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<document id="0E06E9B874A4F0577A8FB586AC36BDD8" ID-DOI="10.5281/zenodo.209419" ID-GBIF-Dataset="d425df20-b024-4ae2-bfd6-3b233f94d15e" ID-ISSN="1175-5326" ID-Zenodo-Dep="209419" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460508995608" checkinUser="plazi" docAuthor="Eléaume, Marc, Bohn, Jens-Michael, Roux, Michel &amp; Améziane, Nadia" docDate="2012" docId="D558AA75E01D4C6B6F85F95F58A4FC8A" docLanguage="en" docName="zt03425p022.pdf" docOrigin="Zootaxa 3425" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Feracrinus heinzelleri Eléaume, Bohn, Roux &amp; Améziane, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="2961D20DE01A4C606F12FFE35C3CFF9E" masterDocTitle="Stalked crinoids (Echinodermata) collected by the R / V Polarstern and Meteor in the south Atlantic and in Antarctica" masterLastPageNumber="22" masterPageNumber="1" pageNumber="8" updateTime="1698313077547" updateUser="plazi">
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<mods:title id="4B82B93B0AD2901D74E072835100E575">Stalked crinoids (Echinodermata) collected by the R / V Polarstern and Meteor in the south Atlantic and in Antarctica</mods:title>
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<mods:namePart id="7A13C4202F94D2A4925399B8070EFAAA">Eléaume, Marc</mods:namePart>
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<mods:namePart id="DFAEE72957A8935DD906FB9486DF2A86">Bohn, Jens-Michael</mods:namePart>
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<mods:namePart id="A9A00B694D2B8169F65D7F3177AB653B">Roux, Michel</mods:namePart>
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<mods:namePart id="F1ACA7D30311D1BF1B990729B683306E">Améziane, Nadia</mods:namePart>
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<mods:date id="81B80D73EDBFADE106CB148356F3F805">2012</mods:date>
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<treatment id="D558AA75E01D4C6B6F85F95F58A4FC8A" ID-DOI="http://doi.org/10.5281/zenodo.6180833" ID-GBIF-Taxon="119522924" ID-Zenodo-Dep="6180833" LSID="urn:lsid:plazi:treatment:D558AA75E01D4C6B6F85F95F58A4FC8A" httpUri="http://treatment.plazi.org/id/D558AA75E01D4C6B6F85F95F58A4FC8A" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8">
<subSubSection id="15EB48E8E01D4C676F85F95F5D1FF968" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="5D4E1B63E01D4C676F85F95F5DD2F94B" blockId="7.[151,494,1723,1782]" box="[151,494,1723,1750]" pageId="7" pageNumber="8">
<heading id="0606AC0FE01D4C676F85F95F5DD2F94B" bold="true" box="[151,494,1723,1750]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="6F85C771E01D4C676F85F95F5DD2F94B" bold="true" box="[151,494,1723,1750]" pageId="7" pageNumber="8">
<taxonomicName id="9AF160E0E01D4C676F85F95F5D98F948" ID-CoL="8G8VH" box="[151,420,1724,1750]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E01D4C676F85F95F5D98F948" bold="true" box="[151,420,1724,1750]" italics="true" pageId="7" pageNumber="8">Feracrinus heinzelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74B67A0AE01D4C676EBEF9585DD2F94B" box="[428,494,1723,1749]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5D4E1B63E01D4C676F85F93D5D1FF968" blockId="7.[151,494,1723,1782]" box="[151,291,1758,1782]" pageId="7" pageNumber="8">
<figureCitation id="C5CA07E6E01D4C676F85F93D5D23F968" box="[151,287,1758,1782]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="8.[151,250,1672,1695]" captionStartId-1="9.[151,250,1758,1781]" captionStartId-2="10.[151,250,1446,1469]" captionTargetBox-0="[151,1435,517,1650]" captionTargetBox-1="[151,1435,301,1744]" captionTargetBox-2="[340,1247,193,1424]" captionTargetId-0="figure@8.[151,1436,517,1651]" captionTargetId-1="figure@9.[151,1436,301,1745]" captionTargetId-2="figure@10.[340,1247,193,1425]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." captionText-1="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." captionText-2="FIGURE 6. Paratype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 2). A: general view; B C: distal columnals of preserved stalk." httpUri-0="https://zenodo.org/record/209423/files/figure.png" httpUri-1="https://zenodo.org/record/209424/files/figure.png" httpUri-2="https://zenodo.org/record/209425/files/figure.png" pageId="7" pageNumber="8">Figures 46</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="15EB48E8E01D4C676F85F8C45854F8FD" pageId="7" pageNumber="8" type="etymology">
<paragraph id="5D4E1B63E01D4C676F85F8C45854F8FD" blockId="7.[151,1436,1830,2034]" pageId="7" pageNumber="8">
<emphasis id="6F85C771E01D4C676F85F8C45D22F8A1" bold="true" box="[151,286,1831,1855]" pageId="7" pageNumber="8">Etymology.</emphasis>
This new species is dedicated to Professor T. Heinzeller Ludwig-Maximilian-University, Munich (
<collectingCountry id="25E65BF3E01D4C676F8CF8A95D30F8FC" box="[158,268,1866,1890]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
), who has contributed substantially to our understanding of crinoid anatomy.
</paragraph>
</subSubSection>
<subSubSection id="15EB48E8E01D4C676FD5F88D5CCAF86C" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="5D4E1B63E01D4C676FD5F88D5CCAF86C" blockId="7.[151,1436,1830,2034]" pageId="7" pageNumber="8">
<emphasis id="6F85C771E01D4C676FD5F88D5D8FF819" bold="true" box="[199,435,1902,1927]" pageId="7" pageNumber="8">Material examined.</emphasis>
<typeStatus id="824AA5C1E01D4C676EAFF88D5E19F818" box="[445,549,1902,1926]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
(catalogue number
<collectionCode id="3BE083A6E01D4C676C1DF88D5F74F818" box="[783,840,1902,1926]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="8">ZSM</collectionCode>
0000/1) and one
<typeStatus id="824AA5C1E01D4C676B0BF88C5846F819" box="[1049,1146,1903,1927]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
(catalogue number
<collectionCode id="3BE083A6E01D4C676A70F88D59A7F818" box="[1378,1435,1902,1926]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="8">ZSM</collectionCode>
0000/2) R/V Polarstern cruise ANT
<date id="294F3DA3E01D4C676D3DF8715E59F834" box="[559,613,1938,1962]" bridgedPair="/" pageId="7" pageNumber="8">IX/3</date>
, station 18/192,
<date id="294F3DA3E01D4C676C0DF8715F9CF834" box="[799,928,1938,1962]" pageId="7" pageNumber="8" value="1991-02-27">27.02.1991</date>
, off Queen Maud Land, 69°40.3 to
<geoCoordinate id="38C57DA4E01D4C676A28F8715C9AF850" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-69.675">69°40.5 S</geoCoordinate>
00°51.1 to
<geoCoordinate id="38C57DA4E01D4C676E2EF8555D85F850" box="[316,441,1974,1998]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="0.9133333">00°54.8 E</geoCoordinate>
,
<quantity id="9A09B686E01D4C676ED5F8555E58F850" box="[455,612,1974,1998]" metricMagnitude="3" metricUnit="m" metricValue="1.3955000000000002" metricValueMax="1.3980000000000001" metricValueMin="1.3930000000000002" pageId="7" pageNumber="8" unit="m" value="1395.5" valueMax="1398.0" valueMin="1393.0">13931398 m</quantity>
, housed in the Bavarian State Museum Collection of Zoology (
<collectionCode id="3BE083A6E01D4C676A2BF855594AF850" box="[1337,1398,1974,1998]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="8">ZSM</collectionCode>
) in Munich.
</paragraph>
</subSubSection>
<subSubSection id="15EB48E8E0124C686FD5FF745E53FE51" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="5D4E1B63E0124C686FD5FF745E53FE51" blockId="8.[151,1436,151,463]" pageId="8" pageNumber="9">
<emphasis id="6F85C771E0124C686FD5FF745D7DFF2E" bold="true" box="[199,321,151,176]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
A small species with robust arms of proximal pattern 1+2 3 4 5+6 7+8 or 5+6 7; beyond Br6 series of successive brachial pairs (a+b c+d e+f…) or brachial pairs alternating with free brachials (a+b c d+e f...); series of up to 4 successive muscular articulations. Pinnules relatively robust and rigid; insertion deep, carved on arm side and impacting the following brachial pair; synarthrial articulation on brachial with strong symmorphy; concave cover plates varying from lanceolate to rounded; genital pinnules with one row of additional H-shaped plates. Ratio of primibrachial width to upper radial height 0.73 to 0.85. Tegmen with oral cone higher than anal cone; orals large and erect around mouth; anal cone conspicuous in CD inter-ray and bearing large bulging plates. Conical dorsal cup with ratio of cup height to upper diameter 1 to1.3; basals fused; radials trapezoidal. Columnal symplexies with 8 crenular units; distal syzygies unknown.
</paragraph>
</subSubSection>
<caption id="098E4BEBE0124C686F85F96B5916F964" httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1435,517,1650]" targetPageId="8">
<paragraph id="5D4E1B63E0124C686F85F96B5916F964" blockId="8.[151,1436,1672,1787]" pageId="8" pageNumber="9">
<emphasis id="6F85C771E0124C686F85F96B5D2FF900" bold="true" box="[151,275,1672,1695]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="9AF160E0E0124C686E88F9695E40F900" box="[410,636,1672,1695]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E0124C686E88F9695E40F900" box="[410,636,1672,1695]" italics="true" pageId="8" pageNumber="9">Feracrinus heinzelleri</emphasis>
</taxonomicName>
<emphasis id="6F85C771E0124C686D91F96A5E80F900" bold="true" box="[643,700,1673,1694]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="74B67A0AE0124C686D91F96A5E80F900" box="[643,700,1673,1694]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(ZSM 0000/1). A: general view with ray broken at Br2/3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation.
</paragraph>
</caption>
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<paragraph id="5D4E1B63E0124C686FD5F8C259A0F870" blockId="8.[151,1437,1825,2030]" pageId="8" pageNumber="9">
<emphasis id="6F85C771E0124C686FD5F8C25E23F8A4" bold="true" box="[199,543,1825,1850]" pageId="8" pageNumber="9">
Description of the
<typeStatus id="824AA5C1E0124C686EA3F8C15E26F8A4" box="[433,538,1826,1850]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</emphasis>
Specimen consisting of proximalmost stalk, theca and partly broken crown (
<figureCitation id="C5CA07E6E0124C686F8DF8A55CE0F8C0" box="[159,220,1862,1886]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="8" pageNumber="9">Fig.4</figureCitation>
A). Tegmen heavily plated and not inflated; tegminal plates of variable shape, some perforated by hydropores; anal cone height
<quantity id="9A09B686E0124C686EF6F8895E00F81C" box="[484,572,1898,1922]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="9" unit="mm" value="2.8">2.8 mm</quantity>
, covered by relatively large bulging plates and conspicuous in CD inter-ray (
<figureCitation id="C5CA07E6E0124C686F8DF86D5CD8F838" box="[159,228,1934,1958]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
B); oral cone height
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with conspicuous, convex and smooth orals surrounding beak-like mouth (
<figureCitation id="C5CA07E6E0124C686A7DF86D5C9BF854" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
C); ambulacral grooves merging into arms at height of Br4. Aboral cup tall conical with large ribs prolonging arm axes; cup height
<quantity id="9A09B686E0124C686E40F8355D95F870" box="[338,425,2006,2030]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="9" unit="mm" value="9.3">9.3 mm</quantity>
, upper diameter
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, lower diameter 2.0 mm, ratio of cup height to upper diameter 1.3.
</paragraph>
<paragraph id="5D4E1B63E0134C696F85FF745E3CFF69" blockId="9.[151,1436,151,247]" pageId="9" pageNumber="10">
Radials trapezoidal, height
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, upper width
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, lower width
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. Ratio of primibrachial width to upper radial height 0.73 to 0,77. Basals fused, height of basal ring
<quantity id="9A09B686E0134C696C6EFF5F5FE8FF4A" box="[892,980,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="10" unit="mm" value="3.7">3.7 mm</quantity>
. Remaining proximal stalk
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long, proximalmost diameter
<quantity id="9A09B686E0134C696EB0FF3C5DC0FF69" box="[418,508,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="10" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<caption id="098E4BEBE0134C696F85F93D5CCDF855" httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1435,301,1744]" targetPageId="9">
<paragraph id="5D4E1B63E0134C696F85F93D5CCDF855" blockId="9.[151,1437,1758,1995]" pageId="9" pageNumber="10">
<emphasis id="6F85C771E0134C696F85F93D5D28F96A" bold="true" box="[151,276,1758,1781]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
Holotype of
<taxonomicName id="9AF160E0E0134C696E8CF9035EBEF96B" box="[414,642,1759,1781]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E0134C696E8CF9035EBEF96B" box="[414,642,1759,1781]" italics="true" pageId="9" pageNumber="10">Feracrinus heinzelleri</emphasis>
</taxonomicName>
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<emphasis id="6F85C771E0134C696D9BF93C5EF8F96A" bold="true" box="[649,708,1759,1780]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
(ZSM 0000/1), pinnule ossicles. AC: views of the two proximalmost pinnulars (p1 and p2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p1 articulated to p2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; GI: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular.
</paragraph>
</caption>
<caption id="098E4BEBE0104C6A6F85FA4559A0FA22" box="[151,1436,1446,1469]" httpUri="https://zenodo.org/record/209425/files/figure.png" pageId="10" pageNumber="11" targetBox="[340,1247,193,1424]" targetPageId="10">
<paragraph id="5D4E1B63E0104C6A6F85FA4559A0FA22" blockId="10.[151,1436,1446,1469]" box="[151,1436,1446,1469]" pageId="10" pageNumber="11">
<emphasis id="6F85C771E0104C6A6F85FA455D2EFA22" bold="true" box="[151,274,1446,1469]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
Paratype of
<taxonomicName id="9AF160E0E0104C6A6E83FA4B5E4EFA23" box="[401,626,1447,1469]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E0104C6A6E83FA4B5E4EFA23" box="[401,626,1447,1469]" italics="true" pageId="10" pageNumber="11">Feracrinus heinzelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74B67A0AE0104C6A6D6AFA445E92FA22" box="[632,686,1447,1468]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
(ZSM 0000/2). A: general view; BC: distal columnals of preserved stalk.
</paragraph>
</caption>
<paragraph id="5D4E1B63E0104C6A6FDBFA0A5871F8F7" blockId="10.[151,1437,1513,2005]" pageId="10" pageNumber="11">
One arm broken at Br23, other arms with the same pattern: 1+2 3 4 5+6 7+8 9+10 11+12 13+14 15 16+17 18, with first pinnule on Br4. Series of successive muscular articulations up to 3. Primibrachial width
<quantity id="9A09B686E0104C6A6BF0F9EE5904F9BB" box="[1250,1336,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="11" unit="mm" value="2.7">2.7 mm</quantity>
decreasing progressively to 2.0 mm at Br5+6. Most complete remaining arm
<quantity id="9A09B686E0104C6A6CB1F9D25FCFF9D7" box="[931,1011,1585,1609]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="mm" value="40.0">40 mm</quantity>
long and bearing 12 pinnules on each side. Pinnules relatively robust and rigid; width decreasing rapidly and progressively from proximal pinnular to distal end. Pinnule insertion deep, carved on arm side and impacting the following brachial pair (
<figureCitation id="C5CA07E6E0104C6A6BE3F99A590BF90F" box="[1265,1335,1657,1681]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
D). Most complete pinnule (24 pinnulars) in medial arm reaching a length of 16.0 mm. Brachial synostoses flat (
<figureCitation id="C5CA07E6E0104C6A6A24F97E5947F92B" box="[1334,1403,1693,1717]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
E). Pinnule articulated on brachial by large transverse symmorphial synarthry with a deep ligament fossae (
<figureCitation id="C5CA07E6E0104C6A6A0AF9225966F947" box="[1304,1370,1729,1753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1695]" captionTargetBox="[151,1435,517,1650]" captionTargetId="figure@8.[151,1436,517,1651]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1). A: general view with ray broken at Br 2 / 3; B: anal sac in CD interray; C: oral cone, view from broken A ray; D: lateral view of pinnule insertion on arm; E: hyposynostosial brachial showing flat synostosial facet; F: episynostosial brachial, distal facet showing pinnule socket (arrow head) and interbrachial muscular synarthry (arrow); G: lateral view of same brachial (side of pinnule insertion). Star: muscular articulation." httpUri="https://zenodo.org/record/209423/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
FG). First pinnular (p1) with a sharply convex proximal facet (
<figureCitation id="C5CA07E6E0104C6A6C27F9065F46F963" box="[821,890,1765,1789]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
AB); p1 and p2 articulated by well-developed muscular synarthry (
<figureCitation id="C5CA07E6E0104C6A6E6EF8EA5D81F8BF" box="[380,445,1801,1825]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
BD); other pinnulars articulated by synostoses (
<figureCitation id="C5CA07E6E0104C6A6CCCF8EA5823F8BF" box="[990,1055,1801,1825]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
EF). Proximal part (up to p12) of genital pinnules inflated, with H-shaped additional plates (
<figureCitation id="C5CA07E6E0104C6A6C30F8CE5F5FF8DB" box="[802,867,1837,1861]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
GI) supporting a row of small upper lateral plates (
<figureCitation id="C5CA07E6E0104C6A6F8DF8B25CDDF8F7" box="[159,225,1873,1897]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
JK); cover plates concave varying from lanceolated to rounded (
<figureCitation id="C5CA07E6E0104C6A6CADF8B2583CF8F7" box="[959,1024,1873,1897]" captionStart="FIGURE 5" captionStartId="9.[151,250,1758,1781]" captionTargetBox="[151,1435,301,1744]" captionTargetId="figure@9.[151,1436,301,1745]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 1), pinnule ossicles. A C: views of the two proximalmost pinnulars (p 1 and p 2); A: proximal adoral view showing facet of muscular transverse synarthry; B: adoral view, proximal transverse synarthry at right, distal synostosis at left, p 1 articulated to p 2 by muscular synarthry (see D); C: distal adoral view showing flat synostosial facet; D: proximal synarthry of p 2; E: pinnular in mid pinnule with synostosial facets; F: distal pinnular with muscular articulations; G I: H-shaped genital plate; G: front view; H: lateral external view, ambulacral side at top; I: lateral internal view; J: profile view of genital, H-shaped and covering plates on genital pinnule; K: detailed view of covering plates; L-M: dissociated proximal cover plate. adp: additional lateral plates; cp: cover plates; gp: genital H-shaped plates; p: pinnular." httpUri="https://zenodo.org/record/209424/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
MN).
</paragraph>
<paragraph id="5D4E1B63E0104C6B6FD5F8965E8BFE6A" blockId="10.[151,1437,1513,2005]" lastBlockId="11.[151,1437,151,788]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="6F85C771E0104C6A6FD5F8965E28F813" bold="true" box="[199,532,1909,1934]" pageId="10" pageNumber="11">
Description of the
<typeStatus id="824AA5C1E0104C6A6EB0F8965E33F813" box="[418,527,1909,1933]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
.
</emphasis>
Young immature specimen completely preserved except distal stalk and one arm broken at Br9+10 (
<figureCitation id="C5CA07E6E0104C6A6E60F87A5D8BF82F" box="[370,439,1945,1969]" captionStart="FIGURE 6" captionStartId="10.[151,250,1446,1469]" captionTargetBox="[340,1247,193,1424]" captionTargetId="figure@10.[340,1247,193,1425]" captionTargetPageId="10" captionText="FIGURE 6. Paratype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 2). A: general view; B C: distal columnals of preserved stalk." httpUri="https://zenodo.org/record/209425/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
A). Five relatively robust and smooth arms with 78 pinnules on each side; arm length about
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with 2728 brachials. Proximal pattern 1+2 3 4 5+6 7, with first pinnule on Br4; successive muscular articulations up to 4. Primibrachial width
<quantity id="9A09B686E0114C6B6D6CFF745EE4FF31" box="[638,728,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="12" unit="mm" value="1.7">1.7 mm</quantity>
progressively decreasing to 1.0 mm beyond Br5+6; distance between arm base and synostosis Br5+6
<quantity id="9A09B686E0114C6B6D72FF5F5E84FF4A" box="[608,696,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="mm" value="3.9">3.9 mm</quantity>
. Pinnule length up to
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for 17 pinnulars. Tegmen with large erect orals; oral cone height
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, anal cone height
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. Aboral cup with fused basals and trapezoidal radials, cup height
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, upper diameter 4.0 mm, lower diameter
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, ratio of cup height to upper diameter about 1. Basal height
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, radial height
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; lower radial width
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, upper radial width 2.0 mm. Ratio of primibrachial width to upper radial width 0.85. Length of preserved stalk of 66 columnals 28.0 mm. Proximal most diameter
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, decreasing to
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at a distance of
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from aboral cup and maintaining the same value to distal end. In distal half of preserved stalk, columnals barrel-shaped with height
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. Columnal symplexies less developed, galleried stereom predominating with larger meshes around lumen; 8 small crenular units of 1 crenula near outer border (
<figureCitation id="C5CA07E6E0114C6B6D23FE3F5E4FFE6A" box="[561,627,476,500]" captionStart="FIGURE 6" captionStartId="10.[151,250,1446,1469]" captionTargetBox="[340,1247,193,1424]" captionTargetId="figure@10.[340,1247,193,1425]" captionTargetPageId="10" captionText="FIGURE 6. Paratype of Feracrinus heinzelleri n. sp. (ZSM 0000 / 2). A: general view; B C: distal columnals of preserved stalk." httpUri="https://zenodo.org/record/209425/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
BC).
</paragraph>
</subSubSection>
<subSubSection id="15EB48E8E0114C6B6FD5FDE358A4FC8A" pageId="11" pageNumber="12" type="discussion">
<paragraph id="5D4E1B63E0114C6B6FD5FDE35FCFFD71" blockId="11.[151,1437,151,788]" pageId="11" pageNumber="12">
<emphasis id="6F85C771E0114C6B6FD5FDE35D07FD86" bold="true" box="[199,315,512,536]" pageId="11" pageNumber="12">Remarks.</emphasis>
<taxonomicName id="9AF160E0E0114C6B6E50FDE25DE8FD86" box="[322,468,512,536]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E0114C6B6E50FDE25DE8FD86" box="[322,468,512,536]" italics="true" pageId="11" pageNumber="12">F. heinzelleri</emphasis>
</taxonomicName>
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<emphasis id="6F85C771E0114C6B6EC9FDE35E26FD86" bold="true" box="[475,538,512,536]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
is a species smaller than
<taxonomicName id="9AF160E0E0114C6B6C28FDE25FFCFD86" box="[826,960,512,536]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="aculeatus">
<emphasis id="6F85C771E0114C6B6C28FDE25FFCFD86" box="[826,960,512,536]" italics="true" pageId="11" pageNumber="12">F. aculeatus</emphasis>
</taxonomicName>
and
<emphasis id="6F85C771E0114C6B6CEAFDE2585AFD86" box="[1016,1126,512,536]" italics="true" pageId="11" pageNumber="12">F. k o s lo w i</emphasis>
. It is distinguished by having a highly conical aboral cup, smooth convex orals, narrow inter-rays (ratio of primibrachial width to upper radial height&gt;0.7), anal cone with relatively large bulging plates and pinnules deeply inserted on arm side impacting the following brachial pair. Stalk articulations are only known in the proximal part of a juvenile specimen, without data on distal syzygies which usually bear specific characters (
<bibRefCitation id="39606692E0114C6B6C09FD6C580CFD39" author="Ameziane" box="[795,1072,655,679]" pageId="11" pageNumber="12" refString="Ameziane, N. &amp; Roux, M. (2011) Stalked crinoids from Southern Tasmanian Seamounts. Part 1: Family Hyocrinidae. Journal of Natural History, 45 (3), 137 - 170." type="journal article" year="2011">Améziane &amp; Roux 2011</bibRefCitation>
).
<taxonomicName id="9AF160E0E0114C6B6B51FD7258E8FD36" box="[1091,1236,656,680]" class="Crinoidea" family="Hyocrinidae" genus="Feracrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="heinzelleri" status="sp. nov.">
<emphasis id="6F85C771E0114C6B6B51FD7258E8FD36" box="[1091,1236,656,680]" italics="true" pageId="11" pageNumber="12">F. heinzelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="74B67A0AE0114C6B6BC9FD73592BFD39" box="[1243,1303,656,679]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
shares with
<emphasis id="6F85C771E0114C6B6F85FD565D39FD52" box="[151,261,692,716]" italics="true" pageId="11" pageNumber="12">F. k o s l o w i</emphasis>
the same pattern of proximal and mid-arm articulations with rare free brachials. These two species live at about the same depths, whereas
<emphasis id="6F85C771E0114C6B6D09FD3A5EA1FD6E" box="[539,669,728,752]" italics="true" pageId="11" pageNumber="12">F. a c u l e a t u s</emphasis>
lives in deeper environments.
</paragraph>
<paragraph id="5D4E1B63E0114C6B6FD5FD1858A4FC8A" blockId="11.[151,1437,151,788]" box="[199,1176,763,788]" pageId="11" pageNumber="12">
<emphasis id="6F85C771E0114C6B6FD5FD185D65FC8A" bold="true" box="[199,345,763,788]" pageId="11" pageNumber="12">Occurrence.</emphasis>
<collectingCountry id="25E65BF3E0114C6B6E72FD1F5DE9FC8A" box="[352,469,764,788]" name="Antarctica" pageId="11" pageNumber="12">Antarctica</collectingCountry>
slope, E of Weddell Sea, at a depth between
<date id="294F3DA3E0114C6B6CDCFD1F5848FC8A" box="[974,1140,764,788]" pageId="11" pageNumber="12" value="1393" valueMax="1398">
1393 and
<quantity id="9A09B686E0114C6B6B2EFD1F5848FC8A" box="[1084,1140,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.398" pageId="11" pageNumber="12" unit="m" value="1398.0">1398</quantity>
</date>
m.
</paragraph>
</subSubSection>
</treatment>
</document>