. G. 1978. A new cyclopoid copepod, Pseudanthessius limatus, associated with an ophiuroid in Panama (Atlantic side). Proceedings of the Biological Society of Washington 91: 242 - 249." type="journal article" year="1978">Humes 1978</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB579FBA2FC36FBF3" author="Kim, I. H." box="[821,926,1044,1068]" pageId="1" pageNumber="250" pagination="367 - 432" refId="ref3026" refString="Kim, I. H. 2000. Poecilostomatoid copepods from an intertidal mud flat in the Yellow Sea. Journal of Natural History 34: 367 - 432." type="journal article" year="2000">Kim 2000</bibRefCitation>
).
<taxonomicName id="68A14D25FFD5465FB5FDFBA2FB16FBF3" authorityName="Stock, Humes, and Gooding" authorityYear="1964" box="[945,1214,1044,1068]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="aestheticus">
<emphasis id="9DD5EAB4FFD5465FB5FDFBA2FB16FBF3" box="[945,1214,1044,1068]" italics="true" pageId="1" pageNumber="250">Pseudanthessius aestheticus</emphasis>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB281FBA3FA92FBF3" authorityName="Kim" authorityYear="2000" box="[1229,1338,1044,1068]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="9DD5EAB4FFD5465FB281FBA3FA92FBF3" box="[1229,1338,1044,1068]" italics="true" pageId="1" pageNumber="250">P. dentatus</emphasis>
</taxonomicName>
,
<taxonomicName id="68A14D25FFD5465FB305FBA3FA3AFBF3" authorityName="Illg" authorityYear="1950" box="[1353,1426,1044,1068]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="9DD5EAB4FFD5465FB305FBA3FA3AFBF3" box="[1353,1426,1044,1068]" italics="true" pageId="1" pageNumber="250">P. latus</emphasis>
</taxonomicName>
, and
<taxonomicName id="68A14D25FFD5465FB52EFB80FC6AFB92" authorityName="Humes" authorityYear="1978" box="[866,962,1077,1101]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="9DD5EAB4FFD5465FB52EFB80FC6AFB92" box="[866,962,1077,1101]" italics="true" pageId="1" pageNumber="250">P. limatus</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="68A14D25FFD5465FB2F4FB80FA91FB92" authority="Uyeno &amp; Rain, 2021" authorityName="Uyeno &amp; Rain" authorityYear="2021" box="[1208,1337,1077,1101]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="chuukensis" status="sp. nov.">
<emphasis id="9DD5EAB4FFD5465FB2F4FB80FA91FB92" box="[1208,1337,1077,1101]" italics="true" pageId="1" pageNumber="250">P. chuukensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="86E657CFFFD5465FB30DFB83FAD0FB93" box="[1345,1400,1077,1100]" pageId="1" pageNumber="250" rank="species">n. sp.</taxonomicNameLabel>
by the leg armature formula: the distal endopodal segment of leg 1 has one spine and five setae in these species (vs. three spines and three setae in the new species); and the endopod of leg 4 has two spines (vs. replaced by two setae in the new species) (
<bibRefCitation id="CB304B57FFD5465FB577FB61FC3DFB30" author="Illg" box="[827,917,1239,1263]" pageId="1" pageNumber="250" pagination="129 - 133" refId="ref2987" refString="Illg, P. L. 1950. A new copepod, Pseudanthessius latus (Cyclopoida: Lichomolgidae), commensal with a marine flatworm. Journal of the Washington Academy of Sciences 40: 129 - 133." type="journal article" year="1950">Illg 1950</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB5D3FB61FBE9FB30" author="Stock, J. H. &amp; Humes, A. G. &amp; Gooding, R. U." box="[927,1089,1239,1263]" pageId="1" pageNumber="250" pagination="1 - 74" refId="ref3230" refString="Stock, J. H., Humes, A. G., and Gooding, R. U. 1964. Copepoda associated with West Indian invertebrates. IV. The genera Octopicola, Pseudanthessius and Meomicola (Cyclopoida, Lichomolgidae). Studies on the Fauna of Curacao 18: 1 - 74." type="journal article" year="1964">Stock et al. 1964</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB206FB6EFB61FB2F" author="Humes, A. G." box="[1098,1225,1240,1264]" pageId="1" pageNumber="250" pagination="242 - 249" refId="ref2820" refString="Humes, A. G. 1978. A new cyclopoid copepod, Pseudanthessius limatus, associated with an ophiuroid in Panama (Atlantic side). Proceedings of the Biological Society of Washington 91: 242 - 249." type="journal article" year="1978">Humes 1978</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB29EFB61FA90FB2F" author="Kim, I. H." box="[1234,1336,1239,1264]" pageId="1" pageNumber="250" pagination="367 - 432" refId="ref3026" refString="Kim, I. H. 2000. Poecilostomatoid copepods from an intertidal mud flat in the Yellow Sea. Journal of Natural History 34: 367 - 432." type="journal article" year="2000">Kim 2000</bibRefCitation>
). In addition to those two characters,
<taxonomicName id="68A14D25FFD5465FB211FB4FFB49FACF" authorityName="Sewell" authorityYear="1949" box="[1117,1249,1272,1296]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="gracilioides">
<emphasis id="9DD5EAB4FFD5465FB211FB4FFB49FACF" box="[1117,1249,1272,1296]" italics="true" pageId="1" pageNumber="250">P. gracilioides</emphasis>
</taxonomicName>
is separated from the new species by having the distal exopodal segment of leg 1 with five inner setae (vs. four setae) and both rami of leg 4 being nearly equal in length (vs. the endopod being about two-thirds as long as the exopod) (
<bibRefCitation id="CB304B57FFD5465FB2C1FACCFAA0FA4D" author="Sewell, R. B. S." box="[1165,1288,1402,1426]" pageId="1" pageNumber="250" pagination="17 - 199" refId="ref3158" refString="Sewell, R. B. S. 1949. The littoral and semi-parasitic Cyclopoida, the Monstrilloida and Notodelphyoida. The John Murray Expedition, 1933 - 34. Scientific Reports 9: 17 - 199." type="journal article" year="1949">Sewell 1949</bibRefCitation>
).
<taxonomicName id="68A14D25FFD5465FB35BFACCFC08FA6C" authorityName="Scott" authorityYear="1909" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="9DD5EAB4FFD5465FB35BFACCFC08FA6C" italics="true" pageId="1" pageNumber="250">Pseudanthessius weberi</emphasis>
</taxonomicName>
and
<taxonomicName id="68A14D25FFD5465FB598FA2DFB87FA6C" authorityName="Claus" authorityYear="1889" box="[980,1071,1435,1459]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="9DD5EAB4FFD5465FB598FA2DFB87FA6C" box="[980,1071,1435,1459]" italics="true" pageId="1" pageNumber="250">P. gracilis</emphasis>
</taxonomicName>
, in which the leg armature formula was insufficiently shown in the original descriptions, differ from
<taxonomicName id="68A14D25FFD5465FB527FA6AFC42FA2B" authority="Uyeno &amp; Rain, 2021" authorityName="Uyeno &amp; Rain" authorityYear="2021" box="[875,1002,1500,1524]" class="Hexanauplia" family="Pseudanthessiidae" genus="Pseudanthessius" kingdom="Animalia" order="Cyclopoida" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="chuukensis" status="sp. nov.">
<emphasis id="9DD5EAB4FFD5465FB527FA6AFC42FA2B" box="[875,1002,1500,1524]" italics="true" pageId="1" pageNumber="250">P. chuukensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="86E657CFFFD5465FB5BCFA6AFB8DFA2C" box="[1008,1061,1500,1523]" pageId="1" pageNumber="250" rank="species">n. sp.</taxonomicNameLabel>
in having the endopod of leg 4 tapering distally with a proximal swelling on the inner margin (vs. not tapering distally without proximal swellings) (
<bibRefCitation id="CB304B57FFD5465FB36FF9ABFA3CF9EA" author="Claus, C." box="[1315,1428,1565,1589]" pageId="1" pageNumber="250" pagination="327 - 370" refId="ref2744" refString="Claus, C. 1889. Uber neue oder wenig bekannte halbparasitische Copepoden insbesondere der Lichomolgiden- und Ascomyzontiden- Gruppe. Arbeiten des Zoologischen Institut der Universitat Wien 8: 327 - 370, pls 21 - 27." type="journal article" year="1889">Claus 1889</bibRefCitation>
;
<bibRefCitation id="CB304B57FFD5465FB579F98BFC08F98A" author="Scott, A." box="[821,928,1597,1621]" pageId="1" pageNumber="250" pagination="1 - 69" refId="ref3121" refString="Scott, A. 1909. The Copepoda of the Siboga Expedition. Part I. Freeswimming, littoral and semi-parasitic Copepoda. Siboga Expeditie 29 a: 1 - 323, pls 1 - 69." type="book chapter" year="1909">Scott 1909</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E7BB652DFFD5465FB502F9E8FC73F949" pageId="1" pageNumber="250" type="etymology">
<paragraph id="AF1E36A6FFD5465FB502F9E8FC73F949" blockId="1.[820,1432,784,1687]" pageId="1" pageNumber="250">
<emphasis id="9DD5EAB4FFD5465FB502F9E8FC6DF9A9" bold="true" box="[846,965,1630,1654]" pageId="1" pageNumber="250">Etymology.</emphasis>
The specific name of the new species refers to the
<typeStatus id="701A8804FFD5465FB516F9C9FC2EF948" box="[858,902,1663,1687]" pageId="1" pageNumber="250">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>