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<document id="7361A8A444BA224BF4E53C4F30912EF0" ID-DOI="10.5252/g2012n3a10" ID-ISSN="1638-9395" ID-Zenodo-Dep="5375769" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630279646815" checkinUser="marcus" docAuthor="Ruiz-Sánchez, Francisco J., Murelaga, Xabier, Larrasoaña, Juan C., Freudenthal, Matthijs &amp; Garcés, Miguel" docDate="2012" docId="0383D54CFFC96D3CA941EA0FFED6FA4C" docLanguage="en" docName="Geodiversitas.2012.34.3.645-663.pdf" docOrigin="Geodiversitas 34 (3)" docSource="http://dx.doi.org/10.5252/g2012n3a10" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Armantomys daamsi" docType="treatment" docVersion="4" lastPageNumber="656" masterDocId="FFBAAD34FFC36D37AA6EEF22FFC9FFCB" masterDocTitle="Hypsodont Myomiminae (Gliridae, Rodentia) from five new localities in the Lower Miocene Tudela Formation (Bardenas Reales, Ebro Basin, Spain) and their bearing on the age of the Agenian-Ramblian boundary" masterLastPageNumber="663" masterPageNumber="645" pageNumber="655" updateTime="1699211341799" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="A9C1677B4646698B4FAA9D6980C57618">Hypsodont Myomiminae (Gliridae, Rodentia) from five new localities in the Lower Miocene Tudela Formation (Bardenas Reales, Ebro Basin, Spain) and their bearing on the age of the Agenian-Ramblian boundary</mods:title>
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<mods:namePart id="9DBE12D3412B3EFFD186611705899603">Ruiz-Sánchez, Francisco J.</mods:namePart>
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<treatment id="0383D54CFFC96D3CA941EA0FFED6FA4C" ID-DOI="http://doi.org/10.5281/zenodo.5464114" ID-Zenodo-Dep="5464114" LSID="urn:lsid:plazi:treatment:0383D54CFFC96D3CA941EA0FFED6FA4C" httpUri="http://treatment.plazi.org/id/0383D54CFFC96D3CA941EA0FFED6FA4C" lastPageId="11" lastPageNumber="656" pageId="10" pageNumber="655">
<subSubSection id="C33037D1FFC96D3DA941EA0FFBC2FA6D" pageId="10" pageNumber="655" type="nomenclature">
<paragraph id="8B95645AFFC96D3DA941EA0FFBC2FA6D" blockId="10.[693,1193,1324,1477]" pageId="10" pageNumber="655">
<heading id="D0DDD336FFC96D3DA941EA0FFBC2FA6D" centered="true" fontSize="11" level="2" pageId="10" pageNumber="655" reason="2">
<taxonomicName id="4C2A1FD9FFC96D3DA941EA0FFBE6FA8C" baseAuthorityName="de Visser" baseAuthorityYear="1991" box="[815,1071,1324,1351]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="655" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B95EB848FFC96D3DA941EA0FFC70FA8C" bold="true" box="[815,953,1325,1351]" italics="true" pageId="10" pageNumber="655">Armantomys</emphasis>
cf.
<emphasis id="B95EB848FFC96D3DA98FEA0EFBE6FA8C" bold="true" box="[993,1071,1324,1351]" italics="true" pageId="10" pageNumber="655">daamsi</emphasis>
</taxonomicName>
(de Visser
<emphasis id="B95EB848FFC96D3DA948EA6FFCF5FAAD" box="[806,828,1357,1382]" italics="true" pageId="10" pageNumber="655">in</emphasis>
Álvarez-Sierra, Daams, Lacomba, López Martínez, Van Der Meulen, Sesé &amp; De Visser, 1991)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33037D1FFC96D3DA900EA89FCDCF9F3" pageId="10" pageNumber="655" type="description">
<paragraph id="8B95645AFFC96D3DA900EA89FC39FA0E" blockId="10.[693,1193,1324,1477]" box="[878,1008,1451,1477]" pageId="10" pageNumber="655">
(
<figureCitation id="131178DFFFC96D3DA918EA89FC7CFA0E" box="[886,949,1451,1477]" captionStart="FIG" captionStartId="8.[113,124,1050,1067]" captionTargetBox="[139,1174,238,952]" captionTargetId="figure-247@8.[116,1212,214,1008]" captionTargetPageId="8" captionText="FIG. 4. — A-E, Armantomys cf. bijmai from Cabezo Hermoso 1 (CH1): A, left M2,CH1-6;B, right M3,CH1-8; C, right P4,CH1-4;D, right D4, CH1-5; E, right m1, CH1-3; F-M, Armantomys daamsi from Cuesta Agujeros 2 (CA2): F, left M1, CA2/1-6; G, right M2, CA2/1-5; H, right M2, CA2/2-6; I, right M3, CA2/1-15; J, right P4, CA2/1-12; K, left m1, CA2/1-8; L, left m1, CA2/1-10; M, right m3, CA2/2-5; N, Armantomys cf. jasperi from Cuesta Agujeros 4 (CA4), fragmentary right m2, CA4/1-27; O, P, Armantomys cf. parsani from Cuesta Agujeros 3 (CA3): O, left M2, CA3-1; P, left P4, CA3-2; Q, R, Armantomys cf. daamsi from Cabezo Carboneras 1 (CC1): Q, right M1, CC1-31; R, left M2, CC1-38; S, cf. Praearmantomys crusafonti from Cabezo Carboneras 1 (CC1), right m3, CC1-22. Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.5375777" httpUri="https://zenodo.org/record/5375777/files/figure.png" pageId="10" pageNumber="655">Fig. 4</figureCitation>
Q-R)
</paragraph>
<paragraph id="8B95645AFFC96D3DA8CBEAC9FCDCF9F3" blockId="10.[677,1211,1515,1592]" pageId="10" pageNumber="655">
LOCALITY AND AGE. — Cabezo Carboneras 1 (CC1), Province of
<collectingRegion id="49EEAAB8FFC96D3DA94CE924FCBDF9D6" box="[802,884,1542,1565]" country="Spain" name="Navarra" pageId="10" pageNumber="655">Navarre</collectingRegion>
,
<collectingCountry id="F33D24CAFFC96D3DA9EEE927FC75F9D6" box="[896,956,1541,1565]" name="Spain" pageId="10" pageNumber="655">Spain</collectingCountry>
. Agenian (Local subzone Y2, MN2).
</paragraph>
</subSubSection>
<subSubSection id="C33037D1FFC96D3DA8CBE975FC8AF968" pageId="10" pageNumber="655" type="materials_examined">
<paragraph id="8B95645AFFC96D3DA8CBE975FC8AF968" blockId="10.[677,1210,1622,1699]" pageId="10" pageNumber="655">
MATERIAL AND MEASUREMENTS (L
<emphasis id="B95EB848FFC96D3DAE6CE975FBDBF9A5" box="[1026,1042,1623,1646]" italics="true" pageId="10" pageNumber="655">×</emphasis>
W). —
<collectionCode id="ED3BFC9FFFC96D3DAE08E974FB47F9A5" box="[1126,1166,1622,1646]" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="10" pageNumber="655">CC</collectionCode>
1-31 (13.76 estimated
<emphasis id="B95EB848FFC96D3DA920E953FC97F943" box="[846,862,1649,1672]" italics="true" pageId="10" pageNumber="655">×</emphasis>
14.04), right M1;
<collectionCode id="ED3BFC9FFFC96D3DAE60E953FBFDF942" box="[1038,1076,1649,1673]" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="10" pageNumber="655">CC</collectionCode>
1-38 (11.71
<emphasis id="B95EB848FFC96D3DAEC4E953FB73F943" box="[1194,1210,1649,1672]" italics="true" pageId="10" pageNumber="655">×</emphasis>
15.80), left M2.
</paragraph>
</subSubSection>
<subSubSection id="C33037D1FFC86D3CAA1FEFFAFDC1FD26" pageId="11" pageNumber="656" type="description">
<paragraph id="8B95645AFFC86D3CAA1FEFFAFECBFF3B" blockId="11.[113,647,216,496]" box="[113,258,216,242]" pageId="11" pageNumber="656">DESCRIPTION</paragraph>
<paragraph id="8B95645AFFC86D3CAA1FEFDAFF51FED9" blockId="11.[113,647,216,496]" box="[113,152,248,274]" pageId="11" pageNumber="656">
<emphasis id="B95EB848FFC86D3CAA1FEFDAFF51FED9" box="[113,152,248,274]" italics="true" pageId="11" pageNumber="656">M1</emphasis>
</paragraph>
<paragraph id="8B95645AFFC86D3CAA1FEE35FE47FE3B" blockId="11.[113,647,216,496]" pageId="11" pageNumber="656">Anteroloph joined to paracone and separated from the protocone by a deep furrow. Protoloph oblique and continuous. Near the paracone the protoloph presents a short, forward directed, spur. Metaloph low connected to protoloph. Postcentroloph short. Posteroloph connected to protocone and metacone.</paragraph>
<paragraph id="8B95645AFFC86D3CAA1FED37FF51FDE4" blockId="11.[111,647,533,749]" box="[113,152,533,559]" pageId="11" pageNumber="656">
<emphasis id="B95EB848FFC86D3CAA1FED37FF51FDE4" box="[113,152,533,559]" italics="true" pageId="11" pageNumber="656">M2</emphasis>
</paragraph>
<paragraph id="8B95645AFFC86D3CAA1FED16FDC1FD26" blockId="11.[111,647,533,749]" pageId="11" pageNumber="656">Anteroloph slightly oblique, connected to the protocone and separated from the paracone by a deep furrow. Protoloph continuous without spurs. Metaloph connected to protoloph, forming a Y-pattern. No centroloph ridges. Posteroloph slightly oblique and high connected to the protocone.</paragraph>
</subSubSection>
<subSubSection id="C33037D1FFC86D3CAA1FEC31FED6FA4C" pageId="11" pageNumber="656" type="discussion">
<paragraph id="8B95645AFFC86D3CAA1FEC31FF1CFCE0" blockId="11.[111,647,787,1415]" box="[113,213,787,813]" pageId="11" pageNumber="656">REMARKS</paragraph>
<paragraph id="8B95645AFFC86D3CAA1FEC10FED6FA4C" blockId="11.[111,647,787,1415]" pageId="11" pageNumber="656">
Two specimens from Cabezo Carboneras 1 (CC1) (
<figureCitation id="131178DFFFC86D3CAA17EC73FF3AFCA7" box="[121,243,849,876]" captionStart="FIG" captionStartId="8.[113,124,1050,1067]" captionTargetBox="[139,1174,238,952]" captionTargetId="figure-247@8.[116,1212,214,1008]" captionTargetPageId="8" captionText="FIG. 4. — A-E, Armantomys cf. bijmai from Cabezo Hermoso 1 (CH1): A, left M2,CH1-6;B, right M3,CH1-8; C, right P4,CH1-4;D, right D4, CH1-5; E, right m1, CH1-3; F-M, Armantomys daamsi from Cuesta Agujeros 2 (CA2): F, left M1, CA2/1-6; G, right M2, CA2/1-5; H, right M2, CA2/2-6; I, right M3, CA2/1-15; J, right P4, CA2/1-12; K, left m1, CA2/1-8; L, left m1, CA2/1-10; M, right m3, CA2/2-5; N, Armantomys cf. jasperi from Cuesta Agujeros 4 (CA4), fragmentary right m2, CA4/1-27; O, P, Armantomys cf. parsani from Cuesta Agujeros 3 (CA3): O, left M2, CA3-1; P, left P4, CA3-2; Q, R, Armantomys cf. daamsi from Cabezo Carboneras 1 (CC1): Q, right M1, CC1-31; R, left M2, CC1-38; S, cf. Praearmantomys crusafonti from Cabezo Carboneras 1 (CC1), right m3, CC1-22. Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.5375777" httpUri="https://zenodo.org/record/5375777/files/figure.png" pageId="11" pageNumber="656">Fig. 4Q, R</figureCitation>
) have been assigned to
<taxonomicName id="4C2A1FD9FFC86D3CAB94EC70FD4EFCA0" baseAuthorityName="de Visser" baseAuthorityYear="1991" box="[506,647,849,876]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B95EB848FFC86D3CAB94EC70FDDBFCA7" box="[506,530,850,876]" italics="true" pageId="11" pageNumber="656">A.</emphasis>
cf.
<emphasis id="B95EB848FFC86D3CA853EC73FD4EFCA0" box="[573,647,849,875]" italics="true" pageId="11" pageNumber="656">daamsi</emphasis>
</taxonomicName>
on the basis of size, the presence of a posterior centroloph in M1 and the kind of connections between metaloph and protoloph in M1,2. The presence of an anterior and/or posterior centroloph is dominant in populations of
<taxonomicName id="4C2A1FD9FFC86D3CAB48ECD2FE6FFBC1" authorityName="de Bruijn" authorityYear="1966" box="[294,422,1008,1034]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="genus">
<emphasis id="B95EB848FFC86D3CAB48ECD2FE6FFBC1" box="[294,422,1008,1034]" italics="true" pageId="11" pageNumber="656">Armantomys</emphasis>
</taxonomicName>
from the Upper Agenian and Lower Ramblian (
<bibRefCitation id="EFBB19ABFFC86D3CABF9EB32FDEEFBE1" author="DAAMS R." box="[407,551,1040,1066]" pageId="11" pageNumber="656" pagination="1 - 62" refId="ref9936" refString="DAAMS R. 1990. - Hypsodont Myomiminae (Gliridae, Rodentia) in the Oligocene-Miocene boundary interval and Miocene of Spain. Scripta Geologica 96: 1 - 62." type="journal article" year="1990">Daams 1990</bibRefCitation>
). On the other hand, in CC1-31 (
<figureCitation id="131178DFFFC86D3CAB14EB0DFE18FB82" box="[378,465,1071,1097]" captionStart="FIG" captionStartId="8.[113,124,1050,1067]" captionTargetBox="[139,1174,238,952]" captionTargetId="figure-247@8.[116,1212,214,1008]" captionTargetPageId="8" captionText="FIG. 4. — A-E, Armantomys cf. bijmai from Cabezo Hermoso 1 (CH1): A, left M2,CH1-6;B, right M3,CH1-8; C, right P4,CH1-4;D, right D4, CH1-5; E, right m1, CH1-3; F-M, Armantomys daamsi from Cuesta Agujeros 2 (CA2): F, left M1, CA2/1-6; G, right M2, CA2/1-5; H, right M2, CA2/2-6; I, right M3, CA2/1-15; J, right P4, CA2/1-12; K, left m1, CA2/1-8; L, left m1, CA2/1-10; M, right m3, CA2/2-5; N, Armantomys cf. jasperi from Cuesta Agujeros 4 (CA4), fragmentary right m2, CA4/1-27; O, P, Armantomys cf. parsani from Cuesta Agujeros 3 (CA3): O, left M2, CA3-1; P, left P4, CA3-2; Q, R, Armantomys cf. daamsi from Cabezo Carboneras 1 (CC1): Q, right M1, CC1-31; R, left M2, CC1-38; S, cf. Praearmantomys crusafonti from Cabezo Carboneras 1 (CC1), right m3, CC1-22. Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.5375777" httpUri="https://zenodo.org/record/5375777/files/figure.png" pageId="11" pageNumber="656">Fig. 4Q</figureCitation>
), a short spur on the anterior side of the protoloph is present. This feature appears in populations of
<taxonomicName id="4C2A1FD9FFC86D3CABA2EB4DFD84FB42" authorityName="de Bruijn" authorityYear="1966" box="[460,589,1135,1161]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="genus">
<emphasis id="B95EB848FFC86D3CABA2EB4DFD84FB42" box="[460,589,1135,1161]" italics="true" pageId="11" pageNumber="656">Armantomys</emphasis>
</taxonomicName>
from Upper Agenian to Middle Aragonian. The presence of spurs on the protoloph of M1,2 is frequent in
<taxonomicName id="4C2A1FD9FFC86D3CAA1DEBECFF1EFB2C" authorityName="Daams" authorityYear="1990" box="[115,215,1230,1256]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="species" species="parsani">
<emphasis id="B95EB848FFC86D3CAA1DEBECFF1EFB2C" box="[115,215,1230,1256]" italics="true" pageId="11" pageNumber="656">A. parsani</emphasis>
</taxonomicName>
and dominant in
<taxonomicName id="4C2A1FD9FFC86D3CABE5EBECFE2CFB2C" baseAuthorityName="Daams" baseAuthorityYear="1990" box="[395,485,1230,1256]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="species" species="jasperi">
<emphasis id="B95EB848FFC86D3CABE5EBECFE2CFB2C" box="[395,485,1230,1256]" italics="true" pageId="11" pageNumber="656">A. jasperi</emphasis>
</taxonomicName>
from the Lower Ramblian; it has been cited in some specimens of
<taxonomicName id="4C2A1FD9FFC86D3CAA1CEA2FFF1EFAEC" baseAuthorityName="de Visser" baseAuthorityYear="1991" box="[114,215,1293,1319]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B95EB848FFC86D3CAA1CEA2FFF1EFAEC" box="[114,215,1293,1319]" italics="true" pageId="11" pageNumber="656">A. daamsi</emphasis>
</taxonomicName>
from the Upper Agenian (
<bibRefCitation id="EFBB19ABFFC86D3CAB83EA2FFDB2FAEC" author="DAAMS R." box="[493,635,1293,1319]" pageId="11" pageNumber="656" pagination="1 - 62" refId="ref9936" refString="DAAMS R. 1990. - Hypsodont Myomiminae (Gliridae, Rodentia) in the Oligocene-Miocene boundary interval and Miocene of Spain. Scripta Geologica 96: 1 - 62." type="journal article" year="1990">Daams 1990</bibRefCitation>
). Therefore, on the basis of morphology, size and the scarcity of the material we assign these specimens to
<taxonomicName id="4C2A1FD9FFC86D3CAAE1EA4EFED1FA4D" baseAuthorityName="de Visser" baseAuthorityYear="1991" box="[143,280,1388,1414]" class="Mammalia" family="Gliridae" genus="Armantomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="656" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B95EB848FFC86D3CAAE1EA4EFF6EFA4D" box="[143,167,1388,1414]" italics="true" pageId="11" pageNumber="656">A.</emphasis>
cf.
<emphasis id="B95EB848FFC86D3CAAA0EA4EFED1FA4D" box="[206,280,1388,1414]" italics="true" pageId="11" pageNumber="656">daamsi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>