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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis &amp; Ribeiro, Guilherme Cunha" docDate="2022" docId="4B6DCC4B12E451DD8560DA080729125A" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) anchoralis Santos &amp; Santos &amp; Ribeiro 2022, sp. nov." docType="treatment" docUuid="87DAED38-FF8B-48DC-8957-1F60A208E890" docUuidSource="ZooBank" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Daubian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>439</mods:start>
<mods:end>494</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:87DAED38-FF8B-48DC-8957-1F60A208E890" httpUri="http://treatment.plazi.org/id/4B6DCC4B12E451DD8560DA080729125A" lastPageNumber="439" pageId="0" pageNumber="439">
<subSubSection pageId="0" pageNumber="439" type="nomenclature">
<paragraph pageId="0" pageNumber="439">
3.3.3.
<taxonomicName LSID="http://zoobank.org/87DAED38-FF8B-48DC-8957-1F60A208E890" authority="Santos &amp; Santos &amp; Ribeiro, 2022" authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) anchoralis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="anchoralis" status="sp. nov." subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) anchoralis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="439">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="439">
<materialsCitation collectingDate="1952-02-12" country="Chile" county="Nahuelbuta" latitude="-37.65" location="Palo Botado" longLatPrecision="1180" longitude="-72.9" municipality="Contulmo" specimenCount="♂" stateProvince="Arauco" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<specimenCount></specimenCount>
,
<collectingCountry name="Chile">Chile</collectingCountry>
,
<collectingRegion country="Chile" name="Araucania">Arauco</collectingRegion>
,
<collectingCounty>Nahuelbuta</collectingCounty>
,
<collectingMunicipality>Contulmo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:4B6DCC4B12E451DD8560DA080729125A:C3071B36C275CA8DE0F501ADDEF849B4" country="Chile" county="Nahuelbuta" latitude="-37.65" longLatPrecision="1180" longitude="-72.9" municipality="Contulmo" name="Palo Botado" stateProvince="Arauco">Palo Botado</location>
[
<geoCoordinate degrees="37" direction="south" minutes="39" orientation="latitude" precision="925" value="-37.65">37°39S</geoCoordinate>
<geoCoordinate degrees="72" direction="west" minutes="54" orientation="longitude" precision="925" value="-72.9">72°54W</geoCoordinate>
],
<collectingDate value="1952-02-12">12-II-1952</collectingDate>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)
</materialsCitation>
*.
<materialsCitation collectingDate="1952-01-30" collectingDateMax="1953-02-02" collectingDateMin="1952-01-30" collectorName="Chiloe Is., Pena" country="Chile" latitude="-42.1" location="Rio Coluco" longLatPrecision="1152" longitude="-73.916664" municipality="Nahuelbuta" specimenCount="1" specimenCount-male="1" stateProvince="Arauco" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<specimenCount type="male">1 ♂</specimenCount>
and 1 [sex unknown],
<collectingCountry name="Chile">Chile</collectingCountry>
,
<collectingRegion country="Chile" name="Araucania">Arauco</collectingRegion>
,
<collectingMunicipality>Nahuelbuta</collectingMunicipality>
[
<geoCoordinate degrees="37" direction="south" minutes="46" orientation="latitude" precision="925" value="-37.766666">37°46S</geoCoordinate>
<geoCoordinate degrees="72" direction="west" minutes="59" orientation="longitude" precision="925" value="-72.98333">72°59W</geoCoordinate>
],
<collectingDate value="1953-02-02">2-II-1953</collectingDate>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM); 1 [sex unknown],
<collectingCountry name="Chile">Chile</collectingCountry>
,
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:4B6DCC4B12E451DD8560DA080729125A:F92B1EE5D3E8C844F6723275A2DBBC05" country="Chile" latitude="-42.1" longLatPrecision="1152" longitude="-73.916664" municipality="Nahuelbuta" name="Rio Coluco" stateProvince="Arauco">Rio Coluco</location>
[
<geoCoordinate degrees="42" direction="south" minutes="6" orientation="latitude" precision="925" value="-42.1">42°6S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="55" orientation="longitude" precision="925" value="-73.916664">73°55W</geoCoordinate>
],
<collectingDate value="1952-01-30">30-I-1952</collectingDate>
,
<normalizedToken originalValue="Peña">
<collectorName>Pena</collectorName>
</normalizedToken>
(USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="etymology">
<paragraph pageId="0" pageNumber="439">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="439">
The word
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus anchoralis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="anchoralis">
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus anchoralis</emphasis>
</taxonomicName>
is Latin for &quot;of the anchor&quot;. The term refers to the shape of the projection of the male 9th tergite, which resembles an anchor.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="439">The species is distinguished by the absence of markings near R1, and R2+3 shorter than R2. Furthermore, this species has a long vertical spur on the sheath of aedeagus, male tergite IX has a distinctive ridge projection, mesal lobes absent and the posterior margin of the male tergite IX is mostly unpigmented.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="description">
<paragraph pageId="0" pageNumber="439">Description.</paragraph>
<paragraph pageId="0" pageNumber="439">
Wing length 6.15 mm, width 1.86 mm. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
: General coloration yellowish brown. Rostrum and palpus dark brown. First segments of antenna pale, remaining segments dark yellow. Head brownish-black. Eyes black. Thorax yellowish-brown, pleura pale. Scutum without stripes. Halter pale with knobs light orange. Coxae and legs brownish. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
(dorsal view Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10B</figureCitation>
, ventral view Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10A</figureCitation>
): Rostrum bifurcated with short appendices; first palpal segment as long as terminal segment, second palpal segment dilated; scape thinner than pedicel, terminal flagellomere dilated, antenna with several long setae. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
(lateral view Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10E</figureCitation>
, dorsal view Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10F</figureCitation>
): a few setae on prescutum. Division line of laterotergite almost vertical; anatergite as tall as katatergite. Halter with dilated knob. Wing (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10D</figureCitation>
) almost clear of markings, except marking covering R2; veins near fork of bM (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10C</figureCitation>
) faded; M1+2 longer than basal deflection of M1; R2+3+4 at least twice as long as R2+3. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439">10G</figureCitation>
): Male tergite IX with posterior margin with irregular ridges; median U-shaped notch with straight projection in middle; tip of projection also ridge shaped; projection and surrounding area darkened. Ventral branch of gonocoxite inclined laterally. Dorsal branch of gonocoxite half-length of ventral branch, atrophied and pointed medially; both branches setose. Lobe of gonostylus with pointed lobule, shaped similar to folded leaf. Clasper of gonostylus with differently shaped branches: lateral branch long, bent near middle; medial branch curved into stub. Mesal lobes absent. Sheath of aedeagus darkened on apical half, angulated, with long pointed projection perpendicular to remainder of sheath.
</paragraph>
<caption doi="10.3897/asp.80.e83035.figure10" httpUri="https://binary.pensoft.net/fig/740563" pageId="0" pageNumber="439" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">Figure 10.</emphasis>
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) anchoralis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="anchoralis" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) anchoralis</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
head (ventral view);
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
head (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
detail of wing;
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
wing;
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
thorax (lateral view);
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
thorax (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
, sheath of aedeagus;
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
, anatergite;
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
, anepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
, anepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
, anteropronotum;
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
, clasper of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
, coxa;
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
, gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
, halter;
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
, knob of the halter;
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
, katepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
, katepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
, katatergite;
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
, left mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
; lobe of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
, meron;
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
, metanepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
, metepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
, mediotergite;
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
, metakatepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
, metanotum;
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
, posterior basalare;
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
, paratergite;
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
, postpronotum;
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
, prescutum;
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
, right mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
, scutum;
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
, scutellum;
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
, male tergite IX.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="remarks">
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="439">
Specimens of this new species were previously identified by C.P. Alexander as
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) sanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="sanus" subGenus="R.">A. (R.) sanus</taxonomicName>
Alexander and
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) insanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="insanus" subGenus="R.">A. (R.) insanus</taxonomicName>
Alexander. This species resembles
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) sanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="sanus" subGenus="R.">A. (R.) sanus</taxonomicName>
but differs mainly in M1+2, shape of projection in male tergite IX, ventral branch of gonocoxite and spur of sheath of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>