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<mods:title id="C96E40CDCEA7844F0D2D3F4F0B5CDF93">Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae)</mods:title>
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Genus 
<taxonomicName id="4C124D1CFFBCFFAA2664D215FD24FFCE" ID-CoL="5ZKC" authority="Lourenco, 1995" authorityName="Lourenco" authorityYear="1995" box="[309,662,876,904]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA2664D215FE67FFC1" bold="true" box="[309,469,876,903]" italics="true" pageId="13" pageNumber="12">Neogrosphus</emphasis>
<bibRefCitation id="EF834B6EFFBCFFAA268DD214FD24FFCE" author="LOURENCO, W. R." box="[476,662,877,904]" journalOrPublisher="Bulletin du Museum National d'Histoire naturelle, Paris (Zoologie, Biologie et Ecologie Animale)" pageId="13" pageNumber="12" pagination="95 - 106" part="4" refId="ref37469" refString="LOURENCO, W. R. 1995 a. Description de trois noveaux genres et quatre nouvelles especes de scorpions Buthidae de Madagascar. Bulletin du Museum National d'Histoire naturelle, Paris (Zoologie, Biologie et Ecologie Animale), (4), 17 A (1 - 2): 95 - 106." title="Description de trois noveaux genres et quatre nouvelles especes de scorpions Buthidae de Madagascar" type="journal article" year="1995">Lourenço, 1995</bibRefCitation>
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<paragraph id="8BAD369FFFBCFFAA27C2D2F7FD56FFE1" blockId="13.[147,740,876,935]" box="[147,740,910,935]" pageId="13" pageNumber="12">
(
<figureCitation id="13292A1AFFBCFFAA27CAD2F7FEAFFFE0" box="[155,285,910,934]" captionStart-0="Figures 21–27" captionStart-1="Figures 28–29" captionStart-2="Figures 30–35" captionStart-3="Figures 36–39" captionStartId-0="14.[113,188,1859,1881]" captionStartId-1="18.[113,190,1953,1975]" captionStartId-2="19.[113,190,1773,1795]" captionStartId-3="20.[113,190,1388,1410]" captionTargetBox-0="[114,1474,227,1840]" captionTargetBox-2="[118,1455,257,1710]" captionTargetBox-3="[121,1471,246,1361]" captionTargetId-0="figure-206@14.[113,1474,222,1851]" captionTargetId-1="figure-131@18.[113,1474,255,1915]" captionTargetId-2="figure-204@19.[113,1474,251,1714]" captionTargetId-3="figure-425@20.[113,1474,234,1383]" captionTargetPageId-0="14" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Figures 21–27. Positions of trichobothria Eb, Eb and petite Eb on manus of pedipalp chela in Grosphus, Neogrosphus and Teruelius gen. 1 2 3 n. Figures 21–24. External views of pedipalp chela, shown under UV fluorescence to highlight trichobothrial areolae. G. hirtus ♂ (21), G. madagascariensis ♀ (22), T. limbatus ♂ (23), T. ankarafantsika ♀ (24). Positions of Eb 1, Eb 2 and Eb 3 and lines joining them shown as white overlays. Scale bars: 2 mm. Figures 25–26. Horizontal histograms comparing ratios, R = d(Eb, Eb)/ d(Eb, Eb), of Eb -Eb distance, to Eb - 123 2 3 1 2 2 3 1 Eb distance, in males (25), and females (26) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), 2 histogram bars mid-range values. Figure 27. Scatter plot of male vs. female ratios R. Ratio is larger in males if points fall above the diagonal 123 (gray) line, larger in females if they fall below it." captionText-1="Figures 28–29. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Horizontal histograms comparing PTC in males (28), and females (29) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), bars mid-range values." captionText-2="Figures 30–35. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Figures 30–31. Logarithmic scatter plots showing scaling of PTC vs. carapace length in males (30), and females (31) of Grosphus, Neogrosphus and Teruelius gen. n. Color lines are least squares linear regression fits of scaling trends for the different genera. Figures 32–33. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (32) and female (33) buthids including both α and β trichobothrial configurations (gray symbols; 1055♂, 1046♀). Dark lines: least squares linear regression fits of scaling trends for larger sample of buthids. Figures 34–35. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (34) and female (35) α-buthids restricted to α trichobothrial configurations (gray symbols; 445♂, 371♀). Dark lines: least squares linear regression fits of scaling trends for α-buthids. Numbers: slopes of regression lines (= allometric exponents). Purple symbols and lines: outgroup Pseudolychas." captionText-3="Figures 36–39. Comparative biometrics of female basal pectinal tooth and hemispermatophore trunk of Grosphus, Neogrosphus and Teruelius gen. n. Figures 36–37. Logarithmic scatter plots showing scaling of length of female basal pectinal tooth vs. carapace length (36), and hemispermatophore trunk length vs. carapace length (37). Color lines: least squares linear regression fits of scaling trends for the different genera. Figures 38–39. Logarithmic scatter plots of female basal pectinal tooth length vs. hemispermatophore trunk length (38), and the ratio of basal pectinal tooth length/ hemispermatophore trunk length (%) vs. hemispermatophore trunk length, showing fraction of trunk spanned by the female tooth (39). Female basal pectinal tooth length measured as chord length, from more proximal corner (along comb axis) at base of tooth, to more distal corner (along comb axis) at tip of tooth. Trunk length measured from proximal edge of basal lobe to pedicel (foot). Color lines: least squares linear regression fits of scaling trends for the different genera." figureDoi-0="http://doi.org/10.5281/zenodo.7143620" figureDoi-1="http://doi.org/10.5281/zenodo.7143622" figureDoi-2="http://doi.org/10.5281/zenodo.7143626" figureDoi-3="http://doi.org/10.5281/zenodo.7143628" httpUri-0="https://zenodo.org/record/7143620/files/figure.png" httpUri-1="https://zenodo.org/record/7143622/files/figure.png" httpUri-2="https://zenodo.org/record/7143626/files/figure.png" httpUri-3="https://zenodo.org/record/7143628/files/figure.png" pageId="13" pageNumber="12">Figs. 25–39</figureCitation>
, 
<figureCitation id="13292A1AFFBCFFAA2679D2F7FEDFFFE0" box="[296,365,910,934]" captionStart="Figures 52–70" captionStartId="24.[113,190,1591,1613]" captionTargetBox="[116,1470,248,1526]" captionTargetId="figure-361@24.[113,1474,248,1527]" captionTargetPageId="24" captionText="Figures 52–70. Hemispermatophores and capsule regions of Grosphus and Neogrosphus. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and capsule with flagellum; capsule region in convex (and/ or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores unless indicated as mirrored left images. Figure 52. G. madagascariensis, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm). Figure 53. G. madagascariensis, capsule, Sc1197, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 54. G. madagascariensis, capsule, Anjiro, G. halleuxi nr topotype, GLPC. Scale bar: 500 μm. Figure 55. G. madagascariensis, capsule, Mandena- Fort Dauphin, G. mandena paratype, MHNG.Scale bar: 500 μm.Figure 56. G. madagascariensis, capsule, Madagascar, det. Vachon, MHNG. Scale bar: 500 μm. Figure 57. G. madagascariensis, capsule, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 58. G. goudoti, whole hemispermatophore. Scale bar: 2 mm. Figure 59. G. goudoti, capsule, Forêt de Bobankota, holotype, MHNG. Scale bar: 500 μm. Figure 60. G. hirtus, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm), Antsiranana, Ramena vill., mirrored left, GLPC, FKCP. Figure 61. G. hirtus, capsule, Mahajamba River, GLPC, FKCP. Scale bar: 500 μm. Figures 62–65. G. hirtus, capsules in convex view.Antsiranana, Ramena vill., mirrored left, GLPC, FKCP (62), Jardin Botanique, MHNG (63), Ranohira-Llakaka, ZMUH, mirrored left (64), Forêt de Vohitaly, MHNG (65). Scale bars: 500 μm. Figure 66. G. hirtus, capsule, Forest Station Ampijoroa, G. garciai holotype, MHNG. Scale bar: 500 μm. Figure 67. G. voahangyae, whole hemispermatophore. Scale bar: 2 mm. Figure 68. G. voahangyae, capsule, Analamy Forest, FMNH. Scale bar: 500 μm. Figure 69. N. griveaudi, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 70. N. griveaudi, capsule, mirrored left, Tsimanampetsotsa National Park, GLPC, FKCP." figureDoi="http://doi.org/10.5281/zenodo.7143636" httpUri="https://zenodo.org/record/7143636/files/figure.png" pageId="13" pageNumber="12">69–70</figureCitation>
, 
<figureCitation id="13292A1AFFBCFFAA2629D2F6FE27FFE1" box="[376,405,911,935]" captionStart="Figures 86–89" captionStartId="27.[114,191,1768,1790]" captionTargetBox="[122,1468,260,1754]" captionTargetId="figure-170@27.[113,1474,234,1762]" captionTargetPageId="27" captionText="Figures 86–89. Hemispermatophores and capsule regions of Grosphus and Neogrosphus. Cross stereoscopic convex views. Scale bars: 500 μm. Figure 86. G. madagascariensis, Sc1197, Andasibe, GLPC, FKCP. Figure 87. N. griveaudi, mirrored left, Tsimanampetsotsa National Park, GLPC, FKCP. Figure 88. G. goudoti, Forêt de Bobankota, holotype, MHNG. Figure 89. G. voahangyae, Analamy Forest, FMNH." figureDoi="http://doi.org/10.5281/zenodo.7143643" httpUri="https://zenodo.org/record/7143643/files/figure.png" pageId="13" pageNumber="12">87</figureCitation>
, 
<figureCitation id="13292A1AFFBCFFAA26F1D2F7FE4DFFE0" box="[416,511,910,934]" captionStart-0="Figures 95–110" captionStart-1="Figures 110–113" captionStart-2="Figures 114–116" captionStart-3="Figures 117–120" captionStart-4="Figure 121" captionStartId-0="18.[113,190,1408,1430]" captionStartId-1="32.[113,190,1684,1706]" captionStartId-2="33.[113,190,1610,1632]" captionStartId-3="35.[113,190,1782,1804]" captionStartId-4="36.[113,181,1915,1937]" captionTargetBox-0="[113,1475,255,1894]" captionTargetBox-1="[113,1474,276,1623]" captionTargetBox-2="[128,1470,242,1585]" captionTargetBox-3="[132,1464,278,1712]" captionTargetBox-4="[120,1469,286,1859]" captionTargetId-0="figure-131@18.[113,1474,255,1915]" captionTargetId-1="figure-16@32.[113,1474,266,1629]" captionTargetId-2="figure-262@33.[113,1474,234,1604]" captionTargetId-3="figure-194@35.[113,1474,274,1732]" captionTargetId-4="figure-94@36.[113,1474,282,1865]" captionTargetPageId-0="18" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="35" captionTargetPageId-4="36" captionText-0="Figures 95–110: Orthochirus mesopotamicus comb. n., from Iraq, Basra (ca. 30°48’N 47°34’E). Figures 95, 97, 100–110. Male, carapace and tergites I–III (95), sternopectinal region and sternites (97). Pedipalp chela, dorsal (100), external (101), and ventrointernal (102) views. Pedipalp patella, dorsal (103), external (104), and ventral (105) views. Pedipalp femur and trochanter, internodorsal (106) and dorsoexternal (107) views. Pedipalp chela, movable finger dentate margin (108). Distal segments of right legs III–IV, retrolateral views (109–110). Figures 96, 98–99. Female, carapace and tergites I–III (96), sternopectinal region and sternites (98), and pedipalp chela dorsoexternal (99). The trichobothrial pattern is indicated in Figures 101–104, 106–107 (white circles)." captionText-1="Figures 110–113. Biometric scaling of spiracles of sternite IV in Grosphus, Neogrosphus and Teruelius gen. n. Figures 110–111. Logarithmic scatter plots showing scaling of single spiracle area/ (carapace length)2 vs. carapace length in males (110), and females (111) of Grosphus, Neogrosphus and Teruelius gen. n. Spiracle length was largest diameter (approximately along mediolateral axis), width was smallest diameter (approximately along rostrocaudal axis). Area was from closed boundary curve: anterior edge of opening sharply delineated, posterior edge sometimes less sharp and fitted with spline curve extrapolating lateral edge curvature and tracing shadow of raised posterior margin. Brown line is least squares linear regression fit of scaling trends for Teruelius. R = Pearson correlation coefficient, and P value as indicated. Plotted points are means, error bars SE. Figures 112–113. Logarithmic scatter plots showing thermal (terrestrial temperature) scaling of spiracle area/ (carapace length)2 in males (112), and females (113) of Grosphus, Neogrosphus and Teruelius gen. n. Gray lines are least squares linear regression fits of scaling trends for all points. R = Pearson correlation coefficient, P value as indicated. Plotted points are means, error bars SE. Land surface temperatures at known collection sites were averaged over all sites for each species. Collection sites were extracted from published data and FMNH records. Temperatures were extracted from published model data of Chabot-Couture et al. (2014), who estimated annual mean daytime temperature by statistical processing and Kriging of remote sensing spectral radiance by AQUA satellite." captionText-2="Figures 114–116. Biometrics of spiracles of sternite IV in Grosphus, Neogrosphus and Teruelius gen. n. Logarithmic scatter plots showing sexual dimorphism of spiracle area/ (carapace length)2 (114), spiracle width/ carapace length (115), and spiracle length/ width (116) in Grosphus, Neogrosphus and Teruelius gen. n. Plotted points are means, error bars SE. Ratios are larger in males if points fall above the diagonal (gray) line, larger in females if they fall below it." captionText-3="Figures 117–120. Distributions of Grosphus, Neogrosphus and Teruelius gen. n. superposed on climate maps. Figure 117. Collection sites superposed on average rainfall map. Figure 118. Collection sites superposed on land temperature map. Figure 119. Distribution of each genus according to average rainfall. Figure 120. Distribution of each genus according to land temperature. Vertical histogram bars in Figs 119–120 are means, error bars SE. P-values and asterisks indicate significant differences of Neogrosphus and Teruelius gen. n. means from Grosphus means. Symbols in bar charts are collection sites collapsed along rainfall axis (119), or temperature axis (120). Inset horizontal histograms show relative (normalized) densities of points vs. rainfall or temperature (vertical axes). Collection sites were extracted from published data and FMNH records. Rainfall and temperatures were extracted from published model data of Chabot-Couture et al. (2014), who estimated January and July rainfalls by interpolation of RFE 2.0 data, and annual mean daytime temperature by statistical processing and Kriging of remote sensing spectral radiance by AQUA satellite. We averaged January and July rainfalls to generate a single map." captionText-4="Figure 121. Bioclimatic zone distributions of Grosphus, Neogrosphus and Teruelius gen. n. in Madagascar. Collection sites were extracted from published data and FMNH records, and their bioclimatic classification visualized by superposition on zone boundaries defined by Cornet (1974). Colored regions show zones as delineated by Cornet, and symbols for genera are placed on the map and on zone keys of Cornet (upper left inset) to show ranges of humidity and temperature tolerance of each genus." figureDoi-1="http://doi.org/10.5281/zenodo.7143657" figureDoi-2="http://doi.org/10.5281/zenodo.7143663" figureDoi-3="http://doi.org/10.5281/zenodo.7143665" figureDoi-4="http://doi.org/10.5281/zenodo.7143669" httpUri-1="https://zenodo.org/record/7143657/files/figure.png" httpUri-2="https://zenodo.org/record/7143663/files/figure.png" httpUri-3="https://zenodo.org/record/7143665/files/figure.png" httpUri-4="https://zenodo.org/record/7143669/files/figure.png" pageId="13" pageNumber="12">106–121</figureCitation>
, 
<figureCitation id="13292A1AFFBCFFAA255BD2F7FDD9FFE0" box="[522,619,910,934]" captionStart-0="Figures 606–609" captionStart-1="Figures 610–619" captionStartId-0="125.[113,190,1988,2010]" captionStartId-1="126.[113,190,1939,1961]" captionTargetBox-0="[123,1452,237,1933]" captionTargetBox-1="[119,1437,234,1911]" captionTargetId-0="figure-27@125.[113,1474,234,1975]" captionTargetId-1="figure-79@126.[119,1452,234,1925]" captionTargetPageId-0="125" captionTargetPageId-1="126" captionText-0="Figures 606–609. Neogrosphus griveaudi. Habitus. Male (606–607) and female (608–609), dorsal (606, 608) and ventral (607, 609) views. Scale bar: 10 mm." captionText-1="Figures 610–619. Neogrosphus griveaudi, male (610–613, 617–619) and female (614–616). Figures 610, 614. Sternopectinal region. Figures 611, 615. Carapace and tergites I–III. Figure 612. Pedipalp chela, movable finger dentate margin. Figures 613, 616. Pedipalp chela in dorsal views. Figure 617. Metasoma V and telson lateral view. Figures 618–619. Right leg III (618) and leg IV (619), tibia, basitarsus and telotarsus, retrolateral views." figureDoi-0="http://doi.org/10.5281/zenodo.7143861" figureDoi-1="http://doi.org/10.5281/zenodo.7143863" httpUri-0="https://zenodo.org/record/7143861/files/figure.png" httpUri-1="https://zenodo.org/record/7143863/files/figure.png" pageId="13" pageNumber="12">606–619</figureCitation>
, 
<tableCitation id="C6900324FFBCFFAA2524D2F7FD6EFFE1" box="[629,732,910,935]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="9.[115,171,1146,1167]" captionStartId-1="10.[113,169,1291,1312]" captionStartId-2="16.[110,166,1277,1298]" captionStartId-3="17.[113,169,1270,1291]" captionTargetId-0="graphics-417@9.[119,1468,237,1129]" captionTargetId-1="graphics-433@10.[121,1460,237,1263]" captionTargetId-2="graphics-479@16.[118,1471,242,1254]" captionTargetId-3="graphics-446@17.[119,1470,248,1242]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="Table 1. Character states for species examined. States of 5 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of trichobothrium d 2 on pedipalp femur; ratio R123 = d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀); and size and shape of female basal pectinal tooth." captionText-1="Table 2. Character states for species examined. States of 6 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of hemispermatophore basal lobe relative to base of flagellum; size of hemispermatophore posterior lobe; shape of spiracles; condition of metasoma I ventromedian carinae; telotarsal setation; and intensity of UV fluorescence." captionText-2="Table 3. Character states for species not examined. States of 6 characters for Grosphus (6 spp., blue panels), Neogrosphus (2 spp., magenta panels) and Teruelius gen. n. (7 spp., yellow panels) not examined in this study: position of trichobothrium d on pedipalp femur; ratio R = 2 123 d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀);size and shape of female basal pectinal tooth; shape of spiracles; and telotarsal setation. Character states inferred from published descriptions and figures. PTC and basal pectinal tooth characters not indicated in cases where either males or females are not known or described. Other characters not indicated in cases where published descriptions did not yield clear information." captionText-3="Table 4. Character states for ‘Grosphus’ group genera and outgroup genus Pseudolychas. States of 11 characters for Grosphus, Neogrosphus, Teruelius gen. n. and Pseudolychas, summarizing data of Tables 1–3. Pseudolychas character states determined by study of materials, and from published descriptions. Primitive states indicated by green panels, derived states by red panels, with polarities determined by comparison to Pseudolychas which is assumed primitive. Unpolarized characters indicated by dark green panels." httpUri-0="http://table.plazi.org/id/DF6D6617FFB8FFAE2722D503FCADF8A1" httpUri-2="http://table.plazi.org/id/DF6D6617FFA1FFB7273FD584FB00F9E5" httpUri-3="http://table.plazi.org/id/DF6D6617FFA0FFB62720D58FFB7EF924" pageId="13" pageNumber="12" tableUuid-0="DF6D6617FFB8FFAE2722D503FCADF8A1" tableUuid-2="DF6D6617FFA1FFB7273FD584FB00F9E5" tableUuid-3="DF6D6617FFA0FFB62720D58FFB7EF924">Tabs. 1–4</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3086514FFBCFFAA2722D2B7FECCF881" pageId="13" pageNumber="12" type="reference_group">
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<emphasis id="B966EA8DFFBCFFAA2722D2B7FF4DFFA0" box="[115,255,974,998]" italics="true" pageId="13" pageNumber="12">Neogrosphus</emphasis>
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, fig. 2, 6; 
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</paragraph>
</subSubSection>
<subSubSection id="C3086514FFBCFFAA2722D596FD7DFB00" pageId="13" pageNumber="12" type="diagnosis">
<paragraph id="8BAD369FFFBCFFAA2722D596FD7DFB00" blockId="13.[115,773,1262,1863]" pageId="13" pageNumber="12">
DIAGNOSIS. A member of the ‘ 
<taxonomicName id="4C124D1CFFBCFFAA26EAD597FD90F940" authorityName="Simon" authorityYear="1880" box="[443,546,1262,1286]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA26EAD597FD90F940" box="[443,546,1262,1286]" italics="true" pageId="13" pageNumber="12">Grosphus</emphasis>
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’ group differentiated as follows: small-sized scorpions, adults ca. 
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in length; pedipalp finger granule rows 8–9 (
<figureCitation id="13292A1AFFBCFFAA2537D457FF2DF920" captionStart="Figures 610–619" captionStartId="126.[113,190,1939,1961]" captionTargetBox="[119,1437,234,1911]" captionTargetId="figure-79@126.[119,1452,234,1925]" captionTargetPageId="126" captionText="Figures 610–619. Neogrosphus griveaudi, male (610–613, 617–619) and female (614–616). Figures 610, 614. Sternopectinal region. Figures 611, 615. Carapace and tergites I–III. Figure 612. Pedipalp chela, movable finger dentate margin. Figures 613, 616. Pedipalp chela in dorsal views. Figure 617. Metasoma V and telson lateral view. Figures 618–619. Right leg III (618) and leg IV (619), tibia, basitarsus and telotarsus, retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.7143863" httpUri="https://zenodo.org/record/7143863/files/figure.png" pageId="13" pageNumber="12">Fig. 612–613, 616</figureCitation>
), movable finger with no more than 3 external subdistal granules; femur trichobothrium 
<emphasis id="B966EA8DFFBCFFAA2697D417FE61F9C0" box="[454,467,1390,1414]" italics="true" pageId="13" pageNumber="12">d</emphasis>
2 located on dorsal, carinal or internal surface; chela manus with petite trichobothrium 
<emphasis id="B966EA8DFFBCFFAA25B1D4F7FD4CF9E0" box="[736,766,1422,1446]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
3 usually well separated from 
<emphasis id="B966EA8DFFBCFFAA26CBD4D7FE0AF980" box="[410,440,1454,1478]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
2, by more than half the distance between 
<emphasis id="B966EA8DFFBCFFAA2782D4B7FF43F9A0" box="[211,241,1486,1510]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
<subScript id="179634DAFFBCFFAA27A0D4A6FF4BF9AB" attach="left" box="[241,249,1503,1517]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
and 
<emphasis id="B966EA8DFFBCFFAA267FD4B7FEFEF9A0" box="[302,332,1486,1510]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
<subScript id="179634DAFFBCFFAA261AD4A6FEE1F9AB" attach="left" box="[331,339,1503,1517]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
(
<figureCitation id="13292A1AFFBCFFAA2633D4B7FE54F9A1" box="[354,486,1486,1511]" captionStart="Figures 21–27" captionStartId="14.[113,188,1859,1881]" captionTargetBox="[114,1474,227,1840]" captionTargetId="figure-206@14.[113,1474,222,1851]" captionTargetPageId="14" captionText="Figures 21–27. Positions of trichobothria Eb, Eb and petite Eb on manus of pedipalp chela in Grosphus, Neogrosphus and Teruelius gen. 1 2 3 n. Figures 21–24. External views of pedipalp chela, shown under UV fluorescence to highlight trichobothrial areolae. G. hirtus ♂ (21), G. madagascariensis ♀ (22), T. limbatus ♂ (23), T. ankarafantsika ♀ (24). Positions of Eb 1, Eb 2 and Eb 3 and lines joining them shown as white overlays. Scale bars: 2 mm. Figures 25–26. Horizontal histograms comparing ratios, R = d(Eb, Eb)/ d(Eb, Eb), of Eb -Eb distance, to Eb - 123 2 3 1 2 2 3 1 Eb distance, in males (25), and females (26) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), 2 histogram bars mid-range values. Figure 27. Scatter plot of male vs. female ratios R. Ratio is larger in males if points fall above the diagonal 123 (gray) line, larger in females if they fall below it." figureDoi="http://doi.org/10.5281/zenodo.7143620" httpUri="https://zenodo.org/record/7143620/files/figure.png" pageId="13" pageNumber="12">Figs. 25–27</figureCitation>
); manus trichobothrium 
<specimenCode id="DBB49EE4FFBCFFAA25BCD4B6FCB7F9AB" box="[749,773,1487,1517]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="12" type="Museum">
<emphasis id="B966EA8DFFBCFFAA25BCD4B6FD4FF9A0" box="[749,765,1487,1510]" italics="true" pageId="13" pageNumber="12">V</emphasis>
<subScript id="179634DAFFBCFFAA25ACD4A6FCB7F9AB" attach="left" box="[765,773,1503,1517]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
</specimenCode>
strongly displaced internally relative to 
<specimenCode id="DBB49EE4FFBCFFAA2564D496FDFCFA4B" box="[565,590,1519,1549]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="12" type="Museum">
<emphasis id="B966EA8DFFBCFFAA2564D496FDF7FA40" box="[565,581,1519,1542]" italics="true" pageId="13" pageNumber="12">V</emphasis>
1
</specimenCode>
; higher pectinal tooth counts: 
<specimenCount id="9D14FD16FFBCFFAA2655D776FEAAFA60" box="[260,280,1551,1574]" pageId="13" pageNumber="12" type="male">♂</specimenCount>
27– 31, 
<specimenCount id="9D14FD16FFBCFFAA2625D776FE3AFA60" box="[372,392,1551,1574]" pageId="13" pageNumber="12" type="female">♀</specimenCount>
27–29 (
<figureCitation id="13292A1AFFBCFFAA268ED777FDD2FA61" box="[479,608,1550,1575]" captionStart-0="Figures 28–29" captionStart-1="Figures 30–35" captionStartId-0="18.[113,190,1953,1975]" captionStartId-1="19.[113,190,1773,1795]" captionTargetBox-1="[118,1455,257,1710]" captionTargetId-0="figure-131@18.[113,1474,255,1915]" captionTargetId-1="figure-204@19.[113,1474,251,1714]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Figures 28–29. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Horizontal histograms comparing PTC in males (28), and females (29) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), bars mid-range values." captionText-1="Figures 30–35. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Figures 30–31. Logarithmic scatter plots showing scaling of PTC vs. carapace length in males (30), and females (31) of Grosphus, Neogrosphus and Teruelius gen. n. Color lines are least squares linear regression fits of scaling trends for the different genera. Figures 32–33. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (32) and female (33) buthids including both α and β trichobothrial configurations (gray symbols; 1055♂, 1046♀). Dark lines: least squares linear regression fits of scaling trends for larger sample of buthids. Figures 34–35. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (34) and female (35) α-buthids restricted to α trichobothrial configurations (gray symbols; 445♂, 371♀). Dark lines: least squares linear regression fits of scaling trends for α-buthids. Numbers: slopes of regression lines (= allometric exponents). Purple symbols and lines: outgroup Pseudolychas." figureDoi-0="http://doi.org/10.5281/zenodo.7143622" figureDoi-1="http://doi.org/10.5281/zenodo.7143626" httpUri-0="https://zenodo.org/record/7143622/files/figure.png" httpUri-1="https://zenodo.org/record/7143626/files/figure.png" pageId="13" pageNumber="12">Figs. 28–31</figureCitation>
); basal pectinal tooth of females wide, oval, only slightly longer than other teeth (
<figureCitation id="13292A1AFFBCFFAA27E9D737FEA5FA20" box="[184,279,1614,1638]" captionStart="Figures 610–619" captionStartId="126.[113,190,1939,1961]" captionTargetBox="[119,1437,234,1911]" captionTargetId="figure-79@126.[119,1452,234,1925]" captionTargetPageId="126" captionText="Figures 610–619. Neogrosphus griveaudi, male (610–613, 617–619) and female (614–616). Figures 610, 614. Sternopectinal region. Figures 611, 615. Carapace and tergites I–III. Figure 612. Pedipalp chela, movable finger dentate margin. Figures 613, 616. Pedipalp chela in dorsal views. Figure 617. Metasoma V and telson lateral view. Figures 618–619. Right leg III (618) and leg IV (619), tibia, basitarsus and telotarsus, retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.7143863" httpUri="https://zenodo.org/record/7143863/files/figure.png" pageId="13" pageNumber="12">Fig. 614</figureCitation>
); hemispermatophore capsule short, posterior lobe rounded, without lanceolate extension (
<figureCitation id="13292A1AFFBCFFAA251BD717FD2BFAC1" box="[586,665,1646,1671]" captionStart="Figures 52–70" captionStartId="24.[113,190,1591,1613]" captionTargetBox="[116,1470,248,1526]" captionTargetId="figure-361@24.[113,1474,248,1527]" captionTargetPageId="24" captionText="Figures 52–70. Hemispermatophores and capsule regions of Grosphus and Neogrosphus. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and capsule with flagellum; capsule region in convex (and/ or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores unless indicated as mirrored left images. Figure 52. G. madagascariensis, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm). Figure 53. G. madagascariensis, capsule, Sc1197, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 54. G. madagascariensis, capsule, Anjiro, G. halleuxi nr topotype, GLPC. Scale bar: 500 μm. Figure 55. G. madagascariensis, capsule, Mandena- Fort Dauphin, G. mandena paratype, MHNG.Scale bar: 500 μm.Figure 56. G. madagascariensis, capsule, Madagascar, det. Vachon, MHNG. Scale bar: 500 μm. Figure 57. G. madagascariensis, capsule, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 58. G. goudoti, whole hemispermatophore. Scale bar: 2 mm. Figure 59. G. goudoti, capsule, Forêt de Bobankota, holotype, MHNG. Scale bar: 500 μm. Figure 60. G. hirtus, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm), Antsiranana, Ramena vill., mirrored left, GLPC, FKCP. Figure 61. G. hirtus, capsule, Mahajamba River, GLPC, FKCP. Scale bar: 500 μm. Figures 62–65. G. hirtus, capsules in convex view.Antsiranana, Ramena vill., mirrored left, GLPC, FKCP (62), Jardin Botanique, MHNG (63), Ranohira-Llakaka, ZMUH, mirrored left (64), Forêt de Vohitaly, MHNG (65). Scale bars: 500 μm. Figure 66. G. hirtus, capsule, Forest Station Ampijoroa, G. garciai holotype, MHNG. Scale bar: 500 μm. Figure 67. G. voahangyae, whole hemispermatophore. Scale bar: 2 mm. Figure 68. G. voahangyae, capsule, Analamy Forest, FMNH. Scale bar: 500 μm. Figure 69. N. griveaudi, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 70. N. griveaudi, capsule, mirrored left, Tsimanampetsotsa National Park, GLPC, FKCP." figureDoi="http://doi.org/10.5281/zenodo.7143636" httpUri="https://zenodo.org/record/7143636/files/figure.png" pageId="13" pageNumber="12">Fig. 70</figureCitation>
); sternites with moderately narrow spiracles (
<figureCitation id="13292A1AFFBCFFAA26B6D7F7FD34FAE0" box="[487,646,1678,1702]" captionStart="Figures 610–619" captionStartId="126.[113,190,1939,1961]" captionTargetBox="[119,1437,234,1911]" captionTargetId="figure-79@126.[119,1452,234,1925]" captionTargetPageId="126" captionText="Figures 610–619. Neogrosphus griveaudi, male (610–613, 617–619) and female (614–616). Figures 610, 614. Sternopectinal region. Figures 611, 615. Carapace and tergites I–III. Figure 612. Pedipalp chela, movable finger dentate margin. Figures 613, 616. Pedipalp chela in dorsal views. Figure 617. Metasoma V and telson lateral view. Figures 618–619. Right leg III (618) and leg IV (619), tibia, basitarsus and telotarsus, retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.7143863" httpUri="https://zenodo.org/record/7143863/files/figure.png" pageId="13" pageNumber="12">Figs. 610, 614</figureCitation>
); metasoma I with ventromedian carinae moderate, finely granulate; telson with elongate vesicle, without subaculear tubercle; legs with ventral surface of telotarsus densely setose, with&gt; 20 long, filiform setae (
<figureCitation id="13292A1AFFBCFFAA2641D677FE1CFB60" box="[272,430,1806,1830]" captionStart="Figures 610–619" captionStartId="126.[113,190,1939,1961]" captionTargetBox="[119,1437,234,1911]" captionTargetId="figure-79@126.[119,1452,234,1925]" captionTargetPageId="126" captionText="Figures 610–619. Neogrosphus griveaudi, male (610–613, 617–619) and female (614–616). Figures 610, 614. Sternopectinal region. Figures 611, 615. Carapace and tergites I–III. Figure 612. Pedipalp chela, movable finger dentate margin. Figures 613, 616. Pedipalp chela in dorsal views. Figure 617. Metasoma V and telson lateral view. Figures 618–619. Right leg III (618) and leg IV (619), tibia, basitarsus and telotarsus, retrolateral views." figureDoi="http://doi.org/10.5281/zenodo.7143863" httpUri="https://zenodo.org/record/7143863/files/figure.png" pageId="13" pageNumber="12">Figs. 618–619</figureCitation>
); telotarsus with dorsal terminal process very small; cuticle with strong UV fluorescence.
</paragraph>
</subSubSection>
<subSubSection id="C3086514FFBCFFAA2722D612FDA5FBA5" pageId="13" pageNumber="12" type="discussion">
<paragraph id="8BAD369FFFBCFFAA2722D612FEF5FBC2" blockId="13.[115,750,1899,2019]" box="[115,327,1899,1924]" pageId="13" pageNumber="12">SUBORDINATE TAXA.</paragraph>
<paragraph id="8BAD369FFFBCFFAA2722D6F2FD5CFBE5" blockId="13.[115,750,1899,2019]" box="[115,750,1931,1955]" pageId="13" pageNumber="12">
<taxonomicName id="4C124D1CFFBCFFAA2722D6F2FD5CFBE5" authority="Lourenco, Wilme &amp; Waeber, 2015" authorityName="Lourenco, Wilme &amp; Waeber" authorityYear="2015" box="[115,750,1931,1955]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="andrafiabe">
<emphasis id="B966EA8DFFBCFFAA2722D6F2FEC8FBE5" box="[115,378,1931,1955]" italics="true" pageId="13" pageNumber="12">Neogrosphus andrafiabe</emphasis>
<bibRefCitation id="EF834B6EFFBCFFAA26D1D6F5FD5CFBE5" author="LOURENCO, W. R. &amp; L. WILME &amp; P. O. WAEBER" box="[384,750,1931,1955]" journalOrPublisher="Comptes Rendus Biologies" pageId="13" pageNumber="12" pagination="37 - 43" part="339" refId="ref40329" refString="LOURENCO, W. R., L. WILME &amp; P. O. WAEBER. 2015. More about the geographical distribution of the Malagasy genus Neogrosphus Lourenco, 1995 (Scorpiones: Buthidae) and description of a vicariant new species. Comptes Rendus Biologies, 339: 37 - 43." title="More about the geographical distribution of the Malagasy genus Neogrosphus Lourenco, 1995 (Scorpiones: Buthidae) and description of a vicariant new species" type="journal article" year="2015">Lourenço, Wilmé &amp; Waeber, 2015</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BAD369FFFBCFFAA2722D6D2FE4AFB85" blockId="13.[115,750,1899,2019]" box="[115,504,1963,1987]" pageId="13" pageNumber="12">
<taxonomicName id="4C124D1CFFBCFFAA2722D6D2FE4AFB85" authority="Lourenco, 1996" authorityName="Lourenco" authorityYear="1996" box="[115,504,1963,1987]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="blanci">
<emphasis id="B966EA8DFFBCFFAA2722D6D2FEFBFB85" box="[115,329,1963,1987]" italics="true" pageId="13" pageNumber="12">Neogrosphus blanci</emphasis>
<bibRefCitation id="EF834B6EFFBCFFAA261ED6D5FE4AFB85" author="LOURENCO, W. R." box="[335,504,1963,1987]" journalOrPublisher="Biogeographie de Madagascar. ORSTOM, Paris" pageId="13" pageNumber="12" pagination="441 - 455" refId="ref37587" refString="LOURENCO, W. R. 1996 a. Origins and affinities of the scorpion fauna of Madagascar. Pp. 441 - 455 in: Lourenco, W. R. (Ed.). Biogeographie de Madagascar. ORSTOM, Paris." title="Origins and affinities of the scorpion fauna of Madagascar" type="book chapter" volumeTitle="Lourenco, W. R" year="1996">Lourenço, 1996</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BAD369FFFBCFFAA2722D6B2FDA5FBA5" blockId="13.[115,750,1899,2019]" box="[115,535,1995,2019]" pageId="13" pageNumber="12">
<taxonomicName id="4C124D1CFFBCFFAA2722D6B2FDA5FBA5" authority="(Vachon, 1969)" baseAuthorityName="Vachon" baseAuthorityYear="1969" box="[115,535,1995,2019]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="griveaudi">
<emphasis id="B966EA8DFFBCFFAA2722D6B2FEDEFBA5" box="[115,364,1995,2019]" italics="true" pageId="13" pageNumber="12">Neogrosphus griveaudi</emphasis>
(
<bibRefCitation id="EF834B6EFFBCFFAA262BD6B2FDA2FBA5" author="Vachon" box="[378,528,1995,2019]" firstAuthor="Vachon" pageId="13" pageNumber="12" pagination="476 - 483" refId="ref44004" refString="VACHON, M. 1969. Grosphus griveaudi, nouvelle espece de Scorpion Buthidae malgache. Bulletin du Museum national d'Histoire naturelle, Paris, (2), 41: 476 - 483." type="journal article" year="1969">Vachon, 1969</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3086514FFBCFFAA2463D195FC77FE41" pageId="13" pageNumber="12" type="discussion">
<paragraph id="8BAD369FFFBCFFAA2463D195FC77FE41" blockId="13.[818,1476,236,519]" pageId="13" pageNumber="12">
REMARKS. 
<taxonomicName id="4C124D1CFFBCFFAA24EED196FBF9FD41" authorityName="Lourenco" authorityYear="1995" box="[959,1099,239,263]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA24EED196FBF9FD41" box="[959,1099,239,263]" italics="true" pageId="13" pageNumber="12">Neogrosphus</emphasis>
</taxonomicName>
shares some primitive characters with 
<taxonomicName id="4C124D1CFFBCFFAA243DD076FC61FD61" authorityName="Simon" authorityYear="1880" box="[876,979,271,295]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA243DD076FC61FD61" box="[876,979,271,295]" italics="true" pageId="13" pageNumber="12">Grosphus</emphasis>
</taxonomicName>
, and some derived characters with 
<taxonomicName id="4C124D1CFFBCFFAA2233D076FA76FD61" authority="Lowe &amp; Kovařík, 2019" authorityName="Lowe &amp; Kovařík" authorityYear="2019" box="[1378,1476,271,295]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B966EA8DFFBCFFAA2233D076FA76FD61" box="[1378,1476,271,295]" italics="true" pageId="13" pageNumber="12">Teruelius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F6FFBCFFAA2463D056FC33FD00" box="[818,897,303,327]" pageId="13" pageNumber="12" rank="genus">
<emphasis id="B966EA8DFFBCFFAA2463D056FCE8FD01" bold="true" box="[818,858,303,327]" pageId="13" pageNumber="12">gen</emphasis>
. 
<emphasis id="B966EA8DFFBCFFAA243AD056FCC8FD01" bold="true" box="[875,890,303,327]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
(summarized in 
<tableCitation id="C6900324FFBCFFAA236ED056FB27FD01" box="[1087,1173,303,327]" captionStart="Table 4" captionStartId="17.[113,169,1270,1291]" captionTargetId="graphics-446@17.[119,1470,248,1242]" captionTargetPageId="17" captionText="Table 4. Character states for ‘Grosphus’ group genera and outgroup genus Pseudolychas. States of 11 characters for Grosphus, Neogrosphus, Teruelius gen. n. and Pseudolychas, summarizing data of Tables 1–3. Pseudolychas character states determined by study of materials, and from published descriptions. Primitive states indicated by green panels, derived states by red panels, with polarities determined by comparison to Pseudolychas which is assumed primitive. Unpolarized characters indicated by dark green panels." httpUri="http://table.plazi.org/id/DF6D6617FFA0FFB62720D58FFB7EF924" pageId="13" pageNumber="12" tableUuid="DF6D6617FFA0FFB62720D58FFB7EF924">Table 4</tableCitation>
). One interpretation of this is that 
<taxonomicName id="4C124D1CFFBCFFAA24DED036FBA9FD21" authorityName="Lourenco" authorityYear="1995" box="[911,1051,335,359]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA24DED036FBA9FD21" box="[911,1051,335,359]" italics="true" pageId="13" pageNumber="12">Neogrosphus</emphasis>
</taxonomicName>
is descended from an intermediate stage in the evolution of 
<taxonomicName id="4C124D1CFFBCFFAA2315D016FB14FDC1" authority="Lowe &amp; Kovařík, 2019" authorityName="Lowe &amp; Kovařík" authorityYear="2019" box="[1092,1190,367,391]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B966EA8DFFBCFFAA2315D016FB14FDC1" box="[1092,1190,367,391]" italics="true" pageId="13" pageNumber="12">Teruelius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557F6FFBCFFAA23FED016FB4EFDC0" box="[1199,1276,367,391]" pageId="13" pageNumber="12" rank="genus">
<emphasis id="B966EA8DFFBCFFAA23FED016FB65FDC1" bold="true" box="[1199,1239,367,391]" pageId="13" pageNumber="12">gen</emphasis>
. 
<emphasis id="B966EA8DFFBCFFAA23B7D016FB47FDC1" bold="true" box="[1254,1269,367,391]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
from a 
<taxonomicName id="4C124D1CFFBCFFAA2205D016FA09FDC1" authorityName="Simon" authorityYear="1880" box="[1364,1467,367,391]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B966EA8DFFBCFFAA2205D016FA09FDC1" box="[1364,1467,367,391]" italics="true" pageId="13" pageNumber="12">Grosphus</emphasis>
</taxonomicName>
- like ancestor. Other characters, such as small size, internal displacement of 
<emphasis id="B966EA8DFFBCFFAA24A9D0C9FBBAFD81" box="[1016,1032,432,455]" italics="true" pageId="13" pageNumber="12">V</emphasis>
<subScript id="179634DAFFBCFFAA2358D0C6FBA3FD8B" attach="right" box="[1033,1041,447,461]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
, reduced dorsal terminal process of telotarsus and elongated telson appear to be autapomorphies for the genus.
</paragraph>
</subSubSection>
</treatment>
</document>