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<mods:title id="038A52BD22E53D13067CE414BDF5615A">Phylogeny and systematics of the Trapeziidae Miers, 1886 (Crustacea: Brachyura), with the description of a new family</mods:title>
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<treatment id="162E87D383681F4FFA7DFB81FE4EF8AE" ID-DOI="http://doi.org/10.5281/zenodo.5675111" ID-GBIF-Taxon="120829199" ID-Zenodo-Dep="5675111" LSID="urn:lsid:plazi:treatment:162E87D383681F4FFA7DFB81FE4EF8AE" httpUri="http://treatment.plazi.org/id/162E87D383681F4FFA7DFB81FE4EF8AE" lastPageNumber="54" pageId="53" pageNumber="54">
<subSubSection id="D69D654E83681F4FFA7DFB81FC87FBD3" box="[264,778,1033,1059]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="9E3836C583681F4FFA7DFB81FC87FBD3" blockId="53.[264,778,1033,1059]" box="[264,778,1033,1059]" pageId="53" pageNumber="54">
<heading id="C57081A983681F4FFA7DFB81FC87FBD3" bold="true" box="[264,778,1033,1059]" fontSize="11" level="5" pageId="53" pageNumber="54" reason="0">
<taxonomicName id="59874D4683681F4FFA7DFB81FC87FBD3" ID-CoL="57Y2N" authority="Eydoux &amp; Souleyet, 1842" authorityName="Eydoux &amp; Souleyet" authorityYear="1842" box="[264,778,1033,1059]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FFA7DFB81FC87FBD3" bold="true" box="[264,778,1033,1059]" pageId="53" pageNumber="54">
<emphasis id="ACF3EAD783681F4FFA7DFB81FE44FBD3" bold="true" box="[264,457,1033,1059]" italics="true" pageId="53" pageNumber="54">Trapezia tigrina</emphasis>
<bibRefCitation id="FA164B3483681F4FFAA4FB81FC87FBD3" author="Eydoux" box="[465,778,1033,1059]" pageId="53" pageNumber="54" refString="Eydoux, F. &amp; Souleyet, F. A. (1842) Crustaces. In: Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette la Bonite, commandee par M. Vaillant. Zoologie, 1 (2), 107 - 328; Atlas, pls. 1 - 150." type="journal article" year="1842">Eydoux &amp; Souleyet, 1842</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="D69D654E83681F4FFA7DFBC7FDF4FB35" pageId="53" pageNumber="54" type="reference_group">
<paragraph id="9E3836C583681F4FFA7DFBC7FD13FB75" blockId="53.[264,1158,1102,1222]" pageId="53" pageNumber="54">
<treatmentCitationGroup id="BE9711EB83681F4FFA7DFBC7FD13FB75" pageId="53" pageNumber="54">
<taxonomicName id="59874D4683681F4FFA7DFBC7FC84FB95" ID-CoL="57Y2N" authority="Eydoux &amp; Souleyet, 1842: 232" authorityName="Eydoux &amp; Souleyet" authorityPageNumber="232" authorityYear="1842" box="[264,777,1102,1126]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FFA7DFBC7FE39FB96" box="[264,436,1103,1126]" italics="true" pageId="53" pageNumber="54">Trapezia tigrina</emphasis>
<treatmentCitation id="1F2610D483681F4FFAC9FBC6FC84FB95" author="Eydoux" box="[444,777,1102,1125]" page="232" pageId="53" pageNumber="54" year="1842">
<bibRefCitation id="FA164B3483681F4FFAC9FBC6FD59FB95" author="Eydoux" box="[444,724,1102,1125]" pageId="53" pageNumber="54" refString="Eydoux, F. &amp; Souleyet, F. A. (1842) Crustaces. In: Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette la Bonite, commandee par M. Vaillant. Zoologie, 1 (2), 107 - 328; Atlas, pls. 1 - 150." type="journal article" year="1842">Eydoux &amp; Souleyet, 1842</bibRefCitation>
: 232
</treatmentCitation>
</taxonomicName>
, pl. 2, fig. 4 (colour) (see Plate 3E).
<taxonomicName id="59874D4683681F4FFA7DFBE7FD17FB75" ID-CoL="7CMPD" authority="Coulon, 1864: 569" authorityName="Coulon" authorityPageNumber="569" authorityYear="1864" box="[264,666,1134,1158]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="ACF3EAD783681F4FFA7DFBE7FE44FB76" box="[264,457,1135,1158]" italics="true" pageId="53" pageNumber="54">Trapezia punctata</emphasis>
<treatmentCitation id="1F2610D483681F4FFAA5FBE6FD17FB75" author="Coulon" box="[464,666,1134,1157]" page="569" pageId="53" pageNumber="54" year="1864">
<bibRefCitation id="FA164B3483681F4FFAA5FBE6FDE8FB75" author="Coulon" box="[464,613,1134,1157]" pageId="53" pageNumber="54" refString="Coulon, L. (1864) Crustaces de la Mer Rouge. Bulletin de la Societe des Sciences Naturelles de Neuchatel, 6, 569 - 570." type="journal article" year="1864">Coulon, 1864</bibRefCitation>
: 569
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E3836C583681F4FFA7DFB07FD69FB55" blockId="53.[264,1158,1102,1222]" box="[264,740,1166,1190]" pageId="53" pageNumber="54">
<treatmentCitationGroup id="BE9711EB83681F4FFA7DFB07FD69FB55" box="[264,740,1166,1190]" pageId="53" pageNumber="54">
<taxonomicName id="59874D4683681F4FFA7DFB07FDD4FB55" ID-CoL="57XZK" authority="Ward, 1939: 13" authorityName="Ward" authorityPageNumber="13" authorityYear="1939" box="[264,601,1166,1190]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="danae">
<emphasis id="ACF3EAD783681F4FFA7DFB07FE20FB56" box="[264,429,1167,1190]" italics="true" pageId="53" pageNumber="54">Trapezia danae</emphasis>
<treatmentCitation id="1F2610D483681F4FFAC1FB06FDD4FB55" author="Ward" box="[436,601,1166,1189]" page="13" pageId="53" pageNumber="54" year="1939">
<bibRefCitation id="FA164B3483681F4FFAC1FB06FDBEFB55" author="Ward" box="[436,563,1166,1189]" pageId="53" pageNumber="54" refString="Ward, M. (1939) The Brachyura of the second Templeton Crocker-American Museum Expedition to the Pacific Ocean. American Museum Novitates, 1049, 1 - 15." type="journal article" year="1939">Ward, 1939</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
, figs. 17, 18.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E3836C583681F4FFA7DFB27FDF4FB35" blockId="53.[264,1158,1102,1222]" box="[264,633,1198,1222]" pageId="53" pageNumber="54">
<treatmentCitationGroup id="BE9711EB83681F4FFA7DFB27FDF4FB35" box="[264,633,1198,1222]" pageId="53" pageNumber="54">
<taxonomicName id="59874D4683681F4FFA7DFB27FDF9FB35" ID-CoL="57Y2P" authority="Serene, 1971: 140" authorityName="Serene" authorityPageNumber="140" authorityYear="1971" box="[264,628,1198,1222]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="ACF3EAD783681F4FFA7DFB27FE2AFB36" box="[264,423,1199,1222]" italics="true" pageId="53" pageNumber="54">Trapezia wardi</emphasis>
<treatmentCitation id="1F2610D483681F4FFAC5FB26FDF9FB35" author="Serene" box="[432,628,1198,1221]" page="140" pageId="53" pageNumber="54" year="1971">
<bibRefCitation id="FA164B3483681F4FFAC5FB26FDB2FB35" author="Serene" box="[432,575,1198,1221]" pageId="53" pageNumber="54" refString="Serene, R. (1971) Observations preliminaires sur des Brachyoures nouveaux ou mal connus du Sud-est Asiatique (Crustacea Decapoda). Bulletin du Museum National d'Histoire Naturelle (Paris) ser. 2, 42 (5), 903 - 918." type="journal article" year="1971">Serène, 1971</bibRefCitation>
: 140
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D69D654E83681F4FFA7DFB64FE4EF8AE" pageId="53" pageNumber="54" type="discussion">
<paragraph id="9E3836C583681F4FFA7DFB64FD72F99E" blockId="53.[264,1323,1260,1886]" pageId="53" pageNumber="54">
<emphasis id="ACF3EAD783681F4FFA7DFB64FEE3FAF5" box="[264,366,1260,1285]" italics="true" pageId="53" pageNumber="54">Remarks</emphasis>
. The identity of
<taxonomicName id="59874D4683681F4FF932FB64FCD5FAF6" authority="Serene, 1971" authorityName="Serene" authorityYear="1971" box="[583,856,1260,1286]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="ACF3EAD783681F4FF932FB64FD27FAF5" box="[583,682,1260,1285]" italics="true" pageId="53" pageNumber="54">T. wardi</emphasis>
<bibRefCitation id="FA164B3483681F4FF9CCFB64FCD5FAF6" author="Serene" box="[697,856,1260,1286]" pageId="53" pageNumber="54" refString="Serene, R. (1971) Observations preliminaires sur des Brachyoures nouveaux ou mal connus du Sud-est Asiatique (Crustacea Decapoda). Bulletin du Museum National d'Histoire Naturelle (Paris) ser. 2, 42 (5), 903 - 918." type="journal article" year="1971">Serène, 1971</bibRefCitation>
</taxonomicName>
(
<typeStatus id="413C886783681F4FF819FB64FC2DFAF6" box="[876,928,1260,1286]" pageId="53" pageNumber="54">type</typeStatus>
locality
<collectingCountry id="E690765583681F4FFF65FB64FBFBFAF6" box="[1040,1142,1260,1286]" name="Vietnam" pageId="53" pageNumber="54">Vietnam</collectingCountry>
) and
<taxonomicName id="59874D4683681F4FFFB4FB64FE1DFADE" authority="Ward, 1939" authorityName="Ward" authorityYear="1939" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="danae">
<emphasis id="ACF3EAD783681F4FFFB4FB64FAA6FAF5" box="[1217,1323,1260,1285]" italics="true" pageId="53" pageNumber="54">T. danae</emphasis>
<bibRefCitation id="FA164B3483681F4FFA7DFA9CFE1DFADE" author="Ward" box="[264,400,1300,1326]" pageId="53" pageNumber="54" refString="Ward, M. (1939) The Brachyura of the second Templeton Crocker-American Museum Expedition to the Pacific Ocean. American Museum Novitates, 1049, 1 - 15." type="journal article" year="1939">Ward, 1939</bibRefCitation>
</taxonomicName>
(
<typeStatus id="413C886783681F4FFAEAFA9CFE5EFADE" box="[415,467,1300,1326]" pageId="53" pageNumber="54">type</typeStatus>
locality
<collectingCountry id="E690765583681F4FF94CFA9CFD07FADE" box="[569,650,1300,1326]" name="Samoa" pageId="53" pageNumber="54">Samoa</collectingCountry>
) with
<taxonomicName id="59874D4683681F4FF9A0FA9CFBF7FADE" authority="Eydoux &amp; Souleyet, 1842" authorityName="Eydoux &amp; Souleyet" authorityYear="1842" box="[725,1146,1300,1326]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FF9A0FA9CFCCCFADD" box="[725,833,1300,1325]" italics="true" pageId="53" pageNumber="54">T. tigrina</emphasis>
<bibRefCitation id="FA164B3483681F4FF83DFA9CFBF7FADE" author="Eydoux" box="[840,1146,1300,1326]" pageId="53" pageNumber="54" refString="Eydoux, F. &amp; Souleyet, F. A. (1842) Crustaces. In: Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette la Bonite, commandee par M. Vaillant. Zoologie, 1 (2), 107 - 328; Atlas, pls. 1 - 150." type="journal article" year="1842">Eydoux &amp; Souleyet, 1842</bibRefCitation>
</taxonomicName>
, is not in question (see
<bibRefCitation id="FA164B3483681F4FFA0FFAB4FD20FAA6" author="Galil" box="[378,685,1340,1366]" pageId="53" pageNumber="54" refString="Galil, B. &amp; Lewinsohn, C. (1984) On the taxonomic status of Trapezia tigrina Eydoux &amp; Souleyet, 1842 (Decapoda, Brachyura). Crustaceana, 46 (2), 166 - 175." type="journal article" year="1984">Galil &amp; Lewinsohn 1984</bibRefCitation>
: 166;
<bibRefCitation id="FA164B3483681F4FF98FFAB4FC12FAA6" author="Castro" box="[762,927,1340,1366]" pageId="53" pageNumber="54" refString="Castro, P. (1997 b) Trapeziid crabs (Brachyura: Xanthoidea: Trapeziidae) of French Polynesia. In: Richer de Forges, B. (Ed.), Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, Benthos). Etudes et Theses, 3, 109 - 139." type="journal article" year="1997" yearSuffix="b">Castro 1997b</bibRefCitation>
: 130).
<taxonomicName id="59874D4683681F4FF883FAB4FAA6FAA6" authority="Coulon, 1864" authorityName="Coulon" authorityYear="1864" box="[1014,1323,1340,1366]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="ACF3EAD783681F4FF883FAB4FBF0FAA5" box="[1014,1149,1340,1365]" italics="true" pageId="53" pageNumber="54">T. punctata</emphasis>
<bibRefCitation id="FA164B3483681F4FFFF2FAB4FAA6FAA6" author="Coulon" box="[1159,1323,1340,1366]" pageId="53" pageNumber="54" refString="Coulon, L. (1864) Crustaces de la Mer Rouge. Bulletin de la Societe des Sciences Naturelles de Neuchatel, 6, 569 - 570." type="journal article" year="1864">Coulon, 1864</bibRefCitation>
</taxonomicName>
(
<typeStatus id="413C886783681F4FFA65FAECFEC9FA8E" box="[272,324,1380,1406]" pageId="53" pageNumber="54">type</typeStatus>
locality Red Sea) is almost certainly identical with
<taxonomicName id="59874D4683681F4FF8A3FAECFBCAFA8D" box="[982,1095,1380,1405]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FF8A3FAECFBCAFA8D" box="[982,1095,1380,1405]" italics="true" pageId="53" pageNumber="54">T. tigrina</emphasis>
</taxonomicName>
, even if Coulon’s description is far too brief. What appears to be Coulon's
<typeStatus id="413C886783681F4FF8D9FA04FC6CFA56" box="[940,993,1420,1446]" pageId="53" pageNumber="54">type</typeStatus>
, a dried specimen, is in the zoology collections of the University of Neuchâtel,
<collectingCountry id="E690765583681F4FF804FA3CFB8CFA3E" box="[881,1025,1460,1486]" name="Switzerland" pageId="53" pageNumber="54">Switzerland</collectingCountry>
. It carries the label &quot;
<taxonomicName id="59874D4683681F4FFF8DFA3CFDB8FA06" authority="Herb." authorityName="Herb." class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783681F4FFF8DFA3CFE6BFA05" italics="true" pageId="53" pageNumber="54">Trapezia rufopunctata</emphasis>
Herb.
</taxonomicName>
&quot; made by Coulon himself. This specimen was very likely used by Coulon in his description of
<taxonomicName id="59874D4683681F4FF908F98CFC8DF9ED" box="[637,768,1540,1565]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="ACF3EAD783681F4FF908F98CFC8DF9ED" box="[637,768,1540,1565]" italics="true" pageId="53" pageNumber="54">T. punctata</emphasis>
</taxonomicName>
, but Coulon apparently realised later that there was an earlier name (
<taxonomicName id="59874D4683681F4FF97AF9A4FD4BF9B5" box="[527,710,1580,1605]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783681F4FF97AF9A4FD4BF9B5" box="[527,710,1580,1605]" italics="true" pageId="53" pageNumber="54">T. rufopunctata</emphasis>
</taxonomicName>
) and his species was only a junior synonym (J.­P. Haenni, pers. comm. through M. Kottelat).
</paragraph>
<paragraph id="9E3836C583681F4FFA4DF9F4FE4EF8AE" blockId="53.[264,1323,1260,1886]" pageId="53" pageNumber="54">
As
<taxonomicName id="59874D4683681F4FFA16F9F4FE5FF965" box="[355,466,1660,1685]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FFA16F9F4FE5FF965" box="[355,466,1660,1685]" italics="true" pageId="53" pageNumber="54">T. tigrina</emphasis>
</taxonomicName>
and
<taxonomicName id="59874D4683681F4FF961F9F4FD46F965" box="[532,715,1660,1685]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783681F4FF961F9F4FD46F965" box="[532,715,1660,1685]" italics="true" pageId="53" pageNumber="54">T. rufopunctata</emphasis>
</taxonomicName>
have very similar colour patterns, and as a result were frequently mistaken for each other, Coulon's eventual identification is not at all surprising. Since
<taxonomicName id="59874D4683681F4FFAC5F944FDE8F915" box="[432,613,1740,1765]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783681F4FFAC5F944FDE8F915" box="[432,613,1740,1765]" italics="true" pageId="53" pageNumber="54">T. rufopunctata</emphasis>
</taxonomicName>
is not known from the Red Sea, Coulon's
<taxonomicName id="59874D4683681F4FFF27F944FB5BF915" box="[1106,1238,1740,1765]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="ACF3EAD783681F4FFF27F944FB5BF915" box="[1106,1238,1740,1765]" italics="true" pageId="53" pageNumber="54">T. punctata</emphasis>
</taxonomicName>
should be
<taxonomicName id="59874D4683681F4FFA5EF97CFE1BF8FD" box="[299,406,1780,1805]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FFA5EF97CFE1BF8FD" box="[299,406,1780,1805]" italics="true" pageId="53" pageNumber="54">T. tigrina</emphasis>
</taxonomicName>
instead. The colour description of
<taxonomicName id="59874D4683681F4FF85BF97CFC3CF8FD" box="[814,945,1780,1805]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="ACF3EAD783681F4FF85BF97CFC3CF8FD" box="[814,945,1780,1805]" italics="true" pageId="53" pageNumber="54">T. punctata</emphasis>
</taxonomicName>
also fits better that of
<taxonomicName id="59874D4683681F4FFFC2F97CFAAEF8FD" box="[1207,1315,1780,1805]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="ACF3EAD783681F4FFFC2F97CFAAEF8FD" box="[1207,1315,1780,1805]" italics="true" pageId="53" pageNumber="54">T. tigrina</emphasis>
</taxonomicName>
, orange dots (&quot;
<emphasis id="ACF3EAD783681F4FFADAF894FDDFF8C5" box="[431,594,1820,1845]" italics="true" pageId="53" pageNumber="54">points orange</emphasis>
&quot;,
<bibRefCitation id="FA164B3483681F4FF91BF894FC81F8C6" author="Coulon" box="[622,780,1820,1846]" pageId="53" pageNumber="54" refString="Coulon, L. (1864) Crustaces de la Mer Rouge. Bulletin de la Societe des Sciences Naturelles de Neuchatel, 6, 569 - 570." type="journal article" year="1864">Coulon 1864</bibRefCitation>
: 569) rather than the distinctly larger spots of
<taxonomicName id="59874D4683681F4FFA7DF8CCFE30F8AD" box="[264,445,1860,1885]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783681F4FFA7DF8CCFE30F8AD" box="[264,445,1860,1885]" italics="true" pageId="53" pageNumber="54">T. rufopunctata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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