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<document id="FFA30346F21B5BEFED148658A1DF07F8" ID-DOI="10.11646/zootaxa.3745.4.1" ID-GBIF-Dataset="04107669-3283-4ee6-b835-3fc467eacd8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="247366" ID-ZooBank="F158CE77-9392-45A4-93F9-28FB1F5FF85A" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460706291994" checkinUser="plazi" docAuthor="Qiu, Jian-Yue, Xu, Hao &amp; Hu, Chun-Lin" docDate="2013" docId="9D180B59FFA5FFA0798AFA09C577F823" docLanguage="en" docName="zt03745p434.pdf" docOrigin="Zootaxa 3745 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmiomorpha (Cosmiomorpha) sauteri Bourgoin 1931" docType="treatment" docVersion="8" lastPageNumber="417" masterDocId="61217321FFAAFFB0791DFFAFC619FFAD" masterDocTitle="Revision of the subgenus Cosmiomorpha (Cosmiomorpha) (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="434" masterPageNumber="401" pageNumber="416" updateTime="1698357144284" updateUser="plazi">
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<mods:title id="2E973712B6297555DC8F814C8AF6FC96">Revision of the subgenus Cosmiomorpha (Cosmiomorpha) (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<mods:namePart id="1F393920F8081A82C41B25C526D2E557">Qiu, Jian-Yue</mods:namePart>
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<mods:namePart id="E8078E545058E0E12703E326CED890AC">Xu, Hao</mods:namePart>
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<treatment id="9D180B59FFA5FFA0798AFA09C577F823" ID-DOI="http://doi.org/10.5281/zenodo.6151408" ID-GBIF-Taxon="119584111" ID-Zenodo-Dep="6151408" LSID="urn:lsid:plazi:treatment:9D180B59FFA5FFA0798AFA09C577F823" httpUri="http://treatment.plazi.org/id/9D180B59FFA5FFA0798AFA09C577F823" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
<subSubSection id="5DABE9C4FFA5FFBF798AFA09C5ADF9C9" pageId="15" pageNumber="416" type="nomenclature">
<paragraph id="150EBA4FFFA5FFBF798AFA09C55CFA6D" blockId="15.[151,875,1446,1506]" box="[151,837,1446,1472]" pageId="15" pageNumber="416">
<heading id="4E460D23FFA5FFBF798AFA09C55CFA6D" bold="true" box="[151,837,1446,1472]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
<taxonomicName id="D2B1C1CCFFA5FFBF798AFA09C55CFA6D" authority="Bourgoin, 1931" authorityName="Bourgoin" authorityYear="1931" box="[151,837,1446,1472]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="sauteri" subGenus="Cosmiomorpha">
<emphasis id="27C5665DFFA5FFBF798AFA09C55CFA6D" bold="true" box="[151,837,1446,1472]" pageId="15" pageNumber="416">
<emphasis id="27C5665DFFA5FFBF798AFA09C748FA6D" bold="true" box="[151,337,1446,1472]" italics="true" pageId="15" pageNumber="416">Cosmiomorpha</emphasis>
(
<emphasis id="27C5665DFFA5FFBF787CFA09C402FA6D" bold="true" box="[353,539,1446,1472]" italics="true" pageId="15" pageNumber="416">Cosmiomorpha</emphasis>
)
<emphasis id="27C5665DFFA5FFBF7B31FA09C466FA6D" bold="true" box="[556,639,1446,1472]" italics="true" pageId="15" pageNumber="416">sauteri</emphasis>
Bourgoin, 1931
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="150EBA4FFFA5FFBF798AFA66C572FA4F" blockId="15.[151,875,1446,1506]" box="[151,875,1481,1506]" pageId="15" pageNumber="416">
(
<figureCitation id="8D8AA6CAFFA5FFBF7982FA66C6E5FA4F" box="[159,252,1481,1506]" captionStart="FIGURES 1 20" captionStartId="4.[151,264,1561,1584]" captionTargetBox="[166,1436,195,1539]" captionTargetId="figure@4.[163,1448,193,1542]" captionTargetPageId="4" captionText="FIGURES 1 20. Mesosternal process of male Cosmiomorpha species. 1 4. Cosmiomorpha fortis new species: 1. Holotype; 2 4. Paratypes; 5. Cosmiomorpha nigripedis new species, holotype; 6. Cosmiomorpha maolanensis new species, holotype; 7. Cosmiomorpha cheni new species, holotype; 8 9. Cosmiomorpha modesta: 8. (Jiangsu); 9. (Guangxi); 10. Cosmiomorpha sauteri (Taiwan); 11 20. Cosmiomorpha decliva: 11. (Sichuan); 12. Cosmiomorpha squamulosa (Gansu), lectotype; 13. (Hunan); 14. (Sichuan); 15. (Guangxi); 16. (Fujian); 17. Cosmiomorpha baryi (N. Vietnam), “ allotype ”; 18. (Jiangxi); 19. (Fujian); 20. (Guangxi). Not to the same scale." httpUri="https://zenodo.org/record/247367/files/figure.png" pageId="15" pageNumber="416">Figs. 10</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7815FA66C748FA4F" box="[264,337,1481,1506]" captionStart="FIGURES 21 34" captionStartId="5.[151,264,1426,1449]" captionTargetBox="[165,1409,199,1391]" captionTargetId="figure@5.[151,1436,195,1405]" captionTargetPageId="5" captionText="FIGURES 21 34. Head of Cosmiomorpha species in dorsal and ventral view. 21 22. Cosmiomorpha modesta; 23 24. Cosmiomorpha decliva; 25 26. Cosmiomorpha sauteri; 27 28. Cosmiomorpha fortis new species, holotype; 29 30. Cosmiomorpha nigripedis new species, holotype; 31 32. Cosmiomorpha maolanensis new species, holotype; 33 34. Cosmiomorpha c heni new species, holotype. Not to the same scale." httpUri="https://zenodo.org/record/247368/files/figure.png" pageId="15" pageNumber="416">2526</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7840FA65C7BCFA4F" box="[349,421,1482,1506]" captionStart="FIGURES 35 47" captionStartId="6.[151,264,985,1008]" captionTargetBox="[157,1430,446,959]" captionTargetId="figure@6.[151,1436,445,964]" captionTargetPageId="6" captionText="FIGURES 35 47. Protibia and basitarsus of male Cosmiomorpha species in dorsal view. 35 36. Cosmiomorpha modesta: 35. (Jiangsu); 36. (Guangxi); 37 38. Cosmiomorpha decliva: 37. (Sichuan); 38. (Guangxi); 39 40. Cosmiomorpha sauteri (Taiwan); 41 43. Cosmiomorpha fortis new species: 41. Holotype; 42 43. Paratypes; 44. Cosmiomorpha nigripedis new species, holotype; 45 46. Cosmiomorpha maolanensis new species: 45. Holotype; 46. Paratype; 47. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247369/files/figure.png" pageId="15" pageNumber="416">3940</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF78ADFA66C7E0FA4F" box="[432,505,1481,1506]" captionStart="FIGURES 48 60" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[160,1427,1181,1768]" captionTargetId="figure@6.[151,1436,1180,1774]" captionTargetPageId="6" captionText="FIGURES 48 60. Protibia and basitarsus of male Cosmiomorpha species in lateral view. 48. Cosmiomorpha modesta (Jiangsu); 49 51. Cosmiomorpha decliva: 49. (Sichuan); 50. (Guangxi); 51. (Fujian); 52 53. Cosmiomorpha sauteri (Taiwan); 54 56. Cosmiomorpha fortis new species: 54. Holotype; 55 56. Paratypes; 57. Cosmiomorpha nigripedis new species, holotype; 58 59. Cosmiomorpha maolanensis new species: 58. Holotype; 59. Paratype; 60. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247370/files/figure.png" pageId="15" pageNumber="416">5253</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7B19FA66C454FA4F" box="[516,589,1481,1506]" captionStart="FIGURES 61 75" captionStartId="7.[151,264,1256,1279]" captionTargetBox="[153,1433,194,1233]" captionTargetId="figure@7.[151,1436,193,1236]" captionTargetPageId="7" captionText="FIGURES 61 75. Scales on pronotal disc of male Cosmiomorpha species 61 62. Cosmiomorpha modesta (Zhejiang); 63 65. Cosmiomorpha decliva: 63. (Fujian); 64. (Sichuan); 65. (Fujian); 66 67. Cosmiomorpha sauteri (Taiwan); 68 72. Cosmiomorpha fortis new species: 68. Holotype; 69 72. Paratype; 73. Cosmiomorpha nigripedis new species, holotype; 74. Cosmiomorpha maolanensis new species, holotype; 75. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247371/files/figure.png" pageId="15" pageNumber="416">6667</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7B45FA66C4A4FA4F" box="[600,701,1481,1506]" captionStart="FIGURES 126 132" captionStartId="16.[151,264,1343,1366]" captionTargetBox="[171,1436,341,1318]" captionTargetId="figure@16.[163,1448,337,1323]" captionTargetPageId="16" captionText="FIGURES 126 132. Cosmiomorpha sauteri Bourgoin, 1931. 126 129. Male; 130 132. Female." httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="15" pageNumber="416">126132</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7BD5FA66C535FA4F" box="[712,812,1481,1506]" captionStart="FIGURES 151 164" captionStartId="24.[151,264,1842,1865]" captionTargetBox="[161,1424,888,1807]" captionTargetId="figure@24.[151,1436,877,1822]" captionTargetPageId="24" captionText="FIGURES 151 164. Parameres of Cosmiomorpha species in dorsal and apical view. 151, 158. Cosmiomorpha modesta; 152, 159. Cosmiomorpha decliva; 153, 160. Cosmiomorpha sauteri; 154, 161. Cosmiomorpha fortis new species, holotype; 155, 162. Cosmiomorpha nigripedis new species, holotype; 156, 163. Cosmiomorpha maolanensis new species, holotype; 157, 164. Cosmiomorpha c heni new species, holotype." httpUri="https://zenodo.org/record/247381/files/figure.png" pageId="15" pageNumber="416">153, 160</figureCitation>
,
<figureCitation id="8D8AA6CAFFA5FFBF7A25FA66C57AFA4F" box="[824,867,1481,1506]" captionStart="FIGURES 165 171" captionStartId="25.[151,264,962,985]" captionTargetBox="[160,1431,227,918]" captionTargetId="figure@25.[152,1437,223,930]" captionTargetPageId="25" captionText="FIGURES 165 171. Lateral view of Cosmiomorpha species. 165. Cosmiomorpha modesta; 166. Cosmiomorpha decliva; 167. Cosmiomorpha sauteri; 168. Cosmiomorpha fortis new species, holotype; 169. Cosmiomorpha nigripedis new species, holotype; 170. Cosmiomorpha maolanensis new species, holotype; 171. Cosmiomorpha c heni new species, holotype." httpUri="https://zenodo.org/record/247382/files/figure.png" pageId="15" pageNumber="416">167</figureCitation>
)
</paragraph>
<paragraph id="150EBA4FFFA5FFBF798AF9BEC5ADF9C9" blockId="15.[151,1436,1553,1637]" pageId="15" pageNumber="416">
<taxonomicName id="D2B1C1CCFFA5FFBF798AF9BEC479F98A" authority="Bourgoin, 1931: 45" authorityName="Bourgoin" authorityPageNumber="45" authorityYear="1931" box="[151,608,1553,1575]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFA5FFBF798AF9BEC79DF98A" box="[151,388,1553,1575]" italics="true" pageId="15" pageNumber="416">Cosmiomorpha sauteri</emphasis>
Bourgoin, 1931: 45
</taxonomicName>
(
<typeStatus id="CA0A04EDFFA5FFBF7B6FF9BEC486F98A" box="[626,671,1553,1575]" pageId="15" pageNumber="416">type</typeStatus>
locality: “Kosempo (
<collectingCountry id="6DA6FADFFFA5FFBF7A94F9BEC5F3F98A" box="[905,1002,1553,1575]" name="Taiwan" pageId="15" pageNumber="416">Formosa</collectingCountry>
)”); Mikšić 1977: 365, plate 6, fig.
<specimenCount id="03B771C6FFA5FFBF7C74F9BEC38CF98A" box="[1385,1429,1553,1575]" pageId="15" pageNumber="416" type="generic">
2
<emphasis id="27C5665DFFA5FFBF7C9FF9BEC38CF98A" box="[1410,1429,1553,1575]" italics="true" pageId="15" pageNumber="416"></emphasis>
</specimenCount>
; Masumoto &amp; Sakai 1988: 116, figs.
<specimenCount id="03B771C6FFA5FFBF7B2BF99FC44FF9EB" box="[566,598,1584,1606]" pageId="15" pageNumber="416" type="generic">
1
<emphasis id="27C5665DFFA5FFBF7B5EF99FC44FF9EB" box="[579,598,1584,1606]" italics="true" pageId="15" pageNumber="416"></emphasis>
</specimenCount>
, 2 ♀; Krajčik, 1998: 19; Sakai &amp; Nagai 1998: 235, plate 60, figs. 672-1
<specimenCount id="03B771C6FFA5FFBF7C27F99FC35EF9EB" box="[1338,1351,1584,1606]" pageId="15" pageNumber="416" type="generic">2</specimenCount>
♂, 672- 3 ♀; Yu
<emphasis id="27C5665DFFA5FFBF7801F9FFC750F9C9" box="[284,329,1614,1637]" italics="true" pageId="15" pageNumber="416">et al</emphasis>
. 1998: 95, plate 3, figs. 1ag
<emphasis id="27C5665DFFA5FFBF7B6BF9E1C490F9C9" box="[630,649,1614,1636]" italics="true" pageId="15" pageNumber="416"></emphasis>
, 1hj ♀; Smetana 2006: 301.
</paragraph>
</subSubSection>
<subSubSection id="5DABE9C4FFA5FFA0798AF93EC549FF78" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="description">
<paragraph id="150EBA4FFFA5FFBF798AF93EC79EF897" blockId="15.[151,1436,1681,2030]" pageId="15" pageNumber="416">
<emphasis id="27C5665DFFA5FFBF798AF93EC6C3F907" bold="true" box="[151,218,1681,1706]" pageId="15" pageNumber="416">Male.</emphasis>
TL: 21.525.0 mm, TW: 11.012.5 mm. Body usually brown or black (
<figureCitation id="8D8AA6CAFFA5FFBF7D1DF93EC2BEF907" box="[1024,1191,1681,1706]" captionStart="FIGURES 126 132" captionStartId="16.[151,264,1343,1366]" captionTargetBox="[171,1436,341,1318]" captionTargetId="figure@16.[163,1448,337,1323]" captionTargetPageId="16" captionText="FIGURES 126 132. Cosmiomorpha sauteri Bourgoin, 1931. 126 129. Male; 130 132. Female." httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="15" pageNumber="416">Figs. 126129</figureCitation>
). Body highly similar to
<taxonomicName id="D2B1C1CCFFA5FFBF79ABF919C733F963" box="[182,298,1717,1742]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFA5FFBF79ABF919C733F963" box="[182,298,1717,1742]" italics="true" pageId="15" pageNumber="416">C. decliva</emphasis>
</taxonomicName>
but dorsal surface brighter; scales much sparser and smaller; protibia slenderer; the deflexed tooth indistinct (
<figureCitation id="8D8AA6CAFFA5FFBF780DF976C785F95F" box="[272,412,1753,1778]" captionStart="FIGURES 48 60" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[160,1427,1181,1768]" captionTargetId="figure@6.[151,1436,1180,1774]" captionTargetPageId="6" captionText="FIGURES 48 60. Protibia and basitarsus of male Cosmiomorpha species in lateral view. 48. Cosmiomorpha modesta (Jiangsu); 49 51. Cosmiomorpha decliva: 49. (Sichuan); 50. (Guangxi); 51. (Fujian); 52 53. Cosmiomorpha sauteri (Taiwan); 54 56. Cosmiomorpha fortis new species: 54. Holotype; 55 56. Paratypes; 57. Cosmiomorpha nigripedis new species, holotype; 58 59. Cosmiomorpha maolanensis new species: 58. Holotype; 59. Paratype; 60. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247370/files/figure.png" pageId="15" pageNumber="416">Figs. 5253</figureCitation>
); the middle and proximal teeth on outer margin usually reduced or absent (
<figureCitation id="8D8AA6CAFFA5FFBF7C18F976C389F95F" box="[1285,1424,1753,1778]" captionStart="FIGURES 35 47" captionStartId="6.[151,264,985,1008]" captionTargetBox="[157,1430,446,959]" captionTargetId="figure@6.[151,1436,445,964]" captionTargetPageId="6" captionText="FIGURES 35 47. Protibia and basitarsus of male Cosmiomorpha species in dorsal view. 35 36. Cosmiomorpha modesta: 35. (Jiangsu); 36. (Guangxi); 37 38. Cosmiomorpha decliva: 37. (Sichuan); 38. (Guangxi); 39 40. Cosmiomorpha sauteri (Taiwan); 41 43. Cosmiomorpha fortis new species: 41. Holotype; 42 43. Paratypes; 44. Cosmiomorpha nigripedis new species, holotype; 45 46. Cosmiomorpha maolanensis new species: 45. Holotype; 46. Paratype; 47. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247369/files/figure.png" pageId="15" pageNumber="416">Figs. 3940</figureCitation>
). Mesometasternal process long with a constriction in the middle, apex truncated (
<figureCitation id="8D8AA6CAFFA5FFBF7D05F952C272F8BB" box="[1048,1131,1789,1814]" captionStart="FIGURES 1 20" captionStartId="4.[151,264,1561,1584]" captionTargetBox="[166,1436,195,1539]" captionTargetId="figure@4.[163,1448,193,1542]" captionTargetPageId="4" captionText="FIGURES 1 20. Mesosternal process of male Cosmiomorpha species. 1 4. Cosmiomorpha fortis new species: 1. Holotype; 2 4. Paratypes; 5. Cosmiomorpha nigripedis new species, holotype; 6. Cosmiomorpha maolanensis new species, holotype; 7. Cosmiomorpha cheni new species, holotype; 8 9. Cosmiomorpha modesta: 8. (Jiangsu); 9. (Guangxi); 10. Cosmiomorpha sauteri (Taiwan); 11 20. Cosmiomorpha decliva: 11. (Sichuan); 12. Cosmiomorpha squamulosa (Gansu), lectotype; 13. (Hunan); 14. (Sichuan); 15. (Guangxi); 16. (Fujian); 17. Cosmiomorpha baryi (N. Vietnam), “ allotype ”; 18. (Jiangxi); 19. (Fujian); 20. (Guangxi). Not to the same scale." httpUri="https://zenodo.org/record/247367/files/figure.png" pageId="15" pageNumber="416">Fig. 10</figureCitation>
). Parameres very similar to
<taxonomicName id="D2B1C1CCFFA5FFBF798AF88DC711F897" box="[151,264,1825,1850]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFA5FFBF798AF88DC711F897" box="[151,264,1825,1850]" italics="true" pageId="15" pageNumber="416">C. decliva</emphasis>
</taxonomicName>
(
<figureCitation id="8D8AA6CAFFA5FFBF7805F88EC763F897" box="[280,378,1825,1850]" captionStart="FIGURES 151 164" captionStartId="24.[151,264,1842,1865]" captionTargetBox="[161,1424,888,1807]" captionTargetId="figure@24.[151,1436,877,1822]" captionTargetPageId="24" captionText="FIGURES 151 164. Parameres of Cosmiomorpha species in dorsal and apical view. 151, 158. Cosmiomorpha modesta; 152, 159. Cosmiomorpha decliva; 153, 160. Cosmiomorpha sauteri; 154, 161. Cosmiomorpha fortis new species, holotype; 155, 162. Cosmiomorpha nigripedis new species, holotype; 156, 163. Cosmiomorpha maolanensis new species, holotype; 157, 164. Cosmiomorpha c heni new species, holotype." httpUri="https://zenodo.org/record/247381/files/figure.png" pageId="15" pageNumber="416">Fig. 153</figureCitation>
).
</paragraph>
<paragraph id="150EBA4FFFA5FFBF79DAF8EAC6F4F80B" blockId="15.[151,1436,1681,2030]" pageId="15" pageNumber="416">
<emphasis id="27C5665DFFA5FFBF79DAF8EAC73CF8F3" bold="true" box="[199,293,1861,1886]" pageId="15" pageNumber="416">Female.</emphasis>
TL: 21.021.5 mm, TW: 10.010.5 mm. Color similar to male (
<figureCitation id="8D8AA6CAFFA5FFBF7D3CF8EAC2D3F8F3" box="[1057,1226,1861,1886]" captionStart="FIGURES 126 132" captionStartId="16.[151,264,1343,1366]" captionTargetBox="[171,1436,341,1318]" captionTargetId="figure@16.[163,1448,337,1323]" captionTargetPageId="16" captionText="FIGURES 126 132. Cosmiomorpha sauteri Bourgoin, 1931. 126 129. Male; 130 132. Female." httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="15" pageNumber="416">Figs. 130132</figureCitation>
). Clypeus shorter, subrectangular. Mesometasternal process short and oval. Legs shorter, protibia with three large teeth on outer margin.
</paragraph>
<paragraph id="150EBA4FFFA5FFA079DAF81EC549FF78" blockId="15.[151,1436,1681,2030]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
<emphasis id="27C5665DFFA5FFBF79DAF81EC726F867" bold="true" box="[199,319,1969,1994]" pageId="15" pageNumber="416">Variation.</emphasis>
In both sexes, there are three general color forms: 1) a single color,
<emphasis id="27C5665DFFA5FFBF7D68F81DC284F864" box="[1141,1181,1970,1993]" italics="true" pageId="15" pageNumber="416">e.g.</emphasis>
, entirely brown, dark orange, reddish brown, black, or some intermediate color (
<figureCitation id="8D8AA6CAFFA5FFBF7A39F87AC221F843" box="[804,1080,2005,2030]" captionStart="FIGURES 126 132" captionStartId="16.[151,264,1343,1366]" captionTargetBox="[171,1436,341,1318]" captionTargetId="figure@16.[163,1448,337,1323]" captionTargetPageId="16" captionText="FIGURES 126 132. Cosmiomorpha sauteri Bourgoin, 1931. 126 129. Male; 130 132. Female." httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="15" pageNumber="416">Figs. 126, 128, 130, 132</figureCitation>
); 2) two or more colors, usually the pronotum is black and the elytra are brown; 3) has two or three markings on pronotum similar to
<taxonomicName id="D2B1C1CCFFBAFFA07C35FF37C385FF1D" box="[1320,1436,151,176]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07C35FF37C385FF1D" box="[1320,1436,151,176]" italics="true" pageId="16" pageNumber="417">C. decliva</emphasis>
</taxonomicName>
(
<figureCitation id="8D8AA6CAFFBAFFA07982FF13C71BFF79" box="[159,258,188,213]" captionStart="FIGURES 126 132" captionStartId="16.[151,264,1343,1366]" captionTargetBox="[171,1436,341,1318]" captionTargetId="figure@16.[163,1448,337,1323]" captionTargetPageId="16" captionText="FIGURES 126 132. Cosmiomorpha sauteri Bourgoin, 1931. 126 129. Male; 130 132. Female." httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="16" pageNumber="417">Fig. 129</figureCitation>
). Scales on dorsal surface vary in size (
<figureCitation id="8D8AA6CAFFBAFFA07BA7FF13C55DFF78" box="[698,836,188,213]" captionStart="FIGURES 61 75" captionStartId="7.[151,264,1256,1279]" captionTargetBox="[153,1433,194,1233]" captionTargetId="figure@7.[151,1436,193,1236]" captionTargetPageId="7" captionText="FIGURES 61 75. Scales on pronotal disc of male Cosmiomorpha species 61 62. Cosmiomorpha modesta (Zhejiang); 63 65. Cosmiomorpha decliva: 63. (Fujian); 64. (Sichuan); 65. (Fujian); 66 67. Cosmiomorpha sauteri (Taiwan); 68 72. Cosmiomorpha fortis new species: 68. Holotype; 69 72. Paratype; 73. Cosmiomorpha nigripedis new species, holotype; 74. Cosmiomorpha maolanensis new species, holotype; 75. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247371/files/figure.png" pageId="16" pageNumber="417">Figs. 6667</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DABE9C4FFBAFFA079DAFF70C406FEB0" pageId="16" pageNumber="417" type="materials_examined">
<paragraph id="150EBA4FFFBAFFA079DAFF70C406FEB0" blockId="16.[151,1436,151,285]" pageId="16" pageNumber="417">
<emphasis id="27C5665DFFBAFFA079DAFF70C7B2FF55" bold="true" box="[199,427,223,248]" pageId="16" pageNumber="417">Material examined</emphasis>
.
<emphasis id="27C5665DFFBAFFA078A7FF70C493FF55" bold="true" box="[442,650,223,248]" pageId="16" pageNumber="417">
<collectingCountry id="6DA6FADFFFBAFFA078A7FF70C405FF55" box="[442,540,223,248]" name="China" pageId="16" pageNumber="417">CHINA</collectingCountry>
:
<collectingCountry id="6DA6FADFFFBAFFA07B37FF70C49FFF55" box="[554,646,223,248]" name="Taiwan" pageId="16" pageNumber="417">Taiwan</collectingCountry>
:
</emphasis>
3♂♂, 2♀♀ (QCCC),
<date id="610F9C8FFFBAFFA07A94FF70C20CFF55" box="[905,1045,223,248]" pageId="16" pageNumber="417" value="2005-07-27">27.VII.2005</date>
, Mamei, Jianshi, Hsinchu County, alt.
<quantity id="D24917AAFFBAFFA079DCFEABC701FEB1" box="[193,280,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="417" unit="m" value="1400.0">1400 m</quantity>
, Chang-Chin Chen leg.
</paragraph>
</subSubSection>
<caption id="41CEEAC7FFBAFFA0798AFA90C273FAF8" box="[151,1130,1343,1366]" httpUri="https://zenodo.org/record/247375/files/figure.png" pageId="16" pageNumber="417" targetBox="[171,1436,341,1318]" targetPageId="16">
<paragraph id="150EBA4FFFBAFFA0798AFA90C273FAF8" blockId="16.[151,1130,1343,1366]" box="[151,1130,1343,1366]" pageId="16" pageNumber="417">
<emphasis id="27C5665DFFBAFFA0798AFA90C72DFAF9" bold="true" box="[151,308,1343,1366]" pageId="16" pageNumber="417">FIGURES 126</emphasis>
<emphasis id="27C5665DFFBAFFA0785DFA90C775FAF9" bold="true" box="[320,364,1343,1364]" pageId="16" pageNumber="417">132.</emphasis>
<taxonomicName id="D2B1C1CCFFBAFFA0786EFA90C51DFAF8" authority="Bourgoin, 1931" authorityName="Bourgoin" authorityYear="1931" box="[371,772,1343,1366]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA0786EFA90C442FAFB" box="[371,603,1343,1366]" italics="true" pageId="16" pageNumber="417">Cosmiomorpha sauteri</emphasis>
Bourgoin, 1931
</taxonomicName>
. 126129. Male; 130132. Female.
</paragraph>
</caption>
<subSubSection id="5DABE9C4FFBAFFA079DAFAD3C40BFA38" box="[199,530,1404,1429]" pageId="16" pageNumber="417" type="distribution">
<paragraph id="150EBA4FFFBAFFA079DAFAD3C40BFA38" blockId="16.[151,1437,1404,1934]" box="[199,530,1404,1429]" pageId="16" pageNumber="417">
<emphasis id="27C5665DFFBAFFA079DAFAD3C779FA38" bold="true" box="[199,352,1404,1429]" pageId="16" pageNumber="417">Distribution.</emphasis>
<collectingCountry id="6DA6FADFFFBAFFA0787AFAD3C7B7FA38" box="[359,430,1404,1429]" name="China" pageId="16" pageNumber="417">China</collectingCountry>
:
<collectingCountry id="6DA6FADFFFBAFFA078A4FAD3C416FA38" box="[441,527,1404,1429]" name="Taiwan" pageId="16" pageNumber="417">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="5DABE9C4FFBAFFA079DAFA0FC577F823" pageId="16" pageNumber="417" type="discussion">
<paragraph id="150EBA4FFFBAFFA079DAFA0FC5CAF988" blockId="16.[151,1437,1404,1934]" pageId="16" pageNumber="417">
<emphasis id="27C5665DFFBAFFA079DAFA0FC722FA14" bold="true" box="[199,315,1440,1465]" pageId="16" pageNumber="417">Remarks.</emphasis>
This species was originally described by Bourgoin (1931) based on a single female from
<collectingCountry id="6DA6FADFFFBAFFA07C5EFA0FC380FA14" box="[1347,1433,1440,1465]" name="Taiwan" pageId="16" pageNumber="417">Taiwan</collectingCountry>
. The
<typeStatus id="CA0A04EDFFBAFFA079CDFA6BC72BFA70" box="[208,306,1476,1501]" pageId="16" pageNumber="417" type="holotype">holotype</typeStatus>
(automatically fixed by monotypy) was not found in BMNH. The
<typeStatus id="CA0A04EDFFBAFFA07D7BFA69C28FFA73" box="[1126,1174,1478,1502]" pageId="16" pageNumber="417">type</typeStatus>
locality is
<emphasis id="27C5665DFFBAFFA07C38FA6BC32AFA70" bold="true" box="[1317,1331,1476,1501]" pageId="16" pageNumber="417"></emphasis>
<collectingCountry id="6DA6FADFFFBAFFA07C2EFA69C381FA70" box="[1331,1432,1478,1501]" name="Taiwan" pageId="16" pageNumber="417">Formosa</collectingCountry>
, Kosempo
<emphasis id="27C5665DFFBAFFA0781FFA47C709F9AC" bold="true" box="[258,272,1512,1537]" pageId="16" pageNumber="417"></emphasis>
(now Jiaxian, Kaohsiung), other localities, such as Taoyuan County, Hualien County, and Nantou County were recorded by Masumoto &amp; Sakai (1988) and Yu
<emphasis id="27C5665DFFBAFFA07A5CF9A1C568F98B" box="[833,881,1549,1574]" italics="true" pageId="16" pageNumber="417">et al</emphasis>
. (1988).
</paragraph>
<paragraph id="150EBA4FFFBAFFA079DAF99FC577F823" blockId="16.[151,1437,1404,1934]" pageId="16" pageNumber="417">
This species is probably the most variable of this subgenus (Sakai &amp; Nagai, 1988; Yu
<emphasis id="27C5665DFFBAFFA07DBDF99DC2CAF9E4" box="[1184,1235,1584,1609]" italics="true" pageId="16" pageNumber="417">et al</emphasis>
. 1988). The third color form mentioned above is very similar to
<taxonomicName id="D2B1C1CCFFBAFFA07BB9F9FAC516F9C3" box="[676,783,1621,1646]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07BB9F9FAC516F9C3" box="[676,783,1621,1646]" italics="true" pageId="16" pageNumber="417">C.decliva</emphasis>
</taxonomicName>
, and it is difficult to distinguish between them, unless the specimens locality is known,
<taxonomicName id="D2B1C1CCFFBAFFA078FBF9D6C44AF93C" box="[486,595,1657,1681]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA078FBF9D6C44AF93C" box="[486,595,1657,1681]" italics="true" pageId="16" pageNumber="417">C. sauteri</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="6DA6FADFFFBAFFA07BE9F9D7C55FF93C" box="[756,838,1656,1681]" name="Taiwan" pageId="16" pageNumber="417">Taiwan</collectingCountry>
while
<taxonomicName id="D2B1C1CCFFBAFFA07A8CF9D6C21BF93C" box="[913,1026,1656,1681]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07A8CF9D6C21BF93C" box="[913,1026,1656,1681]" italics="true" pageId="16" pageNumber="417">C. decliva</emphasis>
</taxonomicName>
is distributed in mainland
<collectingCountry id="6DA6FADFFFBAFFA07C37F9D7C374F93C" box="[1322,1389,1656,1681]" name="China" pageId="16" pageNumber="417">China</collectingCountry>
and northern
<collectingCountry id="6DA6FADFFFBAFFA079E3F932C778F91B" box="[254,353,1693,1718]" name="Vietnam" pageId="16" pageNumber="417">Vietnam</collectingCountry>
. However, a pair of
<taxonomicName id="D2B1C1CCFFBAFFA07B54F931C4A5F91B" box="[585,700,1693,1718]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07B54F931C445F918" box="[585,604,1694,1717]" italics="true" pageId="16" pageNumber="417">C</emphasis>
.
<emphasis id="27C5665DFFBAFFA07B76F932C4A5F91B" box="[619,700,1693,1718]" italics="true" pageId="16" pageNumber="417">decliva</emphasis>
</taxonomicName>
collected from Fuzhou (closest to
<collectingCountry id="6DA6FADFFFBAFFA07D55F932C286F91B" box="[1096,1183,1693,1718]" name="Taiwan" pageId="16" pageNumber="417">Taiwan</collectingCountry>
), are interesting (
<figureCitation id="8D8AA6CAFFBAFFA07C7EF932C6F8F974" captionStart="FIGURES 89 96" captionStartId="11.[151,264,1889,1912]" captionTargetBox="[169,1426,632,1858]" captionTargetId="figure@11.[151,1436,625,1869]" captionTargetPageId="11" captionText="FIGURES 89 96. Cosmiomorpha decliva Janson, 1890. 89 96. Male: 89 90. (Sichuan); 91. (Shanxi); 92. (Henan); 93. (Fujian); 94 96. Female: 94 95. (Fujian); 96. (Henan)." httpUri="https://zenodo.org/record/247372/files/figure.png" pageId="16" pageNumber="417">Figs. 9395</figureCitation>
). Although the male is of large size, the deflexed tooth (
<figureCitation id="8D8AA6CAFFBAFFA07A9FF96FC5C1F974" box="[898,984,1728,1753]" captionStart="FIGURES 48 60" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[160,1427,1181,1768]" captionTargetId="figure@6.[151,1436,1180,1774]" captionTargetPageId="6" captionText="FIGURES 48 60. Protibia and basitarsus of male Cosmiomorpha species in lateral view. 48. Cosmiomorpha modesta (Jiangsu); 49 51. Cosmiomorpha decliva: 49. (Sichuan); 50. (Guangxi); 51. (Fujian); 52 53. Cosmiomorpha sauteri (Taiwan); 54 56. Cosmiomorpha fortis new species: 54. Holotype; 55 56. Paratypes; 57. Cosmiomorpha nigripedis new species, holotype; 58 59. Cosmiomorpha maolanensis new species: 58. Holotype; 59. Paratype; 60. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247370/files/figure.png" pageId="16" pageNumber="417">Fig. 51</figureCitation>
) is short with two tips, just like some individuals of
<taxonomicName id="D2B1C1CCFFBAFFA07825F949C7BEF950" box="[312,423,1766,1789]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA07825F949C7BEF950" box="[312,423,1766,1789]" italics="true" pageId="16" pageNumber="417">C. sauteri</emphasis>
</taxonomicName>
(
<figureCitation id="8D8AA6CAFFBAFFA078ABF94AC413F953" box="[438,522,1765,1790]" captionStart="FIGURES 48 60" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[160,1427,1181,1768]" captionTargetId="figure@6.[151,1436,1180,1774]" captionTargetPageId="6" captionText="FIGURES 48 60. Protibia and basitarsus of male Cosmiomorpha species in lateral view. 48. Cosmiomorpha modesta (Jiangsu); 49 51. Cosmiomorpha decliva: 49. (Sichuan); 50. (Guangxi); 51. (Fujian); 52 53. Cosmiomorpha sauteri (Taiwan); 54 56. Cosmiomorpha fortis new species: 54. Holotype; 55 56. Paratypes; 57. Cosmiomorpha nigripedis new species, holotype; 58 59. Cosmiomorpha maolanensis new species: 58. Holotype; 59. Paratype; 60. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247370/files/figure.png" pageId="16" pageNumber="417">Fig. 53</figureCitation>
). The large dark area on the pronotal disc typical of
<taxonomicName id="D2B1C1CCFFBAFFA07D4EF949C2DCF953" box="[1107,1221,1765,1790]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07D4EF949C27FF950" box="[1107,1126,1766,1789]" italics="true" pageId="16" pageNumber="417">C</emphasis>
.
<emphasis id="27C5665DFFBAFFA07D69F94AC2DCF953" box="[1140,1221,1765,1790]" italics="true" pageId="16" pageNumber="417">decliva</emphasis>
</taxonomicName>
from other regions is much more diffuse and nearly absent in these specimens. Furthermore, the scales are much smaller on the dorsal surface in this pair (
<figureCitation id="8D8AA6CAFFBAFFA07864F882C7D4F8EB" box="[377,461,1837,1862]" captionStart="FIGURES 61 75" captionStartId="7.[151,264,1256,1279]" captionTargetBox="[153,1433,194,1233]" captionTargetId="figure@7.[151,1436,193,1236]" captionTargetPageId="7" captionText="FIGURES 61 75. Scales on pronotal disc of male Cosmiomorpha species 61 62. Cosmiomorpha modesta (Zhejiang); 63 65. Cosmiomorpha decliva: 63. (Fujian); 64. (Sichuan); 65. (Fujian); 66 67. Cosmiomorpha sauteri (Taiwan); 68 72. Cosmiomorpha fortis new species: 68. Holotype; 69 72. Paratype; 73. Cosmiomorpha nigripedis new species, holotype; 74. Cosmiomorpha maolanensis new species, holotype; 75. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247371/files/figure.png" pageId="16" pageNumber="417">Fig. 65</figureCitation>
), the same as in
<taxonomicName id="D2B1C1CCFFBAFFA07B97F881C4E0F8E8" box="[650,761,1838,1861]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA07B97F881C4E0F8E8" box="[650,761,1838,1861]" italics="true" pageId="16" pageNumber="417">C. sauteri</emphasis>
</taxonomicName>
(
<figureCitation id="8D8AA6CAFFBAFFA07A14F882C547F8EB" box="[777,862,1837,1862]" captionStart="FIGURES 61 75" captionStartId="7.[151,264,1256,1279]" captionTargetBox="[153,1433,194,1233]" captionTargetId="figure@7.[151,1436,193,1236]" captionTargetPageId="7" captionText="FIGURES 61 75. Scales on pronotal disc of male Cosmiomorpha species 61 62. Cosmiomorpha modesta (Zhejiang); 63 65. Cosmiomorpha decliva: 63. (Fujian); 64. (Sichuan); 65. (Fujian); 66 67. Cosmiomorpha sauteri (Taiwan); 68 72. Cosmiomorpha fortis new species: 68. Holotype; 69 72. Paratype; 73. Cosmiomorpha nigripedis new species, holotype; 74. Cosmiomorpha maolanensis new species, holotype; 75. Cosmiomorpha cheni new species, holotype." httpUri="https://zenodo.org/record/247371/files/figure.png" pageId="16" pageNumber="417">Fig. 67</figureCitation>
). Due to these similar characters, it is possible that
<taxonomicName id="D2B1C1CCFFBAFFA0798AF8FEC711F8C4" box="[151,264,1873,1897]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA0798AF8FEC711F8C4" box="[151,264,1873,1897]" italics="true" pageId="16" pageNumber="417">C. sauteri</emphasis>
</taxonomicName>
should be considered a subspecies or merely a synonym of
<taxonomicName id="D2B1C1CCFFBAFFA07ADBF8FEC222F8C4" box="[966,1083,1872,1897]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="decliva">
<emphasis id="27C5665DFFBAFFA07ADBF8FEC222F8C4" box="[966,1083,1872,1897]" italics="true" pageId="16" pageNumber="417">C. decliva</emphasis>
</taxonomicName>
. To verify this hypothesis, the
<typeStatus id="CA0A04EDFFBAFFA0798AF8DAC6E0F823" box="[151,249,1909,1934]" pageId="16" pageNumber="417" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D2B1C1CCFFBAFFA07803F8D9C794F820" box="[286,397,1910,1933]" class="Insecta" family="Cetoniidae" genus="Cosmiomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="27C5665DFFBAFFA07803F8D9C794F820" box="[286,397,1910,1933]" italics="true" pageId="16" pageNumber="417">C. sauteri</emphasis>
</taxonomicName>
and more specimens need to be examined.
</paragraph>
</subSubSection>
</treatment>
</document>