treatments-xml/data/4D/02/87/4D0287B1FF92530EFDD71DE72F5A0168.xml

<document id="D181B7AA34548C3D228AC106D04F97D5" ID-CLB-Dataset="30015" ID-DOI="10.5852/ejt.2018.431" ID-GBIF-Dataset="abb4a6c8-d4a2-4153-9372-3196b43899ef" ID-Zenodo-Dep="1254968" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1527596642050" checkinUser="plazi" docAuthor="Boonyanusith, Chaichat, Sanoamuang, La-orsri &amp; Brancelj, Anton" docDate="2018" docId="4D0287B1FF92530EFDD71DE72F5A0168" docLanguage="en" docName="ejt-431_boonyanusith_sanoamuang_brancelj.pdf.imf" docOrigin="European Journal of Taxonomy 431" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Siamcyclops cavernicolus Boonyanusith &amp; Sanoamuang &amp; Brancelj 2018, gen. et sp. nov." docType="treatment" docUuid="33DF0C7F-AFB6-409A-A5B3-45ED0664B27C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="14" masterDocId="B13BFFC9FF915303FFF418732A62057B" masterDocTitle="A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenera of the Bryocyclops and Microcyclops groups" masterLastPageNumber="30" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547541066744" originalUpdateUser="plazi" pageNumber="4" updateTime="1698545469938" updateUser="ExternalLinkService">
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<mods:title id="C828D6B7CB93D2E60E856C36318FDFA9">A new genus and two new species of cave-dwelling cyclopoids (Crustacea, Copepoda) from the epikarst zone of Thailand and up-to-date keys to genera and subgenera of the Bryocyclops and Microcyclops groups</mods:title>
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<mods:namePart id="51D23C8D7E9D4B9AD69F6FA540DACBB1">Boonyanusith, Chaichat</mods:namePart>
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<mods:namePart id="F957AD46790EB6613C1E3C5FC211D3AC">Sanoamuang, La-orsri</mods:namePart>
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<mods:namePart id="42178BBCFE76B289490FF3F8EE0142F3">Brancelj, Anton</mods:namePart>
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<mods:title id="B540999661383019FCAE95DFF2DB60ED">European Journal of Taxonomy</mods:title>
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<mods:date id="CFD0DC940FE7A6C18EEF3138504B4351">2018</mods:date>
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<treatment id="4D0287B1FF92530EFDD71DE72F5A0168" ID-DOI="http://doi.org/10.5281/zenodo.5640768" ID-GBIF-Taxon="144171061" ID-Zenodo-Dep="5640768" LSID="urn:lsid:zoobank.org:act:33DF0C7F-AFB6-409A-A5B3-45ED0664B27C" httpUri="http://treatment.plazi.org/id/4D0287B1FF92530EFDD71DE72F5A0168" lastPageId="13" lastPageNumber="14" pageId="3" pageNumber="4">
<subSubSection id="8DB1652CFF925300FDD71DE72EA900A9" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="C51436A7FF925300FDD71DE72E7300D4" blockId="3.[360,1227,1428,1526]" box="[547,1041,1428,1455]" pageId="3" pageNumber="4">
<heading id="9E5C81CBFF925300FDD71DE72E7300D4" box="[547,1041,1428,1455]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="02AB4D24FF925300FDD71DE7293B00D4" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[547,857,1428,1455]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF925300FDD71DE7293B00D4" bold="true" box="[547,857,1428,1455]" italics="true" pageId="3" pageNumber="4">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF925300FC951DE62E7300D4" box="[865,1041,1429,1455]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="C51436A7FF925300FE9C1DC42EA900A9" blockId="3.[360,1227,1428,1526]" box="[360,1227,1463,1490]" pageId="3" pageNumber="4">urn:lsid:zoobank.org:act:33DF0C7F-AFB6-409A-A5B3-45ED0664B27C</paragraph>
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<subSubSection id="8DB1652CFF925300FD131DA8292F008D" box="[743,845,1499,1526]" pageId="3" pageNumber="4" type="description">
<paragraph id="C51436A7FF925300FD131DA8292F008D" blockId="3.[360,1227,1428,1526]" box="[743,845,1499,1526]" pageId="3" pageNumber="4">
<figureCitation id="5D902A22FF925300FD131DA8292F008D" box="[743,845,1499,1526]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStart-2="Fig. 3" captionStart-3="Fig. 4" captionStart-4="Fig. 5" captionStartId-0="6.[189,232,1929,1955]" captionStartId-1="7.[189,232,1929,1955]" captionStartId-2="9.[189,232,1964,1990]" captionStartId-3="10.[189,232,1965,1991]" captionStartId-4="11.[189,232,1999,2025]" captionTargetBox-0="[191,1387,328,1870]" captionTargetBox-1="[193,1388,317,1881]" captionTargetBox-2="[191,1382,418,1910]" captionTargetBox-3="[191,1393,340,1918]" captionTargetBox-4="[192,1392,1034,1951]" captionTargetId-0="figure@6.[189,1398,323,1886]" captionTargetId-1="figure@7.[189,1398,314,1887]" captionTargetId-2="figure@9.[189,1398,416,1923]" captionTargetId-3="figure@10.[189,1398,338,1921]" captionTargetId-4="figure@11.[189,1398,1032,1957]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." captionText-1="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." captionText-2="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." captionText-3="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." captionText-4="Fig. 5. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. P3. B. P4. Scale bar: 100 μm." httpUri-0="https://zenodo.org/record/1254970/files/figure.png" httpUri-1="https://zenodo.org/record/1254972/files/figure.png" httpUri-2="https://zenodo.org/record/1254974/files/figure.png" httpUri-3="https://zenodo.org/record/1254976/files/figure.png" httpUri-4="https://zenodo.org/record/1254978/files/figure.png" pageId="3" pageNumber="4">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DB1652CFF925300FF491E6E288A03F3" pageId="3" pageNumber="4" type="etymology">
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<heading id="9E5C81CBFF925300FF491E6E2B26034C" bold="true" box="[189,324,1565,1591]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="F7DFEAB5FF925300FF491E6E2B26034C" bold="true" box="[189,324,1565,1591]" pageId="3" pageNumber="4">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF925300FF491E38288A03F3" blockId="3.[189,1399,1565,1673]" pageId="3" pageNumber="4">
The specific epithet 
<taxonomicName id="02AB4D24FF925300FE5F1E382823031E" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[427,577,1611,1637]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cavernicolus">
<emphasis id="F7DFEAB5FF925300FE5F1E382823031E" box="[427,577,1611,1637]" italics="true" pageId="3" pageNumber="4">cavernicolus</emphasis>
</taxonomicName>
meaning ‘organism living in a cave’, indicates its habitat. The epithet is a noun in the nominative singular masculine.
</paragraph>
</subSubSection>
<subSubSection id="8DB1652CFF925300FF491EC32EE70218" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="C51436A7FF925300FF491EC32BC803B1" blockId="3.[189,426,1712,1738]" box="[189,426,1712,1738]" pageId="3" pageNumber="4">
<heading id="9E5C81CBFF925300FF491EC32BC803B1" bold="true" box="[189,426,1712,1738]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="F7DFEAB5FF925300FF491EC32BC803B1" bold="true" box="[189,426,1712,1738]" pageId="3" pageNumber="4">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF925300FF1E1EAC2B3E0382" blockId="3.[189,1399,1759,1891]" box="[234,348,1759,1785]" pageId="3" pageNumber="4">
<emphasis id="F7DFEAB5FF925300FF1E1EAC2B3E0382" bold="true" box="[234,348,1759,1785]" pageId="3" pageNumber="4">Holotype</emphasis>
</paragraph>
<paragraph id="C51436A7FF925300FF491F712EE70218" blockId="3.[189,1399,1759,1891]" pageId="3" pageNumber="4">
<materialsCitation id="75C33CFAFF925300FF491F712EE70218" ID-GBIF-Occurrence="3396402302" collectingDate="2009-11-21" collectionCode="NHM" collectorName="C. Boonyanusith" country="Thailand" elevation="100" httpUri="http://data.nhm.ac.uk/object/433817f0-0a01-4a6b-a6dd-23604fe5c536" latitude="13.625584" location="Chom Phon Cave" longLatPrecision="1" longitude="99.58728" pageId="3" pageNumber="4" specimenCode="NHM 2011.2080" specimenCount="1" specimenCount-female="1" stateProvince="Ratchaburi Province" typeStatus="holotype">
<collectingCountry id="BDBC7637FF925300FF491F712B3B0267" box="[189,345,1794,1820]" name="Thailand" pageId="3" pageNumber="4">THAILAND</collectingCountry>
: 
<specimenCount id="D3ADFD2EFF925300FE991F712BE10267" box="[365,387,1794,1820]" pageId="3" pageNumber="4" type="female">♀</specimenCount>
(adult), 414 µm long, 
<collectingRegion id="076FF845FF925300FD421F7129D00267" box="[694,946,1794,1820]" country="Thailand" name="Ratchaburi" pageId="3" pageNumber="4">Ratchaburi Province</collectingRegion>
, 
<location id="C074607CFF925300FC321F712ECB0267" LSID="urn:lsid:plazi:treatment:4D0287B1FF92530EFDD71DE72F5A0168:C074607CFF925300FC321F712ECB0267" box="[966,1193,1794,1820]" country="Thailand" latitude="13.625584" longLatPrecision="1" longitude="99.58728" name="Chom Phon Cave" pageId="3" pageNumber="4" stateProvince="Ratchaburi Province">Chom Phon Cave</location>
, 
<geoCoordinate id="A09F5060FF925300FB4A1F732F100267" box="[1214,1394,1792,1820]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="13.625584">13°37′32.1″ N</geoCoordinate>
, 
<geoCoordinate id="A09F5060FF925300FF491F502B04023B" box="[189,358,1827,1856]" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="99.58728">99°35′14.2″ E</geoCoordinate>
, 
<elevation id="4E86D194FF925300FE801F562B9D0244" box="[372,511,1829,1856]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="100.0">
<quantity id="02539B42FF925300FE801F562BDD023B" box="[372,447,1829,1856]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
, filtering bottle, 
<date id="B1151067FF925300FD321F5629080244" box="[710,874,1829,1856]" pageId="3" pageNumber="4" value="2009-11-21">
<collectingDate id="A151E98FFF925300FD321F5629080244" box="[710,874,1829,1856]" pageId="3" pageNumber="4" value="2009-11-21">21 Nov. 2009</collectingDate>
</date>
, 
<collectorName id="685E5371FF925300FC821F562E5F0244" box="[886,1085,1829,1855]" pageId="3" pageNumber="4">C. Boonyanusith</collectorName>
leg., completely dissected then mounted on a slide in glycerol and sealed with nail polish (
<specimenCode id="950D9EDCFF925300FC5E1F3A2E1B0218" box="[938,1145,1865,1891]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="4">NHM 2011.2080</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DB1652CFF925307FF1E1FF9287104A2" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="C51436A7FF925300FF1E1FF92B3602DF" blockId="3.[189,1398,1930,2027]" box="[234,340,1930,1956]" pageId="3" pageNumber="4">
<emphasis id="F7DFEAB5FF925300FF1E1FF92B3602DF" bold="true" box="[234,340,1930,1956]" pageId="3" pageNumber="4">Allotype</emphasis>
</paragraph>
<paragraph id="C51436A7FF925300FF491FDD29C60290" blockId="3.[189,1398,1930,2027]" pageId="3" pageNumber="4">
<materialsCitation id="75C33CFAFF925300FF491FDD29C60290" ID-GBIF-Occurrence="1846472192" collectingDate="2009-11-21" collectionCode="NHM" collectorName="C. Boonyanusith" country="Thailand" elevation="100" latitude="13.625584" location="THAILAND" longLatPrecision="1" longitude="99.58728" pageId="3" pageNumber="4" specimenCode="NHM 2011.2081" specimenCount="1" specimenCount-male="1" stateProvince="Ratchaburi Province" typeStatus="holotype">
<collectingCountry id="BDBC7637FF925300FF491FDD2B3B02B3" box="[189,345,1966,1992]" name="Thailand" pageId="3" pageNumber="4">THAILAND</collectingCountry>
: 
<specimenCount id="D3ADFD2EFF925300FE901FDD2B1802B3" box="[356,378,1966,1992]" pageId="3" pageNumber="4" type="male">♂</specimenCount>
(adult), 394 µm long, collected with the 
<typeStatus id="1A108805FF925300FCA91FDE29A502BC" box="[861,967,1965,1991]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, completely dissected, mounted on a slide in glycerol and sealed with nail polish (
<specimenCode id="950D9EDCFF925300FD3D1FA229FA0290" box="[713,920,2001,2027]" collectionCode="NHM" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="4">NHM 2011.2081</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="C51436A7FF955307FF1E19782B05045E" blockId="4.[189,1399,267,473]" box="[234,359,267,293]" pageId="4" pageNumber="5">
<emphasis id="F7DFEAB5FF955307FF1E19782B05045E" bold="true" box="[234,359,267,293]" pageId="4" pageNumber="5">Paratypes</emphasis>
</paragraph>
<paragraph id="C51436A7FF955307FF49195C287104A2" blockId="4.[189,1399,267,473]" pageId="4" pageNumber="5">
<materialsCitation id="75C33CFAFF955307FF49195C29150416" ID-GBIF-Occurrence="3396402301" collectionCode="NHM" country="Thailand" httpUri="http://data.nhm.ac.uk/object/9248624c-fb70-4295-88b5-10425c212a49" location="THAILAND" pageId="4" pageNumber="5" specimenCode="NHM 2011.2082-2084" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<collectingCountry id="BDBC7637FF955307FF49195C2B3B0432" box="[189,345,303,329]" name="Thailand" pageId="4" pageNumber="5">THAILAND</collectingCountry>
: 
<specimenCount id="D3ADFD2EFF955307FE9C195C2BCD0432" box="[360,431,303,329]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
without egg sacs, 
<specimenCount id="D3ADFD2EFF955307FD6E195C28A90432" box="[666,715,303,329]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
with spermatophore, sampled with the 
<typeStatus id="1A108805FF955307FB41195C2F7D0432" box="[1205,1311,303,329]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, whole specimens stored in 70% alcohol (
<specimenCode id="950D9EDCFF955307FDA11920290D0416" box="[597,879,339,365]" pageId="4" pageNumber="5">NHM 2011.2082‒2084</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="75C33CFAFF955307FC701920285304CE" ID-GBIF-Occurrence="1846472193" collectingDate="2009-11-21" collectionCode="KKU" collectorName="C. Boonyanusith" country="Thailand" elevation="100" latitude="13.625584" location="Chom Phon Cave" longLatPrecision="1" longitude="99.58728" pageId="4" pageNumber="5" specimenCode="KKU-COP-2011-002" specimenCount="4" specimenCount-female="4" stateProvince="Ratchaburi Province" typeStatus="paratype">
<specimenCount id="D3ADFD2EFF955307FC70192029D30416" box="[900,945,339,365]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
with spermatophore and egg sac, 
<specimenCount id="D3ADFD2EFF955307FABD19202F140416" box="[1353,1398,339,365]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
with spermatophore, 
<specimenCount id="D3ADFD2EFF955307FE4719042B9604EA" box="[435,500,375,401]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
without egg sacs, sampled with the holotype, whole specimens stored in 70% alcohol (
<specimenCode id="950D9EDCFF955307FED619E8284604CE" box="[290,548,411,437]" pageId="4" pageNumber="5">KKU-COP-2011-002</specimenCode>
);
</materialsCitation>
<materialsCitation id="75C33CFAFF955307FDC319E8287104A2" ID-GBIF-Occurrence="1846472190" collectingDate="2009-11-21" collectionCode="NIB" collectorName="C. Boonyanusith" country="Thailand" elevation="100" latitude="13.625584" location="Chom Phon Cave" longLatPrecision="1" longitude="99.58728" pageId="4" pageNumber="5" specimenCount="4" specimenCount-female="4" stateProvince="Ratchaburi Province" typeStatus="paratype">
<specimenCount id="D3ADFD2EFF955307FDC319E8281B04CE" box="[567,633,411,437]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
, 
<specimenCount id="D3ADFD2EFF955307FD7219E828A504CE" box="[646,711,411,437]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
with egg sac, sampled with the holotype, whole specimens stored in 70% alcohol (NIB).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DB1652CFF955307FF491A72297D0138" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="C51436A7FF955307FF491A722B3F0760" blockId="4.[189,349,513,539]" box="[189,349,513,539]" pageId="4" pageNumber="5">
<heading id="9E5C81CBFF955307FF491A722B3F0760" bold="true" box="[189,349,513,539]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="F7DFEAB5FF955307FF491A722B3F0760" bold="true" box="[189,349,513,539]" pageId="4" pageNumber="5">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF955307FF491A43297D0138" blockId="4.[189,1399,560,1091]" pageId="4" pageNumber="5">
The Chom Phon Cave is located in Chom Bung district, about 
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west of the town of 
<collectingRegion id="076FF845FF955307FB481A432F230731" box="[1212,1345,560,586]" country="Thailand" name="Ratchaburi" pageId="4" pageNumber="5">Ratchaburi</collectingRegion>
. The cave is located in an isolated, small, limestone hill. It is a fossil cave, about 
<quantity id="02539B42FF955307FBC51A262E190714" box="[1073,1147,597,623]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="m" value="290.0">290 m</quantity>
long, with horizontal galleries only and the origin of the sampled water is exclusively drips of percolating water from the epikarst. The cave has two major openings. The first is an entrance located about 
<quantity id="02539B42FF955307FB801AEE2ED207CC" box="[1140,1200,669,695]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</quantity>
above the valley floor at an elevation of 
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a.s.l. The coordinates of the entrance are 
<geoCoordinate id="A09F5060FF955307FBE71ACC2EDC07A0" box="[1043,1214,703,731]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="13.625584">13º37′32.1″ N</geoCoordinate>
, 
<geoCoordinate id="A09F5060FF955307FB3F1ACC2F1007A0" box="[1227,1394,703,731]" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="99.58728">99º35′14.2″ E</geoCoordinate>
. Beyond the entrance is a long horizontal gallery at the same level as the valley floor and accessible without any special equipment except a lamp. The gallery is about 
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wide and 
<quantity id="02539B42FF955307FB5D1B7A2F660658" box="[1193,1284,777,803]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="4" pageNumber="5" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8–10 m</quantity>
high and was formed by a subterranean river. The second opening, about 
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in diameter, is located above the end-hall of the cave. There is a reclining Buddha statue and in a semi-illuminated place there are several plastic and aluminium buckets (= Buddha pots according to 
<bibRefCitation id="A13A4B56FF955307FC601B072EE606F4" box="[916,1156,884,911]" pageId="4" pageNumber="5" refId="ref13250" refString="Brancelj A., Watiroyram S. &amp; Sanoamuang L. 2010. The first record of cave-dwelling Copepoda from Thailand and description of a new species: Elaphoidella namnaoensis n. sp. (Copepoda, Harpacticoida). Crustaceana 83: 779 - 793. https: // doi. org / 10.1163 / 001121610 X 502894" type="journal article">
Brancelj 
<emphasis id="F7DFEAB5FF955307FBF51B062E5C06F4" box="[1025,1086,885,911]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2010
</bibRefCitation>
) collecting dripping water from the cave walls and roots of trees for the whole year around. This is the 
<typeStatus id="1A108805FF955307FB4A1BEA2E9306C8" box="[1214,1265,921,947]" pageId="4" pageNumber="5">type</typeStatus>
locality of 
<taxonomicName id="02AB4D24FF955307FF491BCE2B8F06AC" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[189,493,957,983]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF955307FF491BCE2B8F06AC" box="[189,493,957,983]" italics="true" pageId="4" pageNumber="5">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF955307FE011BCE28C706AC" box="[501,677,957,983]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
On the sampling date (
<date id="B1151067FF955307FC4C1BCE2E3E06AC" box="[952,1116,957,983]" pageId="4" pageNumber="5" value="2009-11-21">21 Nov. 2009</date>
), no water was dripping from the cave walls or tree roots. The volume of water in the plastic buckets was about 5 l and had a brown colour. The water temperature was 23.8°C, pH 8.74, and conductivity 435 µS cm-1. There was some guano and other organic debris in the buckets.
</paragraph>
</subSubSection>
<subSubSection id="8DB1652CFF955308FF491C182E7406CE" lastPageId="11" lastPageNumber="12" pageId="4" pageNumber="5" type="description">
<paragraph id="C51436A7FF955307FF491C182B2C01FE" blockId="4.[189,334,1131,1157]" box="[189,334,1131,1157]" pageId="4" pageNumber="5">
<heading id="9E5C81CBFF955307FF491C182B2C01FE" bold="true" box="[189,334,1131,1157]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="F7DFEAB5FF955307FF491C182B2C01FE" bold="true" box="[189,334,1131,1157]" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF955307FF1E1CE92B2701CF" blockId="4.[189,1399,1178,1781]" box="[234,325,1178,1204]" pageId="4" pageNumber="5">
<emphasis id="F7DFEAB5FF955307FF1E1CE92B2701CF" bold="true" box="[234,325,1178,1204]" pageId="4" pageNumber="5">Female</emphasis>
</paragraph>
<paragraph id="C51436A7FF955307FF491CCD2F53038E" blockId="4.[189,1399,1178,1781]" pageId="4" pageNumber="5">
Body length, measured from tip of rostrum to posterior margin of caudal rami, 391‒414 µm (mean: 404 µm; n = 10); prosome/urosome ratio about 2.0 (
<figureCitation id="5D902A22FF955307FCFF1C9129070187" box="[779,869,1250,1276]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1A</figureCitation>
). Body dorsoventrally compressed. Preserved specimens transparent; naupliar eye not discernible; rostrum small, triangular. Cephalothorax anteriorly oval, as long as wide, with greatest width at posterior end of cephalosome; representing 41% of body length. Posterior margins of Pd1–Pd4 smooth. Integument smooth, not strongly chitinized, with no visible cuticular windows. Body width/length ratio 2.4. Genital double-somite well developed, about 1.4 times as wide as long, with pair of refractile, sclerotized, rounded, dorsolateral lobes (
<figureCitation id="5D902A22FF955307FB5B1DE52F6900CB" box="[1199,1291,1430,1456]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
); as long as following urosomites, including caudal rami; with well-discernible incision between anterior and posterior half of segment and tapering posteriorly (
<figureCitation id="5D902A22FF955307FCD61DAD29C40083" box="[802,934,1502,1528]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1A–C</figureCitation>
). Copulatory pore small, oval, situated near middle of somite; copulatory duct short, narrow and well sclerotized. Seminal receptacle small, representing about ⅓ of double-somite length, with anterior and posterior expansions; both expansions short but anterior one slightly longer, sclerotized and much wider compared to posterior one (
<figureCitation id="5D902A22FF955307FAF81E392F05031F" box="[1292,1383,1610,1636]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1B</figureCitation>
). Ovipores situated lateroposteriorly at about ½ length of somite, covered with reduced P6 (
<figureCitation id="5D902A22FF955307FAF31E1D2F0A03F3" box="[1287,1384,1646,1672]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1D</figureCitation>
). Posterior margin of genital double-somite and two subsequent somites with hyaline fringes with irregular serration both ventrally and dorsally. Anal somite with well developed operculum, reaching ½ length of caudal rami; distal half of free margin serrated; two large sensilla at base of operculum (
<figureCitation id="5D902A22FF955307FB321EA92F40038E" box="[1222,1314,1754,1781]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="C51436A7FF955306FF491F6E2BAC0430" blockId="4.[189,1399,1820,2027]" lastBlockId="5.[189,1398,267,331]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
CAUDAL RAMI (
<figureCitation id="5D902A22FF955307FE841F6E2B97024C" box="[368,501,1821,1847]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="4" pageNumber="5">Fig. 1B–D</figureCitation>
). Slightly divergent; each about 1.5 times as long as wide. Anterolateral accessory seta (I) reduced. Anterolateral seta (II) bare, implanted at ⅔ length of ramus; slightly shorter than ramus. Posterolateral seta (III) slim, about 1.2 times as long as ramus, implanted at ¼ of ramus length; insertion of seta ornamented with few spinules. Outer apical seta (IV) plumose; inner apical seta (V) longest; both setae bipinnatae, without fracture planes; inner one about twice as long as outer one and about 0.5 times as long as body. Apical accessory seta (VI) bare, spiniform, curved outward; about 0.5 times as long as ramus. Dorsal seta (VII) bipinnate, inserted at distal inner corner of ramus, about twice as long as ramus.
</paragraph>
<paragraph id="C51436A7FF945306FF4919072B2B0485" blockId="5.[189,1399,372,510]" pageId="5" pageNumber="6">
ANTENNULE (
<figureCitation id="5D902A22FF945306FEA419072BC504F5" box="[336,423,372,399]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="5" pageNumber="6">Fig. 1E</figureCitation>
). 11-segmented, not reaching posterior margin of cephalothorax. Armature formula: 6.2.5.2.0+I.2.3.2+A.2.2+A.7+A. Fifth segment with short spine ventrally. Penultimate segment with aesthetasc near insertion of outer seta; aesthetasc as long as outer seta. Terminal segment with acrotheck subapically.
</paragraph>
<paragraph id="C51436A7FF945306FF491A542EC90787" blockId="5.[189,1399,551,764]" pageId="5" pageNumber="6">
ANTENNA (
<figureCitation id="5D902A22FF945306FECD1A542BFA073A" box="[313,408,551,578]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
). 4-segmented; with coxobasis and 3-segmented Endp. Coxobasis with one smooth seta on distal inner corner; seta representing Exp absent. Endp-1 with longitudinal row of spinules along distal half of inner margin; with one smooth seta at ½ length of margin. Endp-2 about 1.5 times as long as wide, with longitudinal row of minute spinules; along inner margin five smooth setae increasing in length (three laterally, one subapically, one apically). Endp-3 twice as long as wide, with longitudinal row of minute spinules; seven smooth setae apically (two of them robust and curved).
</paragraph>
<paragraph id="C51436A7FF945306FF491B562BF206D4" blockId="5.[189,1398,804,943]" pageId="5" pageNumber="6">
MANDIBLE (
<figureCitation id="5D902A22FF945306FEB31B572BA90644" box="[327,459,804,831]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="5" pageNumber="6">Fig. 2B–C</figureCitation>
). With coxa and short basis partly fused with coxa. Gnathobase with strong chitinized teeth; ventralmost teeth very robust and slightly obtuse, with pinnate seta dorsally. Basis with one short and two long setae representing Exp and Endp, respectively; long setae about seven times as long as short seta.
</paragraph>
<paragraph id="C51436A7FF945306FF491BAB28B701FC" blockId="5.[189,1399,984,1159]" pageId="5" pageNumber="6">
MAXILLULE (
<figureCitation id="5D902A22FF945306FEB91BAB2BCB0689" box="[333,425,984,1010]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="5" pageNumber="6">Fig. 2D</figureCitation>
). With robust praecoxa and 2-segmented palp; proximal segment of palp coxobasis; distal one Endp. Arthrite of praecoxa with six strong spines laterally, five of them smooth; apically three claw-like spines decreasing in length and one weak seta. Coxobasis with three elements; one robust bipinnate seta accompanied by two weak smooth setae apically. Endp with two setae apically and one seta subapically. Exp represented by one seta.
</paragraph>
<paragraph id="C51436A7FF945306FF491CC329B600FF" blockId="5.[189,1399,1200,1412]" pageId="5" pageNumber="6">
MAXILLA (
<figureCitation id="5D902A22FF945306FEC61CC32BE901B1" box="[306,395,1200,1226]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="5" pageNumber="6">Fig. 2E</figureCitation>
). 5-segmented. Endite of praecoxa prominent, with two plumose setae. Proximal endite of coxa with one plumose seta; distal endite elongate, with two bipinnate setae apically. Basis with clawlike basal endite, with row of spinules along concave margin; two setae at base of claw-like expansion; longest one as long as claw; other one shorter, 0.5 times as long as longer one. Endp 2-segmented; proximal segment with two robust setae; distal segment with one robust seta apically, as long as clawlike expansion on basis, accompanied by two slender, shorter setae.
</paragraph>
<paragraph id="C51436A7FF945306FF491DDD2E5E0369" blockId="5.[189,1399,1453,1554]" pageId="5" pageNumber="6">
MAXILLIPED (
<figureCitation id="5D902A22FF945306FEA01DDE2BCF00B3" box="[340,429,1453,1480]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="5" pageNumber="6">Fig. 2F</figureCitation>
). 4-segmented; syncoxa and basis with two pinnate setae each; basis with one row of spinules on distal outer margin; additional row of spinules laterally. Proximal segment of End with one strong seta. Distal segment armed with three setae, shortest one smooth.
</paragraph>
<paragraph id="C51436A7FF945306FF491E4F285E0390" blockId="5.[189,1399,1595,1771]" pageId="5" pageNumber="6">
P1–P4. With un-ornamented intercoxal sclerite, with deeply concave posterior margins; coxa rectangular; basis relatively triangular with slender outer seta; 2-segmented Exp and Endp (
<figureCitation id="5D902A22FF945306FBA41E132EB10300" box="[1104,1235,1632,1659]" captionStart="Fig. 3" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[191,1382,418,1910]" captionTargetId="figure@9.[189,1398,416,1923]" captionTargetPageId="9" captionText="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="5" pageNumber="6">Fig. 3A–D</figureCitation>
). Coxa of P1– P3 each armed with one plumose seta on inner distal corner. Exp-2 spine/seta formula: 3.3.3.2/5.5.5.5. Endp-2 spine/seta formula: 1.1.1.1/4.4.5.4. Complete armature of P1–P4 as follows (Roman numbers = spines; Arabic numbers = setae):
</paragraph>
<paragraph id="C51436A7FF945306FC881F6B2F3A0298" pageId="5" pageNumber="6">
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<tr id="7B9B34E5FF94ACFCFF201F6B2F3A0249" box="[212,1368,1816,1842]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<th id="384A5D99FF94ACFCFCB01F6B2E400249" box="[836,1058,1816,1842]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FC881F6B29BE0249" bold="true" box="[892,988,1816,1842]" pageId="5" pageNumber="6">Exopod</emphasis>
</th>
<th id="384A5D99FF94ACFCFB781F6B2F3A0249" box="[1164,1368,1816,1842]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FB411F6B2F450249" bold="true" box="[1205,1319,1816,1842]" pageId="5" pageNumber="6">Endopod</emphasis>
</th>
</tr>
<tr id="7B9B34E5FF94ACFCFF201F492F3A022E" box="[212,1368,1850,1877]" gridrow="1" pageId="5" pageNumber="6">
<th id="384A5D99FF94ACFCFF201F492BE7022E" box="[212,389,1850,1877]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FF201F492BE7022E" bold="true" box="[212,389,1850,1877]" pageId="5" pageNumber="6">Swimming leg</emphasis>
</th>
<td id="384A5D99FF94ACFCFE171F492846022E" box="[483,548,1850,1877]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FE171F492846022F" bold="true" box="[483,548,1850,1876]" pageId="5" pageNumber="6">Coxa</emphasis>
</td>
<td id="384A5D99FF94ACFCFD791F4928AC022E" box="[653,718,1850,1877]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FD791F4928AC022F" bold="true" box="[653,718,1850,1876]" pageId="5" pageNumber="6">Basis</emphasis>
</td>
<td id="384A5D99FF94ACFCFCB01F492909022E" box="[836,875,1850,1877]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FCA41F49293D022F" bold="true" box="[848,863,1850,1876]" pageId="5" pageNumber="6">1</emphasis>
</td>
<td id="384A5D99FF94ACFCFC4F1F492E40022E" box="[955,1058,1850,1877]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FC131F492994022F" bold="true" box="[999,1014,1850,1876]" pageId="5" pageNumber="6">2</emphasis>
</td>
<td id="384A5D99FF94ACFCFB781F492EDA022E" box="[1164,1208,1850,1877]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FB6F1F492EC8022F" bold="true" box="[1179,1194,1850,1876]" pageId="5" pageNumber="6">1</emphasis>
</td>
<td id="384A5D99FF94ACFCFB021F492F3A022E" box="[1270,1368,1850,1877]" gridcol="6" gridrow="1" pageId="5" pageNumber="6">
<emphasis id="F7DFEAB5FF945306FAEB1F492F4C022F" bold="true" box="[1311,1326,1850,1876]" pageId="5" pageNumber="6">2</emphasis>
</td>
</tr>
<tr id="7B9B34E5FF94ACFCFF201F2D2F3A0202" box="[212,1368,1886,1913]" gridrow="2" pageId="5" pageNumber="6">
<th id="384A5D99FF94ACFCFF201F2D2BE70202" box="[212,389,1886,1913]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">P1</th>
<td id="384A5D99FF94ACFCFE171F2D28460202" box="[483,548,1886,1913]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFD791F2D28AC0202" box="[653,718,1886,1913]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">1–I</td>
<td id="384A5D99FF94ACFCFCB01F2D29090202" box="[836,875,1886,1913]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">I–0</td>
<td id="384A5D99FF94ACFCFC4F1F2D2E400202" box="[955,1058,1886,1913]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">II–I, 1–4</td>
<td id="384A5D99FF94ACFCFB781F2D2EDA0202" box="[1164,1208,1886,1913]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFB021F2D2F3A0202" box="[1270,1368,1886,1913]" gridcol="6" gridrow="2" pageId="5" pageNumber="6">1–I–3</td>
</tr>
<tr id="7B9B34E5FF94ACFCFF201FF12F3A02E7" box="[212,1368,1922,1948]" gridrow="3" pageId="5" pageNumber="6">
<th id="384A5D99FF94ACFCFF201FF12BE702E7" box="[212,389,1922,1948]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">P2</th>
<td id="384A5D99FF94ACFCFE171FF1284602E7" box="[483,548,1922,1948]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFD791FF128AC02E7" box="[653,718,1922,1948]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">1–0</td>
<td id="384A5D99FF94ACFCFCB01FF1290902E7" box="[836,875,1922,1948]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">I–0</td>
<td id="384A5D99FF94ACFCFC4F1FF12E4002E7" box="[955,1058,1922,1948]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">II–I, 1–4</td>
<td id="384A5D99FF94ACFCFB781FF12EDA02E7" box="[1164,1208,1922,1948]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFB021FF12F3A02E7" box="[1270,1368,1922,1948]" gridcol="6" gridrow="3" pageId="5" pageNumber="6">1–I, 1–2</td>
</tr>
<tr id="7B9B34E5FF94ACFCFF201FD62F3A02BB" box="[212,1368,1957,1984]" gridrow="4" pageId="5" pageNumber="6">
<th id="384A5D99FF94ACFCFF201FD62BE702BB" box="[212,389,1957,1984]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">P3</th>
<td id="384A5D99FF94ACFCFE171FD6284602BB" box="[483,548,1957,1984]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFD791FD628AC02BB" box="[653,718,1957,1984]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">1–0</td>
<td id="384A5D99FF94ACFCFCB01FD6290902BB" box="[836,875,1957,1984]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">I–0</td>
<td id="384A5D99FF94ACFCFC4F1FD62E4002BB" box="[955,1058,1957,1984]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">II–I, 1–4</td>
<td id="384A5D99FF94ACFCFB781FD62EDA02BB" box="[1164,1208,1957,1984]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFB021FD62F3A02BB" box="[1270,1368,1957,1984]" gridcol="6" gridrow="4" pageId="5" pageNumber="6">1–I, 1–3</td>
</tr>
<tr id="7B9B34E5FF94ACFCFF201FBA2F3A0298" box="[212,1368,1993,2019]" gridrow="5" pageId="5" pageNumber="6">
<th id="384A5D99FF94ACFCFF201FBA2BE70298" box="[212,389,1993,2019]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">P4</th>
<td id="384A5D99FF94ACFCFE171FBA28460298" box="[483,548,1993,2019]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">0–0</td>
<td id="384A5D99FF94ACFCFD791FBA28AC0298" box="[653,718,1993,2019]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">1–0</td>
<td id="384A5D99FF94ACFCFCB01FBA29090298" box="[836,875,1993,2019]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">I–0</td>
<td id="384A5D99FF94ACFCFC4F1FBA2E400298" box="[955,1058,1993,2019]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">I–I, 1–4</td>
<td id="384A5D99FF94ACFCFB781FBA2EDA0298" box="[1164,1208,1993,2019]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">0–1</td>
<td id="384A5D99FF94ACFCFB021FBA2F3A0298" box="[1270,1368,1993,2019]" gridcol="6" gridrow="5" pageId="5" pageNumber="6">1–I, 1–2</td>
</tr>
</table>
</paragraph>
<caption id="91D4662FFF975305FF491FFA2E470291" httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1929,1955]" targetBox="[191,1387,328,1870]" targetPageId="6">
<paragraph id="C51436A7FF975305FF491FFA2E470291" blockId="6.[189,1399,1929,2026]" pageId="6" pageNumber="7">
<emphasis id="F7DFEAB5FF975305FF491FFA2B6F02D8" bold="true" box="[189,269,1929,1955]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
<taxonomicName id="02AB4D24FF975305FEE21FFA282B02D8" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[278,585,1929,1955]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF975305FEE21FFA282B02D8" box="[278,585,1929,1955]" italics="true" pageId="6" pageNumber="7">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF975305FDA61FF9296E02DF" box="[594,780,1930,1956]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♀. 
<emphasis id="F7DFEAB5FF975305FC4C1FFA29AF02D8" bold="true" box="[952,973,1929,1955]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, with spermatophores and egg sac, dorsal view. 
<emphasis id="F7DFEAB5FF975305FE351FDF2BB702BD" bold="true" box="[449,469,1964,1990]" pageId="6" pageNumber="7">B</emphasis>
. Urosome, ventral view. 
<emphasis id="F7DFEAB5FF975305FCF21FDF297902BD" bold="true" box="[774,795,1964,1990]" pageId="6" pageNumber="7">C</emphasis>
. Urosome, dorsal view. 
<emphasis id="F7DFEAB5FF975305FBB61FDF2E3502BD" bold="true" box="[1090,1111,1964,1990]" pageId="6" pageNumber="7">D</emphasis>
. Urosome, lateral view. 
<emphasis id="F7DFEAB5FF975305FF491FA32AB30291" bold="true" box="[189,209,2000,2026]" pageId="6" pageNumber="7">E</emphasis>
. Antennule. 
<emphasis id="F7DFEAB5FF975305FE921FA32B1A0291" bold="true" box="[358,376,2000,2026]" pageId="6" pageNumber="7">F</emphasis>
. P5, ventral view. 
<emphasis id="F7DFEAB5FF975305FDA71FBC28080292" bold="true" box="[595,618,1999,2025]" pageId="6" pageNumber="7">G</emphasis>
. P5, lateral view. Scale bars: 100 μm.
</paragraph>
</caption>
<caption id="91D4662FFF965304FF491FFA29860291" httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1929,1955]" targetBox="[193,1388,317,1881]" targetPageId="7">
<paragraph id="C51436A7FF965304FF491FFA29860291" blockId="7.[189,1399,1929,2026]" pageId="7" pageNumber="8">
<emphasis id="F7DFEAB5FF965304FF491FFA2B6A02D8" bold="true" box="[189,264,1929,1955]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
<taxonomicName id="02AB4D24FF965304FEF91FFA285902D8" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[269,571,1929,1955]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF965304FEF91FFA285902D8" box="[269,571,1929,1955]" italics="true" pageId="7" pageNumber="8">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF965304FDB41FF9288E02DF" box="[576,748,1930,1956]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♀. 
<emphasis id="F7DFEAB5FF965304FC7F1FFA29C202D8" bold="true" box="[907,928,1929,1955]" pageId="7" pageNumber="8">A</emphasis>
. Antenna. 
<emphasis id="F7DFEAB5FF965304FBEF1FFA2E3002D8" bold="true" box="[1051,1106,1929,1955]" pageId="7" pageNumber="8">B–C</emphasis>
. Mandible. 
<emphasis id="F7DFEAB5FF965304FB2F1FF92E9202DF" bold="true" box="[1243,1264,1930,1956]" pageId="7" pageNumber="8">D</emphasis>
. Maxillule. 
<emphasis id="F7DFEAB5FF965304FF491FDE2AB302BC" bold="true" box="[189,209,1965,1991]" pageId="7" pageNumber="8">E</emphasis>
. Maxilla. 
<emphasis id="F7DFEAB5FF965304FEA61FDE2B0602BC" bold="true" box="[338,356,1965,1991]" pageId="7" pageNumber="8">F</emphasis>
. Maxilliped. 
<emphasis id="F7DFEAB5FF965304FDF81FDF284102BD" bold="true" box="[524,547,1964,1990]" pageId="7" pageNumber="8">G</emphasis>
. Genital double-somite with spermatophore, ventral view. 
<emphasis id="F7DFEAB5FF965304FB021FDE2F6F02BC" bold="true" box="[1270,1293,1965,1991]" pageId="7" pageNumber="8">H</emphasis>
. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="C51436A7FF99530BFF49197F2E4C04E2" blockId="8.[189,1398,267,409]" pageId="8" pageNumber="9">
P1 (
<figureCitation id="5D902A22FF99530BFF1919782B28045E" box="[237,330,267,294]" captionStart="Fig. 3" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[191,1382,418,1910]" captionTargetId="figure@9.[189,1398,416,1923]" captionTargetPageId="9" captionText="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
). Exp-1 with one smooth, blunt, curved spine on outer corner. Exp-2 about twice as long as wide, apical setae as long as Exp-1 and -2 combined. Endp-1 shorter than wide, with sharply pointed extension on distal outer corner. Endp-2 1.3 times as long as wide, with one seta between 2 claw-like extensions on outer margin; very strong, blunt, hook-shaped spine apically.
</paragraph>
<paragraph id="C51436A7FF99530BFF4919B72940077F" blockId="8.[189,1398,451,517]" pageId="8" pageNumber="9">
P2 (
<figureCitation id="5D902A22FF99530BFF1919B02B2A04A5" box="[237,328,451,478]" captionStart="Fig. 3" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[191,1382,418,1910]" captionTargetId="figure@9.[189,1398,416,1923]" captionTargetPageId="9" captionText="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
). Similar to P1, slightly less robust; apical spine on Endp-2 only slightly curved, as long as segment bearing it. Exp-2 1.5 times as long as wide.
</paragraph>
<paragraph id="C51436A7FF99530BFF491A5C29240714" blockId="8.[189,1399,558,624]" pageId="8" pageNumber="9">
P3 (
<figureCitation id="5D902A22FF99530BFF1B1A5C2B2E0732" box="[239,332,559,585]" captionStart="Fig. 3" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[191,1382,418,1910]" captionTargetId="figure@9.[189,1398,416,1923]" captionTargetPageId="9" captionText="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="8" pageNumber="9">Fig. 3C</figureCitation>
). Coxa, basis and Exp similar to those of P2. Endp-2 1.5 times as long as wide; terminal spine on Endp-2 straight, as long as segment bearing it.
</paragraph>
<paragraph id="C51436A7FF99530BFF491AE928B1065C" blockId="8.[189,1398,666,808]" pageId="8" pageNumber="9">
P4 (
<figureCitation id="5D902A22FF99530BFF1A1AE92B2907CF" box="[238,331,666,692]" captionStart="Fig. 3" captionStartId="9.[189,232,1964,1990]" captionTargetBox="[191,1382,418,1910]" captionTargetId="figure@9.[189,1398,416,1923]" captionTargetPageId="9" captionText="Fig. 3. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. P1. B. P2. C. P3. D. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="8" pageNumber="9">Fig. 3D</figureCitation>
). Exp-2 twice as long as wide, with relatively weak pinnate spines on outer margin; apical setae longer than Exp-1 and 2 combined. Endp-1 large, 1.5 times as wide as long. Endp-2 1.3 times as long as wide, with short apical setae, only slightly longer than segment bearing them; apical spine short, about 0.5 times as long as segment bearing it.
</paragraph>
<paragraph id="C51436A7FF99530BFF491B21283006C2" blockId="8.[189,1399,850,953]" pageId="8" pageNumber="9">
P5 (
<figureCitation id="5D902A22FF99530BFF191B212BD10617" box="[237,435,850,876]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="8" pageNumber="9">Figs 1B–D, F–G</figureCitation>
, 
<figureCitation id="5D902A22FF99530BFE4A1B2128690617" box="[446,523,850,876]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="8" pageNumber="9">2G–H</figureCitation>
). Reduced to 2 cuticular lobes, completely fused to Pd5, with three slender setae. Dorsal lobe broad, with one seta; ventral lobe small, with two unequal setae apically; longer seta about twice as long as shorter one.
</paragraph>
<paragraph id="C51436A7FF99530BFF491B9729E6015F" blockId="8.[189,1398,995,1061]" pageId="8" pageNumber="9">
P6 (
<figureCitation id="5D902A22FF99530BFF1B1B972B2C0685" box="[239,334,996,1022]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
). Small, forming simple cuticular plate, inserted laterodorsally on genital double-somite, with two minute spines ventrally and one short seta dorsally.
</paragraph>
<paragraph id="C51436A7FF99530BFF491C3C286601CD" blockId="8.[189,1399,1103,1206]" pageId="8" pageNumber="9">
SPERMATOPHORE (
<figureCitation id="5D902A22FF99530BFE741C3C28600112" box="[384,514,1103,1129]" captionStart="Fig. 2" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[193,1388,317,1881]" captionTargetId="figure@7.[189,1398,314,1887]" captionTargetPageId="7" captionText="Fig. 2. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Antenna. B–C. Mandible. D. Maxillule. E. Maxilla. F. Maxilliped. G. Genital double-somite with spermatophore, ventral view. H. Genital double-somite with spermatophore, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254972/files/figure.png" pageId="8" pageNumber="9">Fig. 2G–H</figureCitation>
). Paired; each forming 3-dimensional structure running in anterior-posterior direction ventrally, bending laterally at level of seminal receptacle and bending dorsally (as double flipped L); walls very thick.
</paragraph>
<paragraph id="C51436A7FF99530BFF491C922E0D0180" blockId="8.[189,1135,1248,1275]" box="[189,1135,1248,1275]" pageId="8" pageNumber="9">
EGG SAC (
<figureCitation id="5D902A22FF99530BFED91C932BEB0180" box="[301,393,1248,1275]" captionStart="Fig. 1" captionStartId="6.[189,232,1929,1955]" captionTargetBox="[191,1387,328,1870]" captionTargetId="figure@6.[189,1398,323,1886]" captionTargetPageId="6" captionText="Fig. 1. Siamcyclops cavernicolus gen. et sp. nov., holotype, ♀. A. Habitus, with spermatophores and egg sac, dorsal view. B. Urosome, ventral view. C. Urosome, dorsal view. D. Urosome, lateral view. E. Antennule. F. P5, ventral view. G. P5, lateral view. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254970/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
). Extruded from gonopores laterodorsally, with two large eggs.
</paragraph>
<paragraph id="C51436A7FF99530BFF1E1D562B480044" blockId="8.[189,1399,1317,1728]" box="[234,298,1317,1343]" pageId="8" pageNumber="9">
<emphasis id="F7DFEAB5FF99530BFF1E1D562B480044" bold="true" box="[234,298,1317,1343]" pageId="8" pageNumber="9">Male</emphasis>
</paragraph>
<paragraph id="C51436A7FF99530BFF491D3F295503BB" blockId="8.[189,1399,1317,1728]" pageId="8" pageNumber="9">
Body length, measured from tip of rostrum to posterior margin of caudal rami, 381‒402 µm (mean: 391 µm; n = 10); prosome/urosome ratio about 2.0. Habitus (
<figureCitation id="5D902A22FF99530BFC721D01298300F7" box="[902,993,1394,1420]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="8" pageNumber="9">Fig. 4A</figureCitation>
) slightly smaller and more slender than in female. Naupliar eye not discernible; rostrum as in female. Cephalothorax and Pd 2–4 similar to those of female. Cephalothorax anteriorly oval, 1.1 times as long as wide, with greatest width at posterior end, representing 43% of body length. Posterior margins of Pd1‒Pd4 smooth. Body length/width ratio about 2.3. Genital somite large, globular, about 0.9 times as long as rest of urosome, including caudal rami (
<figureCitation id="5D902A22FF99530BFEF61E412BE70336" box="[258,389,1586,1613]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="8" pageNumber="9">Fig. 4A–D</figureCitation>
). Hyaline structures on dorsolateral part of genital somite well developed (
<figureCitation id="5D902A22FF99530BFB1C1E412F080336" box="[1256,1386,1586,1613]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="8" pageNumber="9">Fig. 4A–D</figureCitation>
). Posterior border of genital somite with broad hyaline fringe with irregular serration dorsally. Subsequent three urosomites narrower than genital somite, with irregular serrated free hyaline fringes posteriorly. Anal somite and operculum as in female (
<figureCitation id="5D902A22FF99530BFD531ED5294903BB" box="[679,811,1702,1728]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="8" pageNumber="9">Fig. 4A–D</figureCitation>
).
</paragraph>
<paragraph id="C51436A7FF99530BFF491E992EF20290" blockId="8.[189,1399,1770,2027]" pageId="8" pageNumber="9">
CAUDAL RAMI (
<figureCitation id="5D902A22FF99530BFE9C1E992B8A027F" box="[360,488,1770,1796]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="8" pageNumber="9">Fig. 4B–D</figureCitation>
). More slender than in female, about 1.6 times as long as wide. Anterolateral accessory seta (I) reduced.Anterolateral seta (II) bare, implanted at about ½ length of ramus, slightly shorter than ramus. Posterolateral seta (III) bipinnate, about 1.2 times as long as ramus; insertion ornamented with few spinules. Outer apical seta (IV) plumose; without fracture plane. Inner apical seta (V) longest, plumose, about twice as long as seta IV and about 0.5 times as long as body length, without fracture plane. Apical accessory seta (VI) bare, spiniform, curved outward; about 0.5 times as long as ramus. Dorsal seta (VII) bipinnate, inserted at distal inner corner of ramus, about twice as long as ramus.
</paragraph>
<caption id="91D4662FFF98530AFF491FDF2B7C0292" httpUri="https://zenodo.org/record/1254974/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1964,1990]" targetBox="[191,1382,418,1910]" targetPageId="9">
<paragraph id="C51436A7FF98530AFF491FDF2B7C0292" blockId="9.[189,1399,1964,2025]" pageId="9" pageNumber="10">
<emphasis id="F7DFEAB5FF98530AFF491FDF2B6E02BD" bold="true" box="[189,268,1964,1990]" pageId="9" pageNumber="10">Fig. 3.</emphasis>
<taxonomicName id="02AB4D24FF98530AFEE11FDF282402BD" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[277,582,1964,1990]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF98530AFEE11FDF282402BD" box="[277,582,1964,1990]" italics="true" pageId="9" pageNumber="10">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF98530AFDBB1FDF296402BD" box="[591,774,1964,1990]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♀. 
<emphasis id="F7DFEAB5FF98530AFC441FDF29A702BD" bold="true" box="[944,965,1964,1990]" pageId="9" pageNumber="10">A</emphasis>
. P1. 
<emphasis id="F7DFEAB5FF98530AFBF11FDF2E7B02BD" bold="true" box="[1029,1049,1964,1990]" pageId="9" pageNumber="10">B</emphasis>
. P2. 
<emphasis id="F7DFEAB5FF98530AFBAC1FDF2E0F02BD" bold="true" box="[1112,1133,1964,1990]" pageId="9" pageNumber="10">C</emphasis>
. P3. 
<emphasis id="F7DFEAB5FF98530AFB581FDF2EA302BD" bold="true" box="[1196,1217,1964,1990]" pageId="9" pageNumber="10">D</emphasis>
. P4. Scale bar: 100 μm.
</paragraph>
</caption>
<caption id="91D4662FFF9B5309FF491FDE2F370290" httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1965,1991]" targetBox="[191,1393,340,1918]" targetPageId="10">
<paragraph id="C51436A7FF9B5309FF491FDE2F370290" blockId="10.[189,1398,1965,2027]" pageId="10" pageNumber="11">
<emphasis id="F7DFEAB5FF9B5309FF491FDE2B6F02BC" bold="true" box="[189,269,1965,1991]" pageId="10" pageNumber="11">Fig. 4.</emphasis>
<taxonomicName id="02AB4D24FF9B5309FEE31FDE282B02BC" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[279,585,1965,1991]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9B5309FEE31FDE282B02BC" box="[279,585,1965,1991]" italics="true" pageId="10" pageNumber="11">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9B5309FDA71FDD296F02B3" box="[595,781,1966,1992]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, allotype, ♂. 
<emphasis id="F7DFEAB5FF9B5309FC461FDE29A502BC" bold="true" box="[946,967,1965,1991]" pageId="10" pageNumber="11">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="F7DFEAB5FF9B5309FB2B1FDE2E9102BC" bold="true" box="[1247,1267,1965,1991]" pageId="10" pageNumber="11">B</emphasis>
. Urosome, ventral view. 
<emphasis id="F7DFEAB5FF9B5309FEA81FA32B130291" bold="true" box="[348,369,2000,2026]" pageId="10" pageNumber="11">C</emphasis>
. Urosome, lateral view. 
<emphasis id="F7DFEAB5FF9B5309FD641FA328C70291" bold="true" box="[656,677,2000,2026]" pageId="10" pageNumber="11">D</emphasis>
. Urosome, dorsal view. 
<emphasis id="F7DFEAB5FF9B5309FC371FA329B50291" bold="true" box="[963,983,2000,2026]" pageId="10" pageNumber="11">E</emphasis>
. Antennule. Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="C51436A7FF9A5308FF49197F2F150417" blockId="11.[189,1399,267,364]" pageId="11" pageNumber="12">
ANTENNULE (
<figureCitation id="5D902A22FF9A5308FEA619782BCC045E" box="[338,430,267,294]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="11" pageNumber="12">Fig. 4E</figureCitation>
). 15-segmented, geniculate. Armature formula: 7+3A.4.2.2+A.1.2+A.3.1+A+I.2.2.2.2+A.1.1.7+A. Terminal segment with acrotheck; short spine on eighth segment. Seta on ninth and eleventh segments robust, spiniform; seta on tenth, eleventh and twelfth segments very short, bipinnate.
</paragraph>
<paragraph id="C51436A7FF9A5308FF4919E0297C04D6" blockId="11.[189,798,403,430]" box="[189,798,403,430]" pageId="11" pageNumber="12">ANTENNA, MOUTHPARTS, P1, P2 AND P5. As in female.</paragraph>
<paragraph id="C51436A7FF9A5308FF4919A628500723" blockId="11.[189,1399,468,601]" pageId="11" pageNumber="12">
P3 (
<figureCitation id="5D902A22FF9A5308FF1A19A62B2E0494" box="[238,332,468,495]" captionStart="Fig. 5" captionStartId="11.[189,232,1999,2025]" captionTargetBox="[192,1392,1034,1951]" captionTargetId="figure@11.[189,1398,1032,1957]" captionTargetPageId="11" captionText="Fig. 5. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. P3. B. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254978/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
). Exp as in female. Endp-2 with apical spine modified as spoon-like element; bent inward and ornamented with minute transverse denticles along distal ⅓ of margin; tip of spine bent. Inner subterminal seta modified as claw-like spine, bare and bent toward apical spoon-like spine. Sub-terminal seta on outer margin very short.
</paragraph>
<paragraph id="C51436A7FF9A5308FF491AF32BCC07C6" blockId="11.[189,1398,639,701]" pageId="11" pageNumber="12">
P4 (
<figureCitation id="5D902A22FF9A5308FF041AF32B2F07E1" box="[240,333,639,666]" captionStart="Fig. 5" captionStartId="11.[189,232,1999,2025]" captionTargetBox="[192,1392,1034,1951]" captionTargetId="figure@11.[189,1398,1032,1957]" captionTargetPageId="11" captionText="Fig. 5. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. P3. B. P4. Scale bar: 100 μm." httpUri="https://zenodo.org/record/1254978/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
). Exp and Endp similar to those of female; Endp with relatively longer setae and spines compared to female.
</paragraph>
<paragraph id="C51436A7FF9A5308FF491A96286D065A" blockId="11.[189,1398,740,802]" pageId="11" pageNumber="12">
P6 (
<figureCitation id="5D902A22FF9A5308FF051A972B160785" box="[241,372,740,767]" captionStart="Fig. 4" captionStartId="10.[189,232,1965,1991]" captionTargetBox="[191,1393,340,1918]" captionTargetId="figure@10.[189,1398,338,1921]" captionTargetPageId="10" captionText="Fig. 4. Siamcyclops cavernicolus gen. et sp. nov., allotype, ♂. A. Habitus, dorsal view. B. Urosome, ventral view. C. Urosome, lateral view. D. Urosome, dorsal view. E. Antennule. Scale bars: 100 μm." httpUri="https://zenodo.org/record/1254976/files/figure.png" pageId="11" pageNumber="12">Fig. 4B‒C</figureCitation>
). Positioned ventrally; modified to large cuticular plate with three setae; middle one shortest; ventral one longest.
</paragraph>
<paragraph id="C51436A7FF9A5308FF491B3A2B210618" blockId="11.[189,1398,841,949]" box="[189,323,841,867]" pageId="11" pageNumber="12">
<heading id="9E5C81CBFF9A5308FF491B3A2B210618" bold="true" box="[189,323,841,867]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="F7DFEAB5FF9A5308FF491B3A2B210618" bold="true" box="[189,323,841,867]" pageId="11" pageNumber="12">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF9A5308FF491B0B2E7406CE" blockId="11.[189,1398,841,949]" pageId="11" pageNumber="12">No significant variability was observed in females except minor variation in size, shape and number of spinules on some segments or somites. Variability not observed in males.</paragraph>
</subSubSection>
<caption id="91D4662FFF9A5308FF491FBC2F7A0291" box="[189,1304,1999,2026]" httpUri="https://zenodo.org/record/1254978/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1999,2025]" targetBox="[192,1392,1034,1951]" targetPageId="11">
<paragraph id="C51436A7FF9A5308FF491FBC2F7A0291" blockId="11.[189,1304,1999,2026]" box="[189,1304,1999,2026]" pageId="11" pageNumber="12">
<emphasis id="F7DFEAB5FF9A5308FF491FBC2B680292" bold="true" box="[189,266,1999,2025]" pageId="11" pageNumber="12">Fig. 5.</emphasis>
<taxonomicName id="02AB4D24FF9A5308FEE61FBC28200292" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[274,578,1999,2025]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9A5308FEE61FBC28200292" box="[274,578,1999,2025]" italics="true" pageId="11" pageNumber="12">Siamcyclops cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9A5308FDBD1FA328990291" box="[585,763,2000,2026]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, allotype, ♂. 
<emphasis id="F7DFEAB5FF9A5308FC6D1FBC29CC0292" bold="true" box="[921,942,1999,2025]" pageId="11" pageNumber="12">A</emphasis>
. P3. 
<emphasis id="F7DFEAB5FF9A5308FC1F1FBC299D0292" bold="true" box="[1003,1023,1999,2025]" pageId="11" pageNumber="12">B</emphasis>
. P4. Scale bar: 100 μm.
</paragraph>
</caption>
<subSubSection id="8DB1652CFF9D530EFF4919782F5A0168" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="C51436A7FF9D530FFF491978280D045E" blockId="12.[189,623,267,293]" box="[189,623,267,293]" pageId="12" pageNumber="13">
<heading id="9E5C81CBFF9D530FFF491978280D045E" bold="true" box="[189,623,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="F7DFEAB5FF9D530FFF491978280D045E" bold="true" box="[189,623,267,293]" pageId="12" pageNumber="13">Differential diagnosis and remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C51436A7FF9D530FFF4919492E0807ED" blockId="12.[189,1399,314,662]" pageId="12" pageNumber="13">
Based on the number of segments of the antennules, the progressive oligomerization of P1–P4, and the shape and armature of P5, 
<bibRefCitation id="A13A4B56FF9D530FFDC7192D28AC0403" box="[563,718,349,376]" pageId="12" pageNumber="13" refId="ref14573" refString="Pesce G. L. 1996. Towards a revision of Cyclopinae copepods (Crustacea, Cyclopidae). Fragmenta Entomologica 28: 189 - 200." type="journal article">Pesce (1996)</bibRefCitation>
classified 33 genera in the subfamily 
<taxonomicName id="02AB4D24FF9D530FFB54192D2F440403" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[1184,1318,350,376]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
into 6 groups, including the 
<taxonomicName id="02AB4D24FF9D530FFE3F19F2283E04E0" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[459,604,385,411]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFE3F19F2283E04E0" box="[459,604,385,411]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9D530FFD6B19F2295C04E0" authorityName="Claus" authorityYear="1893" box="[671,830,385,411]" class="Maxillopoda" family="Cyclopidae" genus="Microcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFD6B19F2295C04E0" box="[671,830,385,411]" italics="true" pageId="12" pageNumber="13">Microcyclops</emphasis>
</taxonomicName>
groups, both presented in this paper. Most of the 33 classified genera also include stygobiotic members. Their morphological adaptations for life in subterranean habitats are related to specific environmental characteristics (i.e., lack of light and food scarcity) and include a reduction of body size, 10- or 11-segmented antennules, P1–P4 with three or two segmented Exp, two or one segmented Endp rami, and partly or completely fused P5 to Pd5. The level of reductions express a grade of a transformation of appendages from a plesiomorphic state (P1–P4 with three segmented Exp/Endp and P5 clearly separated from Pd5) to an apomorphic one (P1–P4 with reduced number of segments in Exp/Endp and P5 fused with Pd5) (
<bibRefCitation id="A13A4B56FF9D530FFC261A082E3F07ED" box="[978,1117,635,662]" pageId="12" pageNumber="13" refId="ref14573" refString="Pesce G. L. 1996. Towards a revision of Cyclopinae copepods (Crustacea, Cyclopidae). Fragmenta Entomologica 28: 189 - 200." type="journal article">Pesce 1996</bibRefCitation>
).
</paragraph>
<paragraph id="C51436A7FF9D530FFF491ACD2EAB0162" blockId="12.[189,1399,701,1049]" pageId="12" pageNumber="13">
In stygobiotic 
<taxonomicName id="02AB4D24FF9D530FFE931ACE2B9207AC" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[359,496,701,727]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
, intensive reductions of P5 have a strong effect on establishing a genus level (
<bibRefCitation id="A13A4B56FF9D530FFF311A922BD60780" box="[197,436,737,763]" pageId="12" pageNumber="13" refId="ref14837" refString="Reid J. W. &amp; Ishida T. 2000. Itocyclops, a new genus proposed for Speocyclops yesoensis (Copepoda: Cyclopoida: Cyclopidae). Journal of Crustacean Biology 20: 589 - 596." type="journal article">Reid &amp; Ishida 2000</bibRefCitation>
). For that reason, some other morphological characters should also be included to support the erection of a new genus as well as its position within six groups in the subfamily 
<taxonomicName id="02AB4D24FF9D530FFB1E1B762F110664" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[1258,1395,773,799]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
. Examples of the effect of inclusion of other morphological characters in erecting a new genus and its positioning within six gropus of 
<taxonomicName id="02AB4D24FF9D530FFDB81B3F28B0061D" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[588,722,844,870]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
include the morphologicaly similar genera 
<taxonomicName id="02AB4D24FF9D530FFB111B3F2B2C06F1" authority="Kiefer, 1937" authorityName="Kiefer" authorityYear="1937" class="Maxillopoda" family="Cyclopidae" genus="Speocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFB111B3F2F14061D" box="[1253,1398,844,870]" italics="true" pageId="12" pageNumber="13">Speocyclops</emphasis>
Kiefer, 1937
</taxonomicName>
(a member of 
<taxonomicName id="02AB4D24FF9D530FFE071B0328F006F1" authorityName="Claus" authorityYear="1893" box="[499,658,880,906]" class="Maxillopoda" family="Cyclopidae" genus="Microcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFE071B0328F006F1" box="[499,658,880,906]" italics="true" pageId="12" pageNumber="13">Microcyclops</emphasis>
</taxonomicName>
group) and 
<emphasis id="F7DFEAB5FF9D530FFCEE1B0329C206F1" box="[794,928,880,906]" italics="true" pageId="12" pageNumber="13">Bryocyclop</emphasis>
s Kiefer, 1927 (a member of 
<taxonomicName id="02AB4D24FF9D530FFB111B032F1406F1" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1253,1398,880,906]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFB111B032F1406F1" box="[1253,1398,880,906]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
group). They are similar in body shape and both have very reduced P5 or even have it fused to Pd5 (
<bibRefCitation id="A13A4B56FF9D530FFF311BC42BF006AA" box="[197,402,951,977]" pageId="12" pageNumber="13" refId="ref14504" refString="Pandourski I. 1992. Contribution a l'etude des cyclopoides (Crustacea, Copepoda) des eaux souterraines karstiques de la Bulgarie avec description du Speocyclops rhodopensis sp. n. Acta Zoologica Bulgarica 45: 92 - 101." type="journal article">Pandourski 1992</bibRefCitation>
; 
<bibRefCitation id="A13A4B56FF9D530FFE551BCB28A306AA" box="[417,705,951,978]" pageId="12" pageNumber="13" refId="ref13717" refString="Dussart B. H. &amp; Defaye D. 2001. Introduction to the Copepoda, 2 nd edition. In: Dumont H. J. E. (ed.) Guides to the Identification of the Microinvertebrates of the Continental Waters of the World. Volume 16: 1 - 344. Backhuys Publishers, Leiden." type="journal article">Dussart &amp; Defaye 2001</bibRefCitation>
; 
<bibRefCitation id="A13A4B56FF9D530FFD3B1BC42E5E06AA" box="[719,1084,951,978]" pageId="12" pageNumber="13" refId="ref13956" refString="Galassi D. M. P. &amp; De Laurentiis T. 2004. Little-known cyclopoids from groundwater in Italy: revalidation of Acanthocyclops agamus and redescription of Speocyclops italicus (Crustacea, Copepoda, Cyclopoida). Vie et Milieu 54: 203 - 222." type="journal article">Galassi &amp; De Laurentiis 2004</bibRefCitation>
). The main discrimination character between both genera/groups is the presence/absence of a seta on the inner corner of the coxa in P4. In the genus 
<taxonomicName id="02AB4D24FF9D530FFE6B1B8C28520162" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[415,560,1023,1049]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFE6B1B8C28520162" box="[415,560,1023,1049]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
it is absent, while in the genus 
<taxonomicName id="02AB4D24FF9D530FFC561B8C2E510162" authorityName="Kiefer" authorityYear="1937" box="[930,1075,1023,1049]" class="Maxillopoda" family="Cyclopidae" genus="Speocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFC561B8C2E510162" box="[930,1075,1023,1049]" italics="true" pageId="12" pageNumber="13">Speocyclops</emphasis>
</taxonomicName>
it is present.
</paragraph>
<paragraph id="C51436A7FF9D530FFF491C3229740076" blockId="12.[189,1399,1089,1293]" pageId="12" pageNumber="13">
There are some similar morphological details between the 
<taxonomicName id="02AB4D24FF9D530FFC841C322E6D0120" authorityName="Claus" authorityYear="1893" box="[880,1039,1089,1115]" class="Maxillopoda" family="Cyclopidae" genus="Microcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFC841C322E6D0120" box="[880,1039,1089,1115]" italics="true" pageId="12" pageNumber="13">Microcyclops</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9D530FFBBC1C322EBB0120" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1096,1241,1089,1115]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFBBC1C322EBB0120" box="[1096,1241,1089,1115]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
groups, also. An example is the modification of the P3 Endp-2 spine in the males observed in representatives of 
<taxonomicName id="02AB4D24FF9D530FFF491CFB2B2C01D9" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[189,334,1160,1186]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFF491CFB2B2C01D9" box="[189,334,1160,1186]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
and the Alaskan population of 
<taxonomicName id="02AB4D24FF9D530FFD441CFB2E7E01D8" authority="(Ito, 1953)" authorityName="Ito" authorityYear="1953" baseAuthorityName="Ito" baseAuthorityYear="1953" box="[688,1052,1160,1187]" class="Maxillopoda" family="Cyclopidae" genus="Itocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="yezoensis">
<emphasis id="F7DFEAB5FF9D530FFD441CFB29FF01D9" box="[688,925,1160,1186]" italics="true" pageId="12" pageNumber="13">Itocyclops yezoensis</emphasis>
(Itô, 1953)
</taxonomicName>
(member of the 
<taxonomicName id="02AB4D24FF9D530FFB2C1CFB2F1501D9" authorityName="Claus" authorityYear="1893" box="[1240,1399,1160,1186]" class="Maxillopoda" family="Cyclopidae" genus="Microcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFB2C1CFB2F1501D9" box="[1240,1399,1160,1186]" italics="true" pageId="12" pageNumber="13">Microcyclops</emphasis>
</taxonomicName>
group). However, according to 
<bibRefCitation id="A13A4B56FF9D530FFDC61CDF295001BD" box="[562,818,1196,1222]" pageId="12" pageNumber="13" refId="ref14837" refString="Reid J. W. &amp; Ishida T. 2000. Itocyclops, a new genus proposed for Speocyclops yesoensis (Copepoda: Cyclopoida: Cyclopidae). Journal of Crustacean Biology 20: 589 - 596." type="journal article">Reid &amp; Ishida (2000)</bibRefCitation>
, there are no phylogenetic relationships between the genera 
<taxonomicName id="02AB4D24FF9D530FFEC81CBC2BAF0192" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[316,461,1231,1257]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFEC81CBC2BAF0192" box="[316,461,1231,1257]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9D530FFDF01CBC29130191" authority="Reid &amp; Ishida, 2000" authorityName="Reid &amp; Ishida" authorityYear="2000" box="[516,881,1231,1258]" class="Maxillopoda" family="Cyclopidae" genus="Itocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFDF01CBC281F0192" box="[516,637,1231,1257]" italics="true" pageId="12" pageNumber="13">Itocyclops</emphasis>
<bibRefCitation id="A13A4B56FF9D530FFD771CBC29130191" box="[643,881,1231,1258]" pageId="12" pageNumber="13" refId="ref14837" refString="Reid J. W. &amp; Ishida T. 2000. Itocyclops, a new genus proposed for Speocyclops yesoensis (Copepoda: Cyclopoida: Cyclopidae). Journal of Crustacean Biology 20: 589 - 596." type="journal article">Reid &amp; Ishida, 2000</bibRefCitation>
</taxonomicName>
, and the modification of P3 might be a result of random mutation within the Alaskan population.
</paragraph>
<paragraph id="C51436A7FF9D530FFF491D4628450315" blockId="12.[189,1399,1333,1646]" pageId="12" pageNumber="13">
Absence of a seta on the inner corner of the coxa in P 
<quantity id="02539B42FF9D530FFCD61D46292F0034" box="[802,845,1333,1359]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" pageNumber="13" unit="in" value="4.0">4 in</quantity>
the male and female as well as the modification of the P3 Endp-2 spine in the male place 
<taxonomicName id="02AB4D24FF9D530FFD331D2A29380008" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[711,858,1369,1395]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F7DFEAB5FF9D530FFD331D2A29380008" box="[711,858,1369,1395]" italics="true" pageId="12" pageNumber="13">Siamcyclops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9D530FFC911D2A29B60008" box="[869,980,1369,1395]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
in the 
<taxonomicName id="02AB4D24FF9D530FFBC51D2A2EA00008" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1073,1218,1369,1395]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFBC51D2A2EA00008" box="[1073,1218,1369,1395]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
group. So far, representatives of genera 
<taxonomicName id="02AB4D24FF9D530FFE191D0E281C00EC" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[493,638,1405,1431]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F7DFEAB5FF9D530FFE191D0E281C00EC" box="[493,638,1405,1431]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9D530FFD4D1D0E292E00EC" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[697,844,1405,1431]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F7DFEAB5FF9D530FFD4D1D0E292E00EC" box="[697,844,1405,1431]" italics="true" pageId="12" pageNumber="13">Siamcyclops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9D530FFCA11D0E29A300EC" box="[853,961,1405,1431]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
only differ from other genera within the 
<taxonomicName id="02AB4D24FF9D530FFF121DD32B1500C1" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[230,375,1440,1466]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFF121DD32B1500C1" box="[230,375,1440,1466]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
group by morphological modifications in armature elements on the male P3 Endp-2. The lack of a seta on the inner corner of the coxa of P 
<quantity id="02539B42FF9D530FFD021DB7294000A5" box="[758,802,1476,1502]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" pageNumber="13" unit="in" value="4.0">4 in</quantity>
the male and female, combined with modifications of elements of the P3 Endp- 
<quantity id="02539B42FF9D530FFDFD1D9B28540379" box="[521,566,1512,1538]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="13" unit="in" value="2.0">2 in</quantity>
the male, indicate a close relationship between 
<taxonomicName id="02AB4D24FF9D530FFB981D9B2F420379" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1132,1312,1512,1538]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9D530FFB981D9B2F420379" box="[1132,1312,1512,1538]" italics="true" pageId="12" pageNumber="13">S. cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9D530FFADC1D9B2B74035D" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
and the representatives of 
<taxonomicName id="02AB4D24FF9D530FFDBE1E7828B9035E" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[586,731,1547,1573]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFDBE1E7828B9035E" box="[586,731,1547,1573]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
groups I, II, V and VII (in group VII it is not modification but a difference in the size of the spine between the male and female) (
<bibRefCitation id="A13A4B56FF9D530FFC011E5C2ECA0331" box="[1013,1192,1583,1610]" pageId="12" pageNumber="13" refId="ref14459" refString="Lindberg K. 1956. Les Cyclopides (Crustaces Copepodes) tres evolues en tant qu'habitants des eaux souterraines. Revue des travaux recents concernant les Bryocyclops Kiefer et Speocyclops Kiefer. Actes du Premier Congres International de Speleologie 3: 71 - 83." type="journal article">Lindberg 1956</bibRefCitation>
; Reid 1999; 
<bibRefCitation id="A13A4B56FF9D530FFAC81E5C2A990316" pageId="12" pageNumber="13" refId="ref13864" refString="Fiers F. 2002. The genus Haplocyclops Kiefer, 1952 (Copepoda, Cyclopoida, Cyclopidae): Redescription of the type-species, H. gudrunae, and its congeners. Hydrobiologia 474: 155 - 169. https: // doi. org / 10.1023 / A: 1016579619924" type="journal article">Fiers 2002</bibRefCitation>
; 
<bibRefCitation id="A13A4B56FF9D530FFEF31E2028790315" box="[263,539,1619,1646]" pageId="12" pageNumber="13" refId="ref15316" refString="Watiroyram S., Brancelj A. &amp; Sanoamuang L. 2015. A new cave-dwelling copepod from northeastern Thailand (Cyclopoida: Cyclopidae). The Raffles Bulletin of Zoology 63: 426 - 437." type="journal article">
Watiroyram 
<emphasis id="F7DFEAB5FF9D530FFE6D1E272BB60316" box="[409,468,1619,1645]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="C51436A7FF9D530FFF491EE62BC30219" blockId="12.[189,1399,1685,1890]" pageId="12" pageNumber="13">
A similarity in the shape of the P4 Endp-2 and the presence of six armature elements there (five in other 
<taxonomicName id="02AB4D24FF9D530FFF491ECB2B2C03A9" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[189,334,1720,1746]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFF491ECB2B2C03A9" box="[189,334,1720,1746]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
species) indicate that the new taxon most resembles 
<taxonomicName id="02AB4D24FF9D530FFC381ECA2F1303A8" authority="Monchenko, 1972" authorityName="Monchenko" authorityYear="1972" box="[972,1393,1720,1747]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jankowskajae">
<emphasis id="F7DFEAB5FF9D530FFC381ECA2EED03A9" box="[972,1167,1720,1746]" italics="true" pageId="12" pageNumber="13">B. jankowskajae</emphasis>
Monchenko, 1972
</taxonomicName>
, known from the Kyzylkum Desert in 
<collectingCountry id="BDBC7637FF9D530FFD861EAF2894038D" box="[626,758,1756,1782]" name="Uzbekistan" pageId="12" pageNumber="13">Uzbekistan</collectingCountry>
(formerly part of the USSR). The species was recently moved from the subgenus 
<taxonomicName id="02AB4D24FF9D530FFE021F7329E20261" authority="Monchenko, 1972" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[502,896,1792,1818]" class="Maxillopoda" family="Cyclopidae" genus="Palaeocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="F7DFEAB5FF9D530FFE021F7328C30261" box="[502,673,1792,1818]" italics="true" pageId="12" pageNumber="13">Palaeocyclops</emphasis>
Monchenko, 1972
</taxonomicName>
into 
<taxonomicName id="02AB4D24FF9D530FFC491F732E2C0261" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[957,1102,1792,1818]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="F7DFEAB5FF9D530FFC491F732E2C0261" box="[957,1102,1792,1818]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9D530FFBA11F732EF10261" box="[1109,1171,1792,1818]" pageId="12" pageNumber="13" sensu="stricto">s. str.</taxonomicNameLabel>
(
<bibRefCitation id="A13A4B56FF9D530FFB571F732F580260" box="[1187,1338,1792,1819]" pageId="12" pageNumber="13" refId="ref15216" refString="Walter T. C. 2015. Bryocyclops Kiefer, 1927. In: Walter T. C. &amp; Boxshall G. (eds) World of Copepods Database. Available from http: // marinespecies. org / aphia. php? p = taxdetails &amp; id = 346880 [accessed 13 Sep. 2017]." type="book chapter">Walter 2015</bibRefCitation>
). For that reason, we suggest the establishment of a new group, i.e., group VIII, to accommodate it within the genus 
<taxonomicName id="02AB4D24FF9D530FFEFD1F342BF8021A" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[265,410,1863,1889]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFEFD1F342BF8021A" box="[265,410,1863,1889]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C51436A7FF9D530EFF491FFA2ED70481" blockId="12.[189,1399,1929,2027]" lastBlockId="13.[189,1399,267,506]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
All representatives of the genus 
<taxonomicName id="02AB4D24FF9D530FFDB31FFA28B402D8" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[583,726,1929,1955]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFDB31FFA28B402D8" box="[583,726,1929,1955]" italics="true" pageId="12" pageNumber="13">Bryocyclops</emphasis>
</taxonomicName>
(except 
<taxonomicName id="02AB4D24FF9D530FFCB61FF92EDB02DF" authority="(Menzel, 1926))" authorityName="" authorityYear="1926" baseAuthorityName="Menzel" baseAuthorityYear="1926" box="[834,1209,1929,1956]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bogoriensis">
<emphasis id="F7DFEAB5FF9D530FFCB61FF9298F02D8" box="[834,1005,1929,1955]" italics="true" pageId="12" pageNumber="13">B. bogoriensis</emphasis>
(Menzel, 1926))
</taxonomicName>
and the former member of the subgenus 
<taxonomicName id="02AB4D24FF9D530FFE291FDE28E402BC" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[477,646,1965,1991]" class="Maxillopoda" family="Cyclopidae" genus="Palaeocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9D530FFE291FDE28E402BC" box="[477,646,1965,1991]" italics="true" pageId="12" pageNumber="13">Palaeocyclops</emphasis>
</taxonomicName>
have a pointed prominence on the intercoxal sclerite of P4, while in 
<taxonomicName id="02AB4D24FF9D530FFF2F1FA22BEE0290" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[219,396,2001,2027]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9D530FFF2F1FA22BEE0290" box="[219,396,2001,2027]" italics="true" pageId="12" pageNumber="13">S. cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9D530FFE671FA228200290" box="[403,578,2001,2027]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
it is obtuse. However, the armament on the mandibular palp and distal segment of P4 Exp found in representatives of the genus 
<taxonomicName id="02AB4D24FF9C530EFCA519782982045E" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[849,992,267,293]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFCA519782982045E" box="[849,992,267,293]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
, including a former member of the subgenus 
<emphasis id="F7DFEAB5FF9C530EFEC5195C2BAF0432" box="[305,461,303,329]" italics="true" pageId="13" pageNumber="14">Palaeocylops</emphasis>
, show a pattern of reduction of armature elements. The presence of only one or no seta can be observed in several species within the genus 
<taxonomicName id="02AB4D24FF9C530EFC8019212E610417" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[884,1027,338,364]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFC8019212E610417" box="[884,1027,338,364]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
where the mandibular palp was described or illustrated, as in 
<taxonomicName id="02AB4D24FF9C530EFDF919052EB504F4" authority="Mirabdullayev, Van Damme &amp; Dumont, 2002" authorityName="Mirabdullayev, Van Damme &amp; Dumont" authorityYear="2002" box="[525,1239,373,400]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="saqotraensis">
<emphasis id="F7DFEAB5FF9C530EFDF9190528DD04F4" box="[525,703,374,399]" italics="true" pageId="13" pageNumber="14">B. saqotraensis</emphasis>
Mirabdullayev, 
<collectingRegion id="076FF845FF9C530EFC8F190529CA04EB" box="[891,936,374,400]" country="Turkey" name="Van" pageId="13" pageNumber="14">Van</collectingRegion>
Damme &amp; Dumont, 2002
</taxonomicName>
, 
<taxonomicName id="02AB4D24FF9C530EFB1519052B1704C8" authority="(Menzel, 1926)" authorityName="Menzel" authorityYear="1926" baseAuthorityName="Menzel" baseAuthorityYear="1926" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="F7DFEAB5FF9C530EFB1519052F1504F4" box="[1249,1399,373,399]" italics="true" pageId="13" pageNumber="14">B. muscicola</emphasis>
(Menzel, 1926)
</taxonomicName>
, 
<taxonomicName id="02AB4D24FF9C530EFE7619EA2E5C04C8" authority="Watiroyram, Brancelj &amp; Sanoamuang, 2012" authorityName="Watiroyram, Brancelj &amp; Sanoamuang" authorityYear="2012" box="[386,1086,408,435]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="maewaensis">
<emphasis id="F7DFEAB5FF9C530EFE7619EA285204C9" box="[386,560,409,434]" italics="true" pageId="13" pageNumber="14">B. maewaensis</emphasis>
<bibRefCitation id="A13A4B56FF9C530EFDCC19EB2E5C04C8" box="[568,1086,408,435]" pageId="13" pageNumber="14" refId="ref15275" refString="Watiroyram S., Brancelj A. &amp; Sanoamuang L. 2012. A new Bryocyclops Kiefer (Crustacea: Copepoda: Cyclopoida) from karstic caves in Thailand. The Raffles Bulletin of Zoology 60: 11 - 21." type="journal article">Watiroyram, Brancelj &amp; Sanoamuang, 2012</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9C530EFB8E19EA2F5804C9" authorityName="Monchenko" authorityYear="1972" box="[1146,1338,408,435]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="jankowskajae">
<emphasis id="F7DFEAB5FF9C530EFB8E19EA2EEE04C9" box="[1146,1164,409,434]" italics="true" pageId="13" pageNumber="14">B</emphasis>
. 
<emphasis id="F7DFEAB5FF9C530EFB6819EB2F5804C9" box="[1180,1338,408,434]" italics="true" pageId="13" pageNumber="14">jankowskajae</emphasis>
</taxonomicName>
. The presence of 3 setae on the mandibular palp in 
<taxonomicName id="02AB4D24FF9C530EFD3219CF291C04AD" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[710,894,444,470]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cavernicoulus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9C530EFD3219CF291C04AD" box="[710,894,444,470]" italics="true" pageId="13" pageNumber="14">S.cavernicoulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9C530EFC7019CF2E4C04AD" box="[900,1070,444,470]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
shows a plesiomorphic state of the character, which clearly differentiates it from members of the genus 
<taxonomicName id="02AB4D24FF9C530EFBEB19AC2ECC0482" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1055,1198,479,505]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFBEB19AC2ECC0482" box="[1055,1198,479,505]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C51436A7FF9C530EFF491A6E2F6B07BE" blockId="13.[189,1399,541,709]" pageId="13" pageNumber="14">
Furthermore, the spine/seta formula 3.3.3.2/5.5.5.5 found in the new taxon differs clearly from the formulae of the species within the genus 
<taxonomicName id="02AB4D24FF9C530EFDC21A3328A70721" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[566,709,576,602]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFDC21A3328A70721" box="[566,709,576,602]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
, where the spine/seta formula varied from 2.2.2.2/ 5.4.4.3 (group V) to 2.3.3.3/ 5(4).4.4.3 (group VI), 3.3.3.2/5.4.4.4 (group III), 3.3.3.3/ 5.5.5.4 (groups I and VII), 3.3.3.3(4) /5(4).5.5.4 (group II), and 3.3.3.4 /5.4.4.4 (group IV) (
<bibRefCitation id="A13A4B56FF9C530EFC561AF42E3307DA" box="[930,1105,647,673]" pageId="13" pageNumber="14" refId="ref14424" refString="Lindberg K. 1947. Cyclopides (Crustaces Copepodes) nouveaux de l'Inde. I. Description des premiers Bryocyclops decouverts dans l'Inde. Records of the Indian Museum 45: 47 - 55." type="journal article">Lindberg 1947</bibRefCitation>
; 
<bibRefCitation id="A13A4B56FF9C530EFBA81AF42F0807DA" box="[1116,1386,646,673]" pageId="13" pageNumber="14" refId="ref15316" refString="Watiroyram S., Brancelj A. &amp; Sanoamuang L. 2015. A new cave-dwelling copepod from northeastern Thailand (Cyclopoida: Cyclopidae). The Raffles Bulletin of Zoology 63: 426 - 437." type="journal article">
Watiroyram 
<emphasis id="F7DFEAB5FF9C530EFB181AFB2F4707DA" box="[1260,1317,647,673]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2015
</bibRefCitation>
). It also differs considerably from 
<taxonomicName id="02AB4D24FF9C530EFDCD1AD8289507BF" authorityName="Monchenko" authorityYear="1972" box="[569,759,682,708]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="jankowskajae">
<emphasis id="F7DFEAB5FF9C530EFDCD1AD8289507BF" box="[569,759,682,708]" italics="true" pageId="13" pageNumber="14">B. jankowskajae</emphasis>
</taxonomicName>
whose spine/setae formula is 2.3.3.3/ 5.5.5.4.
</paragraph>
<paragraph id="C51436A7FF9C530EFF491A9B28E706F4" blockId="13.[189,1399,744,911]" pageId="13" pageNumber="14">
Normally, the spermatophores within the subfamily 
<taxonomicName id="02AB4D24FF9C530EFCD51A9B29C50679" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[801,935,744,770]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
are bean- or kidney-shaped. They have also been illustrated in some species of 
<taxonomicName id="02AB4D24FF9C530EFD7B1B782942065E" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[655,800,779,805]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFD7B1B782942065E" box="[655,800,779,805]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
such as 
<taxonomicName id="02AB4D24FF9C530EFC701B7F2ED3065D" authority="Bjornberg, 1985" authorityName="Bjornberg" authorityYear="1985" box="[900,1201,779,806]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="caroli">
<emphasis id="F7DFEAB5FF9C530EFC701B7F2988065E" box="[900,1002,779,805]" italics="true" pageId="13" pageNumber="14">B. caroli</emphasis>
Bjornberg, 1985
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9C530EFB1E1B7F2B520633" authority="Por, 1981" authorityName="Por" authorityYear="1981" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absalomi">
<emphasis id="F7DFEAB5FF9C530EFB1E1B7F2F15065E" box="[1258,1399,779,805]" italics="true" pageId="13" pageNumber="14">B. absalomi</emphasis>
Por, 1981
</taxonomicName>
. The unique, three dimensional L-flip feature of the spermatophore observed in 
<taxonomicName id="02AB4D24FF9C530EFB3F1B5D2F150633" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1227,1399,814,840]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cavernicolus" status="gen. et sp. nov.">
<emphasis id="F7DFEAB5FF9C530EFB3F1B5D2F150633" box="[1227,1399,814,840]" italics="true" pageId="13" pageNumber="14">S.cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9C530EFF491B212B170617" box="[189,373,850,876]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from those of other members of the genus 
<taxonomicName id="02AB4D24FF9C530EFC2F1B212E0E0617" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[987,1132,850,876]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFC2F1B212E0E0617" box="[987,1132,850,876]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
as well as from other members of the subfamily 
<taxonomicName id="02AB4D24FF9C530EFE031B0628E206F4" baseAuthorityName="Pocock" baseAuthorityYear="1924" box="[503,640,885,911]" class="Maxillopoda" family="Cyclopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Cyclopinae">Cyclopinae</taxonomicName>
.
</paragraph>
<paragraph id="C51436A7FF9C530EFF491BC02F5A0168" blockId="13.[189,1399,946,1044]" pageId="13" pageNumber="14">
For the reasons listed above, we propose the establishment of the new genus 
<taxonomicName id="02AB4D24FF9C530EFBA51BC02E8606B6" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[1105,1252,947,973]" class="Maxillopoda" family="Cyclopidae" genus="Siamcyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F7DFEAB5FF9C530EFBA51BC02E8606B6" box="[1105,1252,947,973]" italics="true" pageId="13" pageNumber="14">Siamcyclops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9C530EFB181BC02F3A06B6" box="[1260,1368,947,973]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
to accommodate the new stygobiotic species from 
<collectingCountry id="BDBC7637FF9C530EFD121BA52930068B" box="[742,850,982,1008]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
. The main differences between the three related taxa, 
<taxonomicName id="02AB4D24FF9C530EFF081B8A2BEF0168" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[252,397,1017,1043]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="F7DFEAB5FF9C530EFF081B8A2BEF0168" box="[252,397,1017,1043]" italics="true" pageId="13" pageNumber="14">Bryocyclops</emphasis>
</taxonomicName>
spp., 
<taxonomicName id="02AB4D24FF9C530EFE201B8A296E0168" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[468,780,1017,1043]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="jankowskajae" status="gen. nov.">
<emphasis id="F7DFEAB5FF9C530EFE201B8A296E0168" box="[468,780,1017,1043]" italics="true" pageId="13" pageNumber="14">Bryocyclops jankowskajae</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AB4D24FF9C530EFCB11B8A29A90168" authority="Boonyanusith &amp; Sanoamuang &amp; Brancelj, 2018" authorityName="Boonyanusith &amp; Sanoamuang &amp; Brancelj" authorityYear="2018" box="[837,971,1017,1043]" class="Maxillopoda" family="Cyclopidae" genus="Siamcylops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F7DFEAB5FF9C530EFCB11B8A29A90168" box="[837,971,1017,1043]" italics="true" pageId="13" pageNumber="14">Siamcylops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEC57CEFF9C530EFC251B892E5D016F" box="[977,1087,1018,1044]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
, are listed in 
<tableCitation id="8829031CFF9C530EFB2E1B8A2F500168" box="[1242,1330,1017,1043]" captionStart="Table 1" captionStartId="14.[189,258,268,294]" captionTargetBox="[203,1367,555,1591]" captionText="Table 1. Differential diagnosis between the genus Bryocyclops* Kiefer, 1927, Bryocyclops jankowskajae Monchenko, 1972 and Siamcyclops gen. nov. (Roman numbers = spine; Arabic numbers = seta). * only members of groups I and II (according to Lindberg 1947) are included; details on other groups of spine/seta formulae are in the text † in B. fidjiensis Lindberg, 1954 †† in B. campaneri Rocha &amp; Carvalho Bjornberg, 1987 ††† in B. bogoriensis (Menzel, 1926)" httpUri="http://table.plazi.org/id/91D4662FFF9F530DFF49197F28160772" pageId="13" pageNumber="14" tableUuid="91D4662FFF9F530DFF49197F28160772">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>