165 lines
17 KiB
XML
165 lines
17 KiB
XML
<document id="485336264920FE954FC335FC0885EF08" ID-CLB-Dataset="46518" ID-DOI="10.11646/zootaxa.3760.3.13" ID-GBIF-Dataset="b63cb27f-e35d-43ae-b126-3bad252181a9" ID-ISSN="1175-5326" ID-Zenodo-Dep="225103" ID-ZooBank="E9A2C691-3121-4521-B019-735DD1B32C19" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460379358104" checkinUser="plazi" docAuthor="Santos, Claudiney Biral Dos & Curler, Gregory R." docDate="2014" docId="C23DEF46FFC3FFC9FF4A4D2E587D9D43" docLanguage="en" docName="zt03760p470.pdf" docOrigin="Zootaxa 3760 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoira ferreirai Santos, sp. nov." docType="treatment" docVersion="9" lastPageNumber="467" masterDocId="3E04973EFFC1FFCDFFDD4B0F592A9C27" masterDocTitle="Four new species of Tonnoira Enderlein (Diptera: Psychodidae: Psychodinae) from the Brazilian Atlantic forest" masterLastPageNumber="470" masterPageNumber="463" pageNumber="465" updateTime="1698296622868" updateUser="ExternalLinkService">
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<mods:mods id="FF3B903E88DEE744B77C551212888DE5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AF46DAAEE6BF360E5F40E44AA7FEAA9A">
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<mods:title id="97B4748542B7E6BF7FABAA0B0F063CE3">Four new species of Tonnoira Enderlein (Diptera: Psychodidae: Psychodinae) from the Brazilian Atlantic forest</mods:title>
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</mods:titleInfo>
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<mods:name id="404DBC0839DC90AFE0E0ADEB7E097436" type="personal">
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<mods:roleTerm id="14EADCFFD5977F15F8D7BA90CC5504E0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A287E9EAA2F5F909EAE8AE8C2E113687">Santos, Claudiney Biral Dos</mods:namePart>
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</mods:name>
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<mods:name id="619583477B68EE675F2280FB33A1C5C2" type="personal">
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<mods:role id="039B983D124055ECD9A767C96FB6326F">
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<mods:roleTerm id="92B5D847B549638161379A4B817258F5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D4D9CEF20046F2CD1696E935B622F854">Curler, Gregory R.</mods:namePart>
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<mods:typeOfResource id="D9709F3195DA7701FCD83AC58EF12D76">text</mods:typeOfResource>
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<mods:title id="4E238EE9CFD05F0EC3BD91D45DD7C997">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C79171DA332E151E6164CD8121CCE454">
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<mods:date id="E5F696E82AF79664203784583F6DC0C5">2014</mods:date>
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<mods:detail id="BDBB496338E9B0CD2B62708A3EBC8878" type="volume">
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<mods:number id="795422C7A159175556B59E9CA4A9448A">3760</mods:number>
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</mods:detail>
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<mods:detail id="DF6F3B6ED1323B62C630847F50BDF754" type="issue">
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<mods:number id="63ACA8CADE92D114BF2E66E4AD923A2E">3</mods:number>
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<mods:end id="BEA0BA82B072DA1318AD17920694B529">470</mods:end>
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<mods:classification id="5D18FC3774328F00F43391D8125FE143">journal article</mods:classification>
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<mods:identifier id="87226856A4D1ABF52707D3CE16DA822F" type="CLB-Dataset">46518</mods:identifier>
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<mods:identifier id="EE1C7FDF819BB7D7F18EB17020A6A522" type="DOI">10.11646/zootaxa.3760.3.13</mods:identifier>
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<mods:identifier id="78375D69801CCDBFA1495F88F4EC2503" type="GBIF-Dataset">b63cb27f-e35d-43ae-b126-3bad252181a9</mods:identifier>
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<mods:identifier id="401468D1A0A2716D4C9224FFDDAD261D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="144C127BA9DBBA905A0F54A622E64EF3" type="Zenodo-Dep">225103</mods:identifier>
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<treatment id="C23DEF46FFC3FFC9FF4A4D2E587D9D43" ID-DOI="http://doi.org/10.5281/zenodo.6141183" ID-GBIF-Taxon="119413833" ID-Zenodo-Dep="6141183" LSID="urn:lsid:plazi:treatment:C23DEF46FFC3FFC9FF4A4D2E587D9D43" httpUri="http://treatment.plazi.org/id/C23DEF46FFC3FFC9FF4A4D2E587D9D43" lastPageId="4" lastPageNumber="467" pageId="2" pageNumber="465">
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<subSubSection id="028E0DDBFFC3FFCFFF4A4D2E583B9A7A" pageId="2" pageNumber="465" type="nomenclature">
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<paragraph id="4A2B5E50FFC3FFCFFF4A4D2E5B359A1D" blockId="2.[151,543,1569,1629]" box="[151,543,1569,1594]" pageId="2" pageNumber="465">
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<heading id="1163E93CFFC3FFCFFF4A4D2E5B359A1D" bold="true" box="[151,543,1569,1594]" fontSize="10" level="2" pageId="2" pageNumber="465" reason="2">
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<emphasis id="78E08242FFC3FFCFFF4A4D2E5B359A1D" bold="true" box="[151,543,1569,1594]" pageId="2" pageNumber="465">
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<taxonomicName id="8D9425D3FFC3FFCFFF4A4D2E58949A1D" ID-CoL="9HRY9" authority="Santos" authorityName="Santos" box="[151,446,1569,1594]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="ferreirai" status="sp. nov.">
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<emphasis id="78E08242FFC3FFCFFF4A4D2E58439A1D" bold="true" box="[151,361,1569,1594]" italics="true" pageId="2" pageNumber="465">Tonnoira ferreirai</emphasis>
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Santos
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</taxonomicName>
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<taxonomicNameLabel id="63D33F39FFC3FFCFFE184D2E5B359A1D" box="[453,543,1569,1594]" pageId="2" pageNumber="465" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4A2B5E50FFC3FFCFFF4A4D4B583B9A7A" blockId="2.[151,543,1569,1629]" box="[151,273,1604,1629]" pageId="2" pageNumber="465">
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(
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<figureCitation id="D2AF42D5FFC3FFCFFF424D4B58209A7A" box="[159,266,1604,1629]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1789,1812]" captionTargetBox="[151,1436,373,1768]" captionTargetId="figure@3.[151,1436,373,1769]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Tonnoira ferreirai sp. nov., holotype ♂. 6. Head, frontal view. 7. Wing. 8. Hypandrium, gonopods, aedeagal complex and parameres, dorsal view. 9. Epandrium and epandrial clasping lobes, dorsal view." httpUri="https://zenodo.org/record/225105/files/figure.png" pageId="2" pageNumber="465">Figs. 6–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="028E0DDBFFC3FFCFFF4A4D835C0E9AD7" pageId="2" pageNumber="465" type="diagnosis">
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<paragraph id="4A2B5E50FFC3FFCFFF4A4D835C0E9AD7" blockId="2.[151,1436,1676,2034]" pageId="2" pageNumber="465">
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<emphasis id="78E08242FFC3FFCFFF4A4D83583B9A82" bold="true" box="[151,273,1676,1701]" pageId="2" pageNumber="465">Diagnosis.</emphasis>
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Eye bridge divided medially by 0.3 facet diameter. Wing with base of R2+3 not connected to R4, CuA not attached basally or apically. Gonostylus bifurcated near the base with lateral branch about ¼ the length of mesal branch. Epandrial clasping lobes tapered from base to apex, strongly curved apically, with two tenacula.
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</paragraph>
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</subSubSection>
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<subSubSection id="028E0DDBFFC3FFCEFF1A4DF45BAA9D67" lastPageId="3" lastPageNumber="466" pageId="2" pageNumber="465" type="description">
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<paragraph id="4A2B5E50FFC3FFCEFF1A4DF45BAA9D67" blockId="2.[151,1436,1676,2034]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="466" pageId="2" pageNumber="465">
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<emphasis id="78E08242FFC3FFCFFF1A4DF4587B9B33" bold="true" box="[199,337,1787,1812]" pageId="2" pageNumber="465">Description</emphasis>
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. Male. Length total
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<quantity id="8D6CF3B5FFC3FFCFFDE94DF35BA69B34" box="[564,652,1788,1812]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="465" unit="mm" value="3.3">3.3 mm</quantity>
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. Head: eye bridge divided medially by 0.3 facet diameters; interocular suture inverted Y-shaped; antenna with 16 antennomeres; scape cylindrical, 1.7 times the length of pedicel; pedicel spherical; flagellum with 14 flagellomeres; flagellomeres cylindrical; basal flagellomere 1.1 times the length of second; apiculus present on flagellomere 14; palpus formula = 1.0: 2.3: 2.6: 2.6 (fig. 6); apical palp segment not striated. Wing with Sc short, with apex unattached; base of R2+3 not connected to R4; R5 unattached basally, terminating slightly posterior to wing apex; CuA1 unattached basally; CuA2 free (fig. 7). Terminalia: Gonocoxites 1.6 times the length of gonostyli, contiguous with hypandrium; hypandrium narrow, arcuate in dorsal view; gonostylus bifurcated near base with lateral branch about ¼ the length of medial branch; basiphallus ovoid; parameres slightly curved, directed posterolaterally, tapering to apex, shorter than distiphallus (fig. 8); epandrium shorter than wide, appearing U-shaped with the pseudospiracular opening; hypoproct micropilose, with posterior margin rounded; epandrial clasping lobes tapered from base to apex, strongly curved near apices, each with one apical and one subapical tenaculum (fig. 9).
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</paragraph>
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</subSubSection>
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<caption id="1EEB0ED8FFC2FFCEFF4A4DF25D619B14" httpUri="https://zenodo.org/record/225105/files/figure.png" pageId="3" pageNumber="466" targetBox="[151,1436,373,1768]" targetPageId="3">
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<paragraph id="4A2B5E50FFC2FFCEFF4A4DF25D619B14" blockId="3.[151,1436,1789,1843]" pageId="3" pageNumber="466">
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<emphasis id="78E08242FFC2FFCEFF4A4DF258119B34" bold="true" box="[151,315,1789,1812]" pageId="3" pageNumber="466">FIGURES 6–9.</emphasis>
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<taxonomicName id="8D9425D3FFC2FFCEFE9E4DF058D19B33" box="[323,507,1790,1812]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="ferreirai" status="sp. nov.">
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<emphasis id="78E08242FFC2FFCEFE9E4DF058D19B33" box="[323,507,1790,1812]" italics="true" pageId="3" pageNumber="466">Tonnoira ferreirai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="63D33F39FFC2FFCEFDDE4DF05B7E9B33" box="[515,596,1791,1812]" pageId="3" pageNumber="466" rank="species">
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<emphasis id="78E08242FFC2FFCEFDDE4DF05B659B33" bold="true" box="[515,591,1791,1812]" pageId="3" pageNumber="466">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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, holotype ♂.
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<emphasis id="78E08242FFC2FFCEFD394DF15BDD9B34" bold="true" box="[740,759,1790,1811]" pageId="3" pageNumber="466">6.</emphasis>
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Head, frontal view.
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<emphasis id="78E08242FFC2FFCEFC164DF05AF49B33" bold="true" box="[971,990,1791,1812]" pageId="3" pageNumber="466">7.</emphasis>
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Wing.
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<emphasis id="78E08242FFC2FFCEFBF74DF15D179B34" bold="true" box="[1066,1085,1790,1811]" pageId="3" pageNumber="466">8.</emphasis>
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Hypandrium, gonopods, aedeagal complex and parameres, dorsal view.
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<emphasis id="78E08242FFC2FFCEFDC94C125B0D9B15" bold="true" box="[532,551,1821,1842]" pageId="3" pageNumber="466">9.</emphasis>
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Epandrium and epandrial clasping lobes, dorsal view.
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</paragraph>
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</caption>
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<subSubSection id="028E0DDBFFC2FFCEFF1A4C50582A9BE7" pageId="3" pageNumber="466" type="materials_examined">
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<paragraph id="4A2B5E50FFC2FFCEFF1A4C50582A9BE7" blockId="3.[151,1436,1887,2021]" pageId="3" pageNumber="466">
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<emphasis id="78E08242FFC2FFCEFF1A4C50589D9B5F" bold="true" box="[199,439,1887,1912]" pageId="3" pageNumber="466">Material examined.</emphasis>
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<collectingCountry id="32831EC0FFC2FFCEFE194C6F5B069B5F" box="[452,556,1888,1912]" name="Brazil" pageId="3" pageNumber="466">BRAZIL</collectingCountry>
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, state of Espírito Santo, Santa Teresa, Reserva Biológia Augusto Ruschi, (
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<geoCoordinate id="2FA03897FFC2FFCEFF7D4C8C59D69BBB" box="[160,252,1923,1948]" direction="south" orientation="latitude" pageId="3" pageNumber="466" precision="925" value="-19.9">19º54’S</geoCoordinate>
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,
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<geoCoordinate id="2FA03897FFC2FFCEFED44C8B58479BBB" box="[265,365,1924,1948]" direction="west" orientation="longitude" pageId="3" pageNumber="466" precision="925" value="-40.55">40º33’W</geoCoordinate>
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,
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<specimenCount id="5C9295D9FFC2FFCEFEA74C8B58E49BBB" box="[378,462,1923,1948]" pageId="3" pageNumber="466" type="generic">
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<quantity id="8D6CF3B5FFC2FFCEFEA74C8B58E09BBB" box="[378,458,1923,1948]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="466" unit="m" value="820.0">820 m</quantity>
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.
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</specimenCount>
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a.s.l.),
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<typeStatus id="952FE0F2FFC2FFCEFDCA4C8C5B539BBB" box="[535,633,1923,1948]" pageId="3" pageNumber="466" type="holotype">holotype</typeStatus>
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male,
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<date id="3E2A7890FFC2FFCEFD1A4C8C5A6B9BBB" box="[711,833,1923,1948]" pageId="3" pageNumber="466" value="2011-09-28">28.ix.2011</date>
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, coll. C.B. Santos, five
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<typeStatus id="952FE0F2FFC2FFCEFB8C4C8B5D9B9BBB" box="[1105,1201,1924,1948]" pageId="3" pageNumber="466" type="paratype">paratype</typeStatus>
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males, same data as
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<typeStatus id="952FE0F2FFC2FFCEFF4A4CA859D79BE7" box="[151,253,1959,1984]" pageId="3" pageNumber="466" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="028E0DDBFFC2FFC9FF1A4CC35D399C97" lastPageId="4" lastPageNumber="467" pageId="3" pageNumber="466" type="etymology">
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<paragraph id="4A2B5E50FFC2FFC9FF1A4CC35D399C97" blockId="3.[151,1436,1887,2021]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="467" pageId="3" pageNumber="466">
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<emphasis id="78E08242FFC2FFCEFF1A4CC3586C9BC2" bold="true" box="[199,326,1996,2021]" pageId="3" pageNumber="466">Etymology</emphasis>
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. This species is named for Dr. Adelson Luiz Ferreira, Universidade Federal do Espírito Santo, in honor of his contributions to the study of
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<taxonomicName id="8D9425D3FFC5FFC9FDB54B985BDC9C97" box="[616,758,151,176]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
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in Espírito Santo,
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<collectingCountry id="32831EC0FFC5FFC9FC1A4B985D259C97" box="[967,1039,151,176]" name="Brazil" pageId="4" pageNumber="467">Brazil</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="028E0DDBFFC5FFC9FF1A4BB3587D9D43" pageId="4" pageNumber="467" type="discussion">
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<paragraph id="4A2B5E50FFC5FFC9FF1A4BB3587D9D43" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="467">
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<emphasis id="78E08242FFC5FFC9FF1A4BB358659CF2" bold="true" box="[199,335,188,213]" pageId="4" pageNumber="467">Comments.</emphasis>
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<taxonomicName id="8D9425D3FFC5FFC9FE854BB25B0C9CF3" box="[344,550,188,212]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ferreirai">
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<emphasis id="78E08242FFC5FFC9FE854BB25B0C9CF3" box="[344,550,188,212]" italics="true" pageId="4" pageNumber="467">Tonnoira ferreirai</emphasis>
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</taxonomicName>
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can be readily distinguished from other species by the narrowly divided eye bridge with four facet rows and the bifurcate gonostyli with markedly shorter lateral branches.
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<taxonomicName id="8D9425D3FFC5FFC9FB664BEE5C169CDF" box="[1211,1340,223,248]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="andradei">
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<emphasis id="78E08242FFC5FFC9FB664BEE5C169CDF" box="[1211,1340,223,248]" italics="true" pageId="4" pageNumber="467">T. andradei</emphasis>
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</taxonomicName>
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also has bifurcate gonostyli, but with lateral branches longer than medial branches. Overall,
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<taxonomicName id="8D9425D3FFC5FFC9FB9A4A0A5DEB9D3B" box="[1095,1217,260,284]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ferreirai">
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<emphasis id="78E08242FFC5FFC9FB9A4A0A5DEB9D3B" box="[1095,1217,260,284]" italics="true" pageId="4" pageNumber="467">T. ferreirai</emphasis>
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</taxonomicName>
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appears to be most similar morphologically to
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<taxonomicName id="8D9425D3FFC5FFC9FE0C4A265B7E9D67" box="[465,596,295,320]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="andradei">
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<emphasis id="78E08242FFC5FFC9FE0C4A265B7E9D67" box="[465,596,295,320]" italics="true" pageId="4" pageNumber="467">T. andradei</emphasis>
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</taxonomicName>
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. The apiculus on flagellomere 14 of
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<taxonomicName id="8D9425D3FFC5FFC9FBD84A265DBF9D67" box="[1029,1173,296,320]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ferreirana">
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<emphasis id="78E08242FFC5FFC9FBD84A265DBF9D67" box="[1029,1173,296,320]" italics="true" pageId="4" pageNumber="467">T. ferreirana</emphasis>
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</taxonomicName>
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is similar to that of
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<taxonomicName id="8D9425D3FFC5FFC9FA5A4A2659DC9D42" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="brisolaii">
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<emphasis id="78E08242FFC5FFC9FA5A4A2659DC9D42" italics="true" pageId="4" pageNumber="467">T. brisolaii</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D2AF42D5FFC5FFC9FED84A4358609D43" box="[261,330,332,357]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1445,1468]" captionTargetBox="[196,1391,193,1424]" captionTargetId="figure@2.[196,1392,193,1425]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Tonnoira brisolaii sp. nov., holotype ♂. 1. Head, frontal view. 2. Flagellomeres 13 – 14. 3. Wing. 4. Hypandrium, gonopods, aedeagal complex and parameres, dorsal view. 5. Epandrium and epandrial clasping lobes, dorsal view." httpUri="https://zenodo.org/record/225104/files/figure.png" pageId="4" pageNumber="467">Fig. 2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |