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<document id="6D3D8726EF23753CB9A34233C1E784A1" ID-DOI="10.21248/contrib.entomol.66.2.329-346" ID-ISSN="0005-805X" ID-Zenodo-Dep="4754143" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620838964136" checkinUser="carolina" docAuthor="Prokofiev, Artem M. &amp; Zorn, Carsten" docDate="2016" docId="7A52E960FF8E5014FCBFFE37FEC4A82F" docLanguage="en" docName="ContribEntomol.66.2.329-346.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1909" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Mimela krasava Prokofiev &amp; Zorn 2016, spec. nov." docType="treatment" docVersion="3" lastPageNumber="338" masterDocId="866B9118FF86501DFFDAFFC4FFCBA00E" masterDocTitle="Review of the Mimela species of the Dalat Plateau in southern Vietnam (Coleoptera, Scarabaeidae, Rutelinae)" masterLastPageNumber="346" masterPageNumber="329" pageNumber="337" updateTime="1698916946317" updateUser="plazi">
<mods:mods id="EAA0503FE3E0F226E038AC63BCEB6CF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07DA6B461D8F467D8A1AAA790B57738A">
<mods:title id="73E2CCD1B28E1A6D8E8B9CB265B28E29">Review of the Mimela species of the Dalat Plateau in southern Vietnam (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="4ACC33769C7C274F1B2741BA92DDD989" type="personal">
<mods:role id="AA8644DEFDDE37E506A0BDC39D559E52">
<mods:roleTerm id="9272E0E9A799BC860F30E3BEE4FDDA4D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="605BCB9C1CF3F6D6B5F1EBA15BA1B873">Prokofiev, Artem M.</mods:namePart>
</mods:name>
<mods:name id="44A664DBCDDDE0D5A37B7AC907B346DA" type="personal">
<mods:role id="B6AE4338A0B2DB57E57FA5F21DF9E5B3">
<mods:roleTerm id="49B02D522DB6E03BE0395E4EA02E5EC0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41B91B883FF96CF481F68DB573BF50FB">Zorn, Carsten</mods:namePart>
</mods:name>
<mods:typeOfResource id="EF72A204487B9A2B220500005130A063">text</mods:typeOfResource>
<mods:relatedItem id="28CA6160719DDD9D77C919F0C1D9E618" type="host">
<mods:titleInfo id="2BC7E25EEA44C37941D748539EF67142">
<mods:title id="D8178F8BD14984D5883D04362A464569">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="731255AFE306C10FE06D4352E5B6A742">
<mods:date id="194FD26CAC27C826BC692E15C4C7350B">2016</mods:date>
<mods:detail id="9807DE788E0CD4629A23FEE12FB73C00" type="pubDate">
<mods:number id="A7D7AF3D526349DBDB31006F959EF889">2016-12-20</mods:number>
</mods:detail>
<mods:detail id="69A5C1E264C392AF2848BB68F7F7B31F" type="volume">
<mods:number id="95BF3FF80A7D76FC872885ACEF5D3812">66</mods:number>
</mods:detail>
<mods:detail id="0465945519D8DFF5B4D9CFD32664FCA6" type="issue">
<mods:number id="166C74DEC376CD90E973BFD82CD26036">2</mods:number>
</mods:detail>
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<mods:start id="83F20CA8E2CA741D0E97FB9EEFF9A31E">329</mods:start>
<mods:end id="400A5BC08D0DC59D648154AE9AB71F10">346</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="B4263456CB3FFB775A422131530F0453">
<mods:url id="28B5B312BB72EB82E74A0B6E996C2730">https://www.contributions-to-entomology.org/article/view/1909</mods:url>
</mods:location>
<mods:classification id="F261FFA8F99063C7B6FE440DB8B317FF">journal article</mods:classification>
<mods:identifier id="C2D74B50E9475BC20099E6824B6F98FD" type="DOI">10.21248/contrib.entomol.66.2.329-346</mods:identifier>
<mods:identifier id="32F92B36D3DDBDDE66E7F2A7281C5EBA" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="6E197BF2CFC152ED3A3996B8EE2B300F" type="Zenodo-Dep">4754143</mods:identifier>
</mods:mods>
<treatment id="7A52E960FF8E5014FCBFFE37FEC4A82F" LSID="urn:lsid:plazi:treatment:7A52E960FF8E5014FCBFFE37FEC4A82F" httpUri="http://treatment.plazi.org/id/7A52E960FF8E5014FCBFFE37FEC4A82F" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337">
<subSubSection id="BAE10BFDFF8E5015FCBFFE37FBBCA200" box="[869,1143,499,526]" pageId="8" pageNumber="337" type="nomenclature">
<paragraph id="F2445876FF8E5015FCBFFE37FBBCA200" blockId="8.[869,1143,499,559]" box="[869,1143,499,526]" pageId="8" pageNumber="337">
<heading id="A90CEF1AFF8E5015FCBFFE37FBBCA200" box="[869,1143,499,526]" centered="true" fontSize="12" level="2" pageId="8" pageNumber="337" reason="2">
<taxonomicName id="35FB23F5FF8E5015FCBFFE37FBC3A200" authority="Prokofiev &amp; Zorn, 2016" authorityName="Prokofiev &amp; Zorn" authorityYear="2016" box="[869,1032,499,526]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="337" phylum="Arthropoda" rank="species" species="krasava" status="sp. nov.">
<emphasis id="C08F8464FF8E5015FCBFFE37FBC3A200" box="[869,1032,499,526]" italics="true" pageId="8" pageNumber="337">Mimela krasava</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DBBC391FFF8E5015FBD4FE37FBBCA200" box="[1038,1143,499,526]" pageId="8" pageNumber="337" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8E5015FCBFFDDDFC28A221" box="[869,995,537,559]" pageId="8" pageNumber="337" type="description">
<paragraph id="F2445876FF8E5015FCBFFDDDFC28A221" blockId="8.[869,1143,499,559]" box="[869,995,537,559]" pageId="8" pageNumber="337">
(
<figureCitation id="6AC044F3FF8E5015FCB0FDDDFC54A221" box="[874,927,537,559]" captionStart="Figs 1–14" captionStartId="5.[151,191,1895,1917]" captionTargetBox="[168,1383,245,1865]" captionTargetId="figure-147@5.[151,1398,232,1884]" captionTargetPageId="5" captionText="Figs 1–14: Mimela spp.– 1, 2 M. amphichroma spec. nov.(1 greenform,holotype; 2 violet form, paratype,male).3 M. campyloclados spec. nov., holotype. 4 M. dalatocoerulea spec. nov., holotype. 5 M. dehaani, male, Vietnam: Lamdong prov. 6 M. galbinata spec. nov., holotype. 7 M. germana spec. nov., holotype. 8 M. indosinica spec. nov., holotype. 9 M. krasava spec. nov., holotype. 10 M. langbianica spec. nov., holotype. 11 M. pacholatkoi spec. nov., holotype. 12 M. signaticollis, male, Vietnam: Nha Trang. 13, 14 M. vietnamensis spec. nov. (13. male, holotype. 14. female, paratype)." figureDoi="http://doi.org/10.5281/zenodo.4754147" httpUri="https://zenodo.org/record/4754147/files/figure.png" pageId="8" pageNumber="337">Figs 9</figureCitation>
,
<figureCitation id="6AC044F3FF8E5015FC7DFDDDFC15A221" box="[935,990,537,559]" captionStart="Figs 44–60" captionStartId="16.[208,248,1893,1915]" captionTargetBox="[224,1401,251,1864]" captionTargetId="figure-41@16.[208,1418,239,1884]" captionTargetPageId="16" captionText="Figs 44–60: Mimela spp., aedeagi. – 44–46: M. krasava spec. nov., holotype (44 lateral view. 45 parameres, dorsal view. 46 parameres, ventral view). 47–50: M. langbianica spec. nov., holotype (47 lateral view, left side. 48 lateral view of parameres, right side. 49 dorsal view of parameres. 50 ventral view of parameres). 51–53: M. pacholatkoi spec. nov., holotype (51 lateral view. 52 parameres, dorsal view. 53 parameres, ventral view). 54 M. ohausi, Laos: Xieng Khouang prov., lateral view. 55–57: M. signaticollis, Vietnam: Nha Trang (55 lateral view. 56 parameres, dorsal view. 57 parameres, ventral view). 58–60: M. vietnamensis spec. nov., holotype (58 lateral view. 59 parameres, dorsal view. 60 parameres, ventral view). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4754155" httpUri="https://zenodo.org/record/4754155/files/figure.png" pageId="8" pageNumber="337">44–46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8E5015FCE5FD8DFBC7A34F" pageId="8" pageNumber="337" type="materials_examined">
<paragraph id="F2445876FF8E5015FCE5FD8DFBC7A2CF" blockId="8.[831,1437,585,834]" pageId="8" pageNumber="337">
<materialsCitation id="4293522BFF8E5015FCE5FD8DFBC7A2CF" collectingDate="2012-06-05" collectionCode="CAP" collectorName="A. M. Prokofiev" country="Vietnam" elevation="775" latitude="12.235556" location="Khanh Vinh distr." longLatPrecision="21" longitude="108.77056" pageId="8" pageNumber="337" specimenCount="2" stateProvince="Khanh Hoa" typeStatus="holotype">
<emphasis id="C08F8464FF8E5015FCE5FD8DFC6EA26F" bold="true" box="[831,933,585,609]" pageId="8" pageNumber="337">
<typeStatus id="2D40E6D4FF8E5015FCE5FD8DFC6EA26F" box="[831,933,585,609]" pageId="8" pageNumber="337" type="holotype">Holotype</typeStatus>
</emphasis>
:  “
<collectingCountry id="8AEC18E6FF8E5015FC0EFD8DFB9BA26F" box="[980,1104,585,609]" name="Vietnam" pageId="8" pageNumber="337">VIETNAM</collectingCountry>
,
<collectingRegion id="303F9694FF8E5015FB83FD8DFADBA26F" box="[1113,1296,585,609]" country="Vietnam" name="Khanh Hoa" pageId="8" pageNumber="337">Khanh Hoa prov.</collectingRegion>
,
<location id="F7240EADFF8E5015FAC3FD8DFCBCA28F" LSID="urn:lsid:plazi:treatment:7A52E960FF8E5014FCBFFE37FEC4A82F:F7240EADFF8E5015FAC3FD8DFCBCA28F" country="Vietnam" latitude="12.235556" longLatPrecision="21" longitude="108.77056" name="Khanh Vinh distr." pageId="8" pageNumber="337" stateProvince="Khanh Hoa">Khanh Vinh distr.</location>
, ~
<locationDeviation id="E3F64154FF8E5015FC78FDADFB8CA28F" box="[930,1095,617,641]" location="Dalat" pageId="8" pageNumber="337">
<quantity id="3503F593FF8E5015FC78FDADFC2FA28F" box="[930,996,617,641]" metricMagnitude="4" metricUnit="m" metricValue="7.2" pageId="8" pageNumber="337" unit="km" value="72.0">72 km</quantity>
E Dalat
</locationDeviation>
,
<geoCoordinate id="97CF3EB1FF8E5015FB8DFDADFB1DA28F" box="[1111,1238,617,641]" degrees="12" direction="north" minutes="14" orientation="latitude" pageId="8" pageNumber="337" precision="15" seconds="08" value="12.235556">12°14'08&quot; N</geoCoordinate>
,
<geoCoordinate id="97CF3EB1FF8E5015FB3FFDADFAA7A28F" box="[1253,1388,617,641]" degrees="108" direction="east" minutes="46" orientation="longitude" pageId="8" pageNumber="337" precision="15" seconds="14" value="108.77056">108°46'14&quot; E</geoCoordinate>
, alt.
<quantity id="3503F593FF8E5015FCE5FD4DFC73A2AF" box="[831,952,649,673]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="8" pageNumber="337" unit="m" value="775.0" valueMax="800.0" valueMin="750.0">
<elevation id="79D6BF45FF8E5015FCE5FD4DFC73A2AF" box="[831,952,649,673]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="8" pageNumber="337" unit="m" value="775.0" valueMax="800.0" valueMin="750.0">750–800 m</elevation>
</quantity>
, at light,
<date id="86457EB6FF8E5015FBCCFD4DFBB9A2AF" box="[1046,1138,649,673]" pageId="8" pageNumber="337" value="2012-06-05">
<collectingDate id="9601875EFF8E5015FBCCFD4DFBB9A2AF" box="[1046,1138,649,673]" pageId="8" pageNumber="337" value="2012-06-05">5.6.2012</collectingDate>
</date>
,
<collectorName id="5F0E3DA0FF8E5015FBA1FD4DFAD6A2AF" box="[1147,1309,649,673]" pageId="8" pageNumber="337">A.M. Prokofiev</collectorName>
leg. | [
<typeStatus id="2D40E6D4FF8E5015FABAFD4DFCA6A2CC" pageId="8" pageNumber="337" type="holotype">holotype</typeStatus>
label]” (
<collectionCode id="94EAC0B3FF8E5015FC1DFD6DFBCBA2CF" box="[967,1024,681,705]" pageId="8" pageNumber="337">CAP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F2445876FF8E5015FCE5FD0DFBC7A34F" blockId="8.[831,1437,585,834]" pageId="8" pageNumber="337">
<materialsCitation id="4293522BFF8E5015FCE5FD0DFBC3A34F" collectingDate="2012-06-04" collectionCode="CAP" collectorName="A. M. Prokofiev" country="Vietnam" elevation="916" latitude="12.2237" location="Khanh Vinh distr." longLatPrecision="1" longitude="108.75892" pageId="8" pageNumber="337" specimenCount="2" stateProvince="Khanh Hoa" typeStatus="paratype">
<emphasis id="C08F8464FF8E5015FCE5FD0DFC6BA2EF" bold="true" box="[831,928,713,737]" pageId="8" pageNumber="337">
<typeStatus id="2D40E6D4FF8E5015FCE5FD0DFC6BA2EF" box="[831,928,713,737]" pageId="8" pageNumber="337" type="paratype">Paratype</typeStatus>
</emphasis>
:  “
<collectingCountry id="8AEC18E6FF8E5015FC0AFD0DFB87A2EF" box="[976,1100,713,737]" name="Vietnam" pageId="8" pageNumber="337">VIETNAM</collectingCountry>
,
<collectingRegion id="303F9694FF8E5015FB8FFD0DFAC4A2EF" box="[1109,1295,713,737]" country="Vietnam" name="Khanh Hoa" pageId="8" pageNumber="337">Khanh Hoa prov.</collectingRegion>
,
<location id="F7240EADFF8E5015FAC2FD0DFCBCA30F" LSID="urn:lsid:plazi:treatment:7A52E960FF8E5014FCBFFE37FEC4A82F:F7240EADFF8E5015FAC2FD0DFCBCA30F" country="Vietnam" latitude="12.2237" longLatPrecision="1" longitude="108.75892" name="Khanh Vinh distr." pageId="8" pageNumber="337" stateProvince="Khanh Hoa">Khanh Vinh distr.</location>
, ~
<quantity id="3503F593FF8E5015FC41FD2DFC1AA30F" box="[923,977,745,769]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="337" unit="km" value="3.0">3 km</quantity>
above felling,
<geoCoordinate id="97CF3EB1FF8E5015FBABFD2DFB31A30F" box="[1137,1274,745,769]" degrees="12" direction="north" minutes="13.422" orientation="latitude" pageId="8" pageNumber="337" precision="1" value="12.2237">12°13.422' N</geoCoordinate>
,
<geoCoordinate id="97CF3EB1FF8E5015FADDFD2DFA53A30F" box="[1287,1432,745,769]" degrees="108" direction="east" minutes="45.535" orientation="longitude" pageId="8" pageNumber="337" precision="1" value="108.75892">108°45.535' E</geoCoordinate>
, alt.
<quantity id="3503F593FF8E5015FCBDFCCDFC67A32F" box="[871,940,777,801]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="8" pageNumber="337" unit="m" value="916.0">
<elevation id="79D6BF45FF8E5015FCBDFCCDFC67A32F" box="[871,940,777,801]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="8" pageNumber="337" unit="m" value="916.0">916 m</elevation>
</quantity>
, at light,
<date id="86457EB6FF8E5015FBD5FCCDFBA1A32F" box="[1039,1130,777,801]" pageId="8" pageNumber="337" value="2012-06-04">
<collectingDate id="9601875EFF8E5015FBD5FCCDFBA1A32F" box="[1039,1130,777,801]" pageId="8" pageNumber="337" value="2012-06-04">4.6.2012</collectingDate>
</date>
,
<collectorName id="5F0E3DA0FF8E5015FBAFFCCDFAD2A32F" box="[1141,1305,777,801]" pageId="8" pageNumber="337">A.M. Prokofiev</collectorName>
leg. | [
<typeStatus id="2D40E6D4FF8E5015FABBFCCEFCA6A34C" pageId="8" pageNumber="337" type="paratype">paratype</typeStatus>
label]” (
<collectionCode id="94EAC0B3FF8E5015FC1DFCEDFBCBA34F" box="[967,1024,809,833]" pageId="8" pageNumber="337">CAP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8E5014FCE5FCADFEABA5AF" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337" type="description">
<paragraph id="F2445876FF8E5015FCE5FCADFCB9A4CF" blockId="8.[831,1437,873,2081]" pageId="8" pageNumber="337">
<emphasis id="C08F8464FF8E5015FCE5FCADFC0AA38F" bold="true" box="[831,961,873,897]" pageId="8" pageNumber="337">Description</emphasis>
(
<typeStatus id="2D40E6D4FF8E5015FC09FCADFBFCA38F" box="[979,1079,873,897]" pageId="8" pageNumber="337" type="holotype">holotype</typeStatus>
): Length
<quantity id="3503F593FF8E5015FB7BFCADFB3EA38F" box="[1185,1269,873,897]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="337" unit="mm" value="8.5">8.5 mm</quantity>
, greatest width
<quantity id="3503F593FF8E5015FCE5FC4DFC58A3AF" box="[831,915,905,929]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="337" unit="mm" value="4.5">4.5 mm</quantity>
. Reddish-brown above, pronotum with strong dark-green tint and broad yellowish side margins; apical protuberances of elytra with oblong blackish marks descending toward side border; sterna and pygidium testaceous, abdomen much darker, reddish-brown; legs testaceous, except most parts of protibia and upper and inner parts of meso- and metatibia, which are metallic green-brown, tarsi reddish-brown to blackish, hind tarsi darker; antennae reddish-brown, club darker; pilosity pale.
</paragraph>
<paragraph id="F2445876FF8E5014FCE5FB0DFD16A1EF" blockId="8.[831,1437,873,2081]" lastBlockId="9.[151,757,231,1217]" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337">
Clypeus nearly trapezoidal, with sides convergent anteriad, anterior angles broadly rounded, not distinct; anterior margin straight,weakly raised,narrow in frontal view. Outer part along outer margin distinctly impressed, finely punctured, convex central and basal parts of clypeus coarsely, rugosely punctate. Fronto-clypeal suture distinct. Inner margin of eye with fine marginal line, with a single erect seta inserted before its mid-length on each side (perhaps other setae worn off). Eyes somewhat bulging. Anterior half of frons rugosely punctate, slightly impressed at center, punctures becoming smaller and much sparser toward vertex. Pronotum 1.9 times broader than long, sides almost parallel in basal half, distinctly convergent in anterior half; anterior angles acute; posterior angles obtuse, rounded. Anterior margin of pronotum with complete marginal line, concave; posterior margin of pronotum strongly convex in middle third, straightened in outer thirds, basis with complete marginal line. Sides of pronotum with few long erect distant hairs. Pronotum regularly and distinctly, moderately dense punctured, spaces between punctures on disc more or less equal with their diameter, punctures becoming smaller toward the margins. Scutellum subtriangular with apex bluntly pointed, irregularly punctured; punctures similar to those on the disc of pronotum. Elytra with distinct punctate rows having deep simple punctures and slightly convex interspaces; second interspace with irregular punctation narrowed into a single row in posteri- ormost quarter, 4
<superScript id="058EF53EFF8F5014FE90FF23FE9CA0FB" attach="none" box="[330,343,231,245]" fontSize="6" pageId="9" pageNumber="338">th</superScript>
, 6
<superScript id="058EF53EFF8F5014FEB5FF23FEB7A0FB" attach="left" box="[367,380,231,245]" fontSize="6" pageId="9" pageNumber="338">th</superScript>
and 8
<superScript id="058EF53EFF8F5014FE60FF23FE0CA0FB" attach="left" box="[442,455,231,245]" fontSize="6" pageId="9" pageNumber="338">th</superScript>
interspaces with a single row of punctures; punctures in even interspaces the same as in punctate rows; odd interspaces with minute somewhat irregular punctures. Sides of elytra with marginal line, with elytral epipleura thickened under humeral umbo, bearing short distant setae, longer and more closely spaced under humeral umbo. Pygidium weakly convex, rugosely punctate, with a double row of rather long erect hairs at apex.
</paragraph>
<paragraph id="F2445876FF8F5014FF4DFE2DFDB0A36F" blockId="9.[151,757,231,1217]" pageId="9" pageNumber="338">Prosternal process well-developed, plough-shaped, bluntly pointed anteriorly, its ventral margin conspicuously sloping anteriad, posterior ventral angle with small but distinct anteriorly recurved tooth. Mesometasternal process indistinct. Sides of sterna somewhat rugosely punctate, with short and rather sparse adpressed hairs; disc of metasternum glabrous, impressed. Abdominal sternites coarsely punctured, punctures smaller and denser at upper sides, each sternite with a transverse row of well spaced rather long semierect setae, which are conspicuously longer on penultimate sternite; last visible sternite with similar setae along distal border.</paragraph>
<paragraph id="F2445876FF8F5014FF4DFCADFE32A4AF" blockId="9.[151,757,231,1217]" pageId="9" pageNumber="338">Protibia bidentate, apical tooth long, basal tooth welldeveloped, inner spur attached at level of basal tooth. Last joint of fore tarsi thickened apically, indented at mid-length of its concave margin. Inner fore claw broadened, with lobes slightly divergent, lower lobe 2.5 times broader than upper at base, its ventral margin distinctly convex. Outer claw of middle tarsi cleft. Metatarsal joints distinctly thicker than pro- and mesotarsal joints. Hind tibiae fusiform. Long spur of hind tibiae evenly doublecurved, narrow but blunt at apex.</paragraph>
<paragraph id="F2445876FF8F5014FF4DFB6DFE4FA4CF" blockId="9.[151,757,231,1217]" box="[151,388,1193,1217]" pageId="9" pageNumber="338">
Aedeagus:
<figureCitation id="6AC044F3FF8F5014FED0FB6DFE4BA4CF" box="[266,384,1193,1217]" captionStart="Figs 44–60" captionStartId="16.[208,248,1893,1915]" captionTargetBox="[224,1401,251,1864]" captionTargetId="figure-41@16.[208,1418,239,1884]" captionTargetPageId="16" captionText="Figs 44–60: Mimela spp., aedeagi. – 44–46: M. krasava spec. nov., holotype (44 lateral view. 45 parameres, dorsal view. 46 parameres, ventral view). 47–50: M. langbianica spec. nov., holotype (47 lateral view, left side. 48 lateral view of parameres, right side. 49 dorsal view of parameres. 50 ventral view of parameres). 51–53: M. pacholatkoi spec. nov., holotype (51 lateral view. 52 parameres, dorsal view. 53 parameres, ventral view). 54 M. ohausi, Laos: Xieng Khouang prov., lateral view. 55–57: M. signaticollis, Vietnam: Nha Trang (55 lateral view. 56 parameres, dorsal view. 57 parameres, ventral view). 58–60: M. vietnamensis spec. nov., holotype (58 lateral view. 59 parameres, dorsal view. 60 parameres, ventral view). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4754155" httpUri="https://zenodo.org/record/4754155/files/figure.png" pageId="9" pageNumber="338">Figs 44–46</figureCitation>
.
</paragraph>
<paragraph id="F2445876FF8F5014FF4DFB2DFD11A56C" blockId="9.[151,756,1257,1378]" pageId="9" pageNumber="338">
<emphasis id="C08F8464FF8F5014FF4DFB2DFF35A50F" bold="true" box="[151,254,1257,1281]" pageId="9" pageNumber="338">Variation</emphasis>
: Punctation of pronotum somewhat coarser and punctation of scutellum denser and more regular in the second male; black apical marks on elytra indistinct and not covering apical protuberances in the
<typeStatus id="2D40E6D4FF8F5014FDADFA8EFD1DA56C" box="[631,726,1354,1378]" pageId="9" pageNumber="338" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="F2445876FF8F5014FF4DFA4DFEABA5AF" blockId="9.[151,352,1417,1441]" box="[151,352,1417,1441]" pageId="9" pageNumber="338">
<emphasis id="C08F8464FF8F5014FF4DFA4DFF2DA5AF" bold="true" box="[151,230,1417,1441]" pageId="9" pageNumber="338">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8F5014FF4DFA0DFDD3A74F" pageId="9" pageNumber="338" type="diagnosis">
<paragraph id="F2445876FF8F5014FF4DFA0DFDD3A74F" blockId="9.[151,757,1481,1857]" pageId="9" pageNumber="338">
<emphasis id="C08F8464FF8F5014FF4DFA0DFE43A5EF" bold="true" box="[151,392,1481,1505]" pageId="9" pageNumber="338">Differential diagnosis</emphasis>
: Small size, color and punctation pattern of elytra make this new species look extremely similar to
<taxonomicName id="35FB23F5FF8F5014FED4F9CDFD9FA62F" authority="ARROW, 1910" authorityName="ARROW" authorityYear="1910" box="[270,596,1545,1569]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="felicia">
<emphasis id="C08F8464FF8F5014FED4F9CDFE7DA62F" box="[270,438,1545,1569]" italics="true" pageId="9" pageNumber="338">Anomala felicia</emphasis>
<bibRefCitation id="966A2587FF8F5014FE18F9CDFD9FA62F" author="ARROW, G. J." box="[450,596,1545,1569]" pageId="9" pageNumber="338" pagination="64 - 72" refId="ref13693" refString="ARROW, G. J. 1910: On a few new Bornean beetles of the Rutelid genera Mimela and Anomala. - The Annals and Magazine of natural History, including Zoology, Botany and Geology, London (series 8) 6: 64 - 72." type="journal article" year="1910">ARROW, 1910</bibRefCitation>
</taxonomicName>
from Borneo, however, the well developed prosternal process places this species in the genus
<taxonomicName id="35FB23F5FF8F5014FE6FF98DFDCEA66F" authorityName="KIRBY" authorityYear="1825" box="[437,517,1609,1633]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="C08F8464FF8F5014FE6FF98DFDCEA66F" box="[437,517,1609,1633]" italics="true" pageId="9" pageNumber="338">Mimela</emphasis>
</taxonomicName>
. The closest relatives appear to be the small Sundanian species
<taxonomicName id="35FB23F5FF8F5014FDAAF9AEFEAEA6AF" authority="BURMEISTER, 1844" authorityName="BURMEISTER" authorityYear="1844" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="discoidea">
<emphasis id="C08F8464FF8F5014FDAAF9AEFD3FA68F" box="[624,756,1641,1665]" italics="true" pageId="9" pageNumber="338">M. discoidea</emphasis>
<bibRefCitation id="966A2587FF8F5014FF4DF94DFEAEA6AF" author="BURMEISTER, H." box="[151,357,1673,1697]" pageId="9" pageNumber="338" refId="ref13781" refString="BURMEISTER, H. 1844: Handbuch der Entomologie. Vol. 4 (1), Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Berlin: Enslin, ii + 587 pp." type="book" year="1844">BURMEISTER, 1844</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="35FB23F5FF8F5014FEAFF94EFDADA6AF" authority="OHAUS, 1913" authorityName="OHAUS" authorityYear="1913" box="[373,614,1673,1697]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="C08F8464FF8F5014FEAFF94EFE00A6AF" box="[373,459,1673,1697]" italics="true" pageId="9" pageNumber="338">M. vethi</emphasis>
<bibRefCitation id="966A2587FF8F5014FE0CF94DFDADA6AF" author="OHAUS, F." box="[470,614,1673,1697]" pageId="9" pageNumber="338" pagination="29 - 37" refId="ref14279" refString="OHAUS, F. 1913: Neue Indomalayische Ruteliden. - Tijdschrift voor Entomologie 56: 29 - 37." type="journal article" year="1913">OHAUS, 1913</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="35FB23F5FF8F5014FD7EF94DFE59A6CF" authority="LANSBERGE, 1879" authorityName="LANSBERGE" authorityYear="1879" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="C08F8464FF8F5014FD7EF94DFF06A6CF" italics="true" pageId="9" pageNumber="338">Mimela nana</emphasis>
<bibRefCitation id="966A2587FF8F5014FF09F96DFE59A6CF" author="LANSBERGE, G. W. VAN" box="[211,402,1705,1729]" pageId="9" pageNumber="338" refId="ref13987" refString="LANSBERGE, G. W. VAN 1879: Diagnoses de quelques especes nouvelles de Buprestidae et de Scarabaeides de la Malaisie. - Annales de la Societe entomologique de Belgique (Compte Rendus) 22: CXLVII - CLV." type="book" year="1879">LANSBERGE, 1879</bibRefCitation>
</taxonomicName>
. The aedeagus of
<taxonomicName id="35FB23F5FF8F5014FD97F96DFD3FA6CF" authorityName="Prokofiev &amp; Zorn" authorityYear="2016" box="[589,756,1705,1729]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="krasava">
<emphasis id="C08F8464FF8F5014FD97F96DFD3FA6CF" box="[589,756,1705,1729]" italics="true" pageId="9" pageNumber="338">Mimela krasava</emphasis>
</taxonomicName>
is very different from those species in having apically somewhat raised and pointed parameres. Moreover, all known similar
<taxonomicName id="35FB23F5FF8F5014FEE5F8CDFE2FA72F" box="[319,484,1801,1825]" class="Insecta" family="Rutelidae" genus="Mimela" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C08F8464FF8F5014FEE5F8CDFE44A72F" box="[319,399,1801,1825]" italics="true" pageId="9" pageNumber="338">Mimela</emphasis>
species
</taxonomicName>
from
<collectingCountry id="8AEC18E6FF8F5014FDF7F8CDFD46A72F" box="[557,653,1801,1825]" name="Malaysia" pageId="9" pageNumber="338">Malaysia</collectingCountry>
, Sumatra and Java are more or less testaceous.
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8F5014FF4DF8ADFEF4A7AF" pageId="9" pageNumber="338" type="distribution">
<paragraph id="F2445876FF8F5014FF4DF8ADFEF4A7AF" blockId="9.[151,756,1897,1953]" pageId="9" pageNumber="338">
<emphasis id="C08F8464FF8F5014FF4DF8ADFEEBA78F" bold="true" box="[151,288,1897,1921]" pageId="9" pageNumber="338">Distribution</emphasis>
: So far known only from the Dalat Plateau, South
<collectingCountry id="8AEC18E6FF8F5014FF06F84DFEF0A7AF" box="[220,315,1929,1953]" name="Vietnam" pageId="9" pageNumber="338">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BAE10BFDFF8F5014FF4DF80DFEC4A82F" pageId="9" pageNumber="338" type="etymology">
<paragraph id="F2445876FF8F5014FF4DF80DFEC4A82F" blockId="9.[151,756,1993,2081]" pageId="9" pageNumber="338">
<emphasis id="C08F8464FF8F5014FF4DF80DFEC5A7EF" bold="true" box="[151,270,1993,2017]" pageId="9" pageNumber="338">Etymology</emphasis>
: The species epithet is derived from a Russian word meaning “beautiful”, to be used as noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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