275 lines
30 KiB
XML
275 lines
30 KiB
XML
<document id="F7A2D558D3D331C1ED3C959193DBD3FE" ID-DOI="10.5281/zenodo.5161214" ID-ISSN="1942-1354" ID-Zenodo-Dep="5161214" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628116654208" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2011" docId="4B0D3211FFB6FF9DFF49FE393AE72F5E" docLanguage="en" docName="InsectaMundi.0191.1-13.pdf" docOrigin="Insecta Mundi 2011 (191)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cicindela antatsima var. fotsy Alluaud 1913" docType="treatment" docVersion="4" lastPageNumber="4" masterDocId="B7344A69FFB3FF98FF89FFFA3B7B2A65" masterDocTitle="Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1699035013485" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="074CCDB72A72FEE333B5E5454ED4DF93">Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="91FEACD67A95F1FC6F714281B9D03B55">Mawdsley, Jonathan R.</mods:namePart>
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<mods:title id="049D086B17F67C3E12AA730425AE4806">Insecta Mundi</mods:title>
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<mods:part id="65492089C50017205A39ECDC03D4EC2A">
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<mods:date id="1C2D9CF4DF4A6719608D42F72EC48033">2011</mods:date>
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<mods:number id="271E544D852A141814FEA3EC48B676BA">2011-09-02</mods:number>
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<mods:number id="9E0EC7789D00164F3801F157A7335D61">2011</mods:number>
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<mods:identifier id="0FA723A2A83DF242198E5EB3366A3C45" type="DOI">10.5281/zenodo.5161214</mods:identifier>
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<treatment id="4B0D3211FFB6FF9DFF49FE393AE72F5E" ID-DOI="http://doi.org/10.5281/zenodo.5164920" ID-Zenodo-Dep="5164920" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E" httpUri="http://treatment.plazi.org/id/4B0D3211FFB6FF9DFF49FE393AE72F5E" lastPageNumber="4" pageId="5" pageNumber="4">
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<subSubSection id="8BBED08CFFB6FF9DFF49FE393E412BBE" box="[192,1338,451,475]" pageId="5" pageNumber="4" type="nomenclature">
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<paragraph id="C31B8307FFB6FF9DFF49FE393E412BBE" blockId="5.[192,1338,451,475]" box="[192,1338,451,475]" pageId="5" pageNumber="4">
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<heading id="9853346BFFB6FF9DFF49FE393E412BBE" box="[192,1338,451,475]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="8">
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<emphasis id="F1D05F15FFB6FF9DFF49FE393A322BBE" bold="true" box="[192,329,451,475]" pageId="5" pageNumber="4">Synonym.</emphasis>
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<taxonomicName id="04A4F884FFB6FF9DFEC6FE3938C12BBE" ID-CoL="5YT5R" authority="Alluaud (1913: 495)" authorityName="Alluaud" authorityPageNumber="495" authorityYear="1913" box="[335,954,451,475]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="variety" species="antatsima" variety="fotsy">
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<emphasis id="F1D05F15FFB6FF9DFEC6FE3939322BBE" box="[335,585,451,475]" italics="true" pageId="5" pageNumber="4">Cicindela antatsima</emphasis>
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var.
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<emphasis id="F1D05F15FFB6FF9DFD0FFE3939C42BBE" box="[646,703,451,475]" italics="true" pageId="5" pageNumber="4">fotsy</emphasis>
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<bibRefCitation id="A735FEF6FFB6FF9DFD4CFE3938C12BBE" author="Alluaud, C." box="[709,954,451,475]" pageId="5" pageNumber="4" pagination="494 - 495" refId="ref9629" refString="Alluaud, C. 1913. Diagnose d'une race nouvelle de Cicindela antatsima Alluaud (Col. Cicindelidae). Bulletin de la Societe Entomologique de France 1913: 494 - 495." type="journal article" year="1913">Alluaud (1913: 495)</bibRefCitation>
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</taxonomicName>
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. Synonymy by
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<bibRefCitation id="A735FEF6FFB6FF9DFBFDFE3E3E4D2BBE" author="Horn, W." box="[1140,1334,451,475]" pageId="5" pageNumber="4" pagination="7 - 28" refId="ref9919" refString="Horn, W. 1934. Les Cicindelides de Madagascar, Premiere partie, Catalogue bibligraphique et synonymique. Memoires de l'Academie Malgache 20: 7 - 28." type="journal article" year="1934">Horn (1934: 19)</bibRefCitation>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8BBED08CFFB6FF9DFF49FDF93A3F283E" pageId="5" pageNumber="4" type="materials_examined">
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<paragraph id="C31B8307FFB6FF9DFF49FDF93A3F283E" blockId="5.[192,1440,515,603]" pageId="5" pageNumber="4">
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<emphasis id="F1D05F15FFB6FF9DFF49FDF93AFD287E" bold="true" box="[192,390,515,539]" pageId="5" pageNumber="4">Type material.</emphasis>
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<typeStatus id="1C1F3DA5FFB6FF9DFE02FDF93A94287E" box="[395,495,515,539]" pageId="5" pageNumber="4" type="syntype">Syntype</typeStatus>
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male of
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<taxonomicName id="04A4F884FFB6FF9DFDD9FDFE3990287E" authorityName="Alluaud" authorityYear="1902" box="[592,747,516,539]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antatsima">
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<emphasis id="F1D05F15FFB6FF9DFDD9FDFE3990287E" box="[592,747,516,539]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
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</taxonomicName>
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, labeled “
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<collectingCountry id="BBB3C397FFB6FF9DFCD6FDF93888287E" box="[863,1011,515,539]" name="Madagascar" pageId="5" pageNumber="4">Madagascar</collectingCountry>
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(Sud)/Bassin du Mandraré/
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<bibRefCitation id="A735FEF6FFB6FF9DFAB7FDF93B87285E" author="Alluaud, C." pageId="5" pageNumber="4" pagination="509" refId="ref9544" refString="Alluaud, C. 1900. Liste des Insectes Coleopteres de la Region Malgache. p. vii + 509. In: A. Grandidier (ed.). Histoire Physique, Naturelle et Politique de Madagascar. Volume XXI. Histoire Naturelle des Coleopteres. I (l), p. vii + 509." type="book chapter" year="1900">Alluaud 1900</bibRefCitation>
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44” (
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<collectionCode id="A5B51BC2FFB6FF9DFEB2FDD93AFE285E" box="[315,389,547,571]" pageId="5" pageNumber="4">DEIC</collectionCode>
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, examined);
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<typeStatus id="1C1F3DA5FFB6FF9DFDABFDD939F3285E" box="[546,648,547,571]" pageId="5" pageNumber="4" type="syntype">Syntype</typeStatus>
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female of
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<taxonomicName id="04A4F884FFB6FF9DFC8EFDDE3F5A285E" box="[775,1057,547,571]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="variety" species="antatsima" variety="fotsy">
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<emphasis id="F1D05F15FFB6FF9DFC8EFDDE38D3285E" box="[775,936,548,571]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
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var.
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<emphasis id="F1D05F15FFB6FF9DFC61FDD93F5A285E" box="[1000,1057,547,571]" italics="true" pageId="5" pageNumber="4">fotsy</emphasis>
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</taxonomicName>
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, labeled “Mananpatra” (
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<collectionCode id="A5B51BC2FFB6FF9DFAD8FDD93EE0285E" box="[1361,1435,547,571]" pageId="5" pageNumber="4">DEIC</collectionCode>
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, examined).
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</paragraph>
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</subSubSection>
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<subSubSection id="8BBED08CFFB6FF9DFF49FD7939642E1E" pageId="5" pageNumber="4" type="diagnosis">
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<paragraph id="C31B8307FFB6FF9DFF49FD793F0C295E" blockId="5.[192,1440,643,827]" pageId="5" pageNumber="4">
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<emphasis id="F1D05F15FFB6FF9DFF49FD793A2A28FE" bold="true" box="[192,337,643,667]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
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Length 10.5-12.5 mm. Dorsal coloration as shown in
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<figureCitation id="5B9F9F82FFB6FF9DFC71FD793FEB28FE" box="[1016,1168,643,667]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="5" pageNumber="4">Figures 1, 2</figureCitation>
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. The extensive white coloration on the elytra separates this species from sympatric species of
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<taxonomicName id="04A4F884FFB6FF9DFBA4FD593FC128DE" authorityName="Jeannel. Characters" authorityYear="1946" box="[1069,1210,675,699]" class="Insecta" family="Carabidae" genus="Chaetodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="F1D05F15FFB6FF9DFBA4FD593FC128DE" box="[1069,1210,675,699]" italics="true" pageId="5" pageNumber="4">Chaetodera</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="04A4F884FFB6FF9DFB7CFD593E2628DE" box="[1269,1373,675,699]" class="Insecta" family="Carabidae" genus="Lophyra" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="F1D05F15FFB6FF9DFB7CFD593E2628DE" box="[1269,1373,675,699]" italics="true" pageId="5" pageNumber="4">Lophyra</emphasis>
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</taxonomicName>
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. It is most closely related to the Madagascan
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<taxonomicName id="04A4F884FFB6FF9DFD27FD3E385628BE" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[686,813,708,731]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
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<emphasis id="F1D05F15FFB6FF9DFD27FD3E385628BE" box="[686,813,708,731]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
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</taxonomicName>
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, from which it may be distinguished by the elytral color characteristics mentioned in the key to species (the elytral white areas are broadly expanded in
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<taxonomicName id="04A4F884FFB6FF9DFA0CFD1E3A45297E" authorityName="Alluaud" authorityYear="1902" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antatsima">
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<emphasis id="F1D05F15FFB6FF9DFA0CFD1E3A45297E" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
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</taxonomicName>
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, covering much of the disc, whereas the white areas on the elytra of
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<taxonomicName id="04A4F884FFB6FF9DFB3AFCFE3E4D297E" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[1203,1334,772,795]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
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<emphasis id="F1D05F15FFB6FF9DFB3AFCFE3E4D297E" box="[1203,1334,772,795]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
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</taxonomicName>
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are less extensive and tend to form alternating bands with the dark ground coloration).
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</paragraph>
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<paragraph id="C31B8307FFB6FF9DFF49FC9939642E1E" blockId="5.[192,1440,867,1147]" pageId="5" pageNumber="4">
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<emphasis id="F1D05F15FFB6FF9DFF49FC993A6D291E" bold="true" box="[192,278,867,891]" pageId="5" pageNumber="4">Notes.</emphasis>
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<bibRefCitation id="A735FEF6FFB6FF9DFE92FC9E3AD0291E" author="Horn, W." box="[283,427,867,891]" pageId="5" pageNumber="4" pagination="7 - 28" refId="ref9919" refString="Horn, W. 1934. Les Cicindelides de Madagascar, Premiere partie, Catalogue bibligraphique et synonymique. Memoires de l'Academie Malgache 20: 7 - 28." type="journal article" year="1934">Horn (1934)</bibRefCitation>
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and
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<bibRefCitation id="A735FEF6FFB6FF9DFE6CFC9939D6291E" author="Olsoufieff, G." box="[485,685,867,891]" pageId="5" pageNumber="4" pagination="31 - 76" refId="ref10675" refString="Olsoufieff, G. 1934. Les Cicindelides de Madagascar, Premiere partie, Essai de revision systematique et biologie. Memoires d l'Academie Malgache 20: 31 - 76 + 3 pls." type="book chapter" year="1934">Olsoufieff (1934)</bibRefCitation>
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considered this species to be a regional variant or subspecies of
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<taxonomicName id="04A4F884FFB6FF9DFF49FC7E3A4429FE" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[192,319,900,923]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
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<emphasis id="F1D05F15FFB6FF9DFF49FC7E3A4429FE" box="[192,319,900,923]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
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</taxonomicName>
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, while
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<bibRefCitation id="A735FEF6FFB6FF9DFE10FC79392B29FE" author="Jeannel, R. G." box="[409,592,899,923]" pageId="5" pageNumber="4" refId="ref9981" refString="Jeannel, R. G. 1946. Faune de l'Empire Francaise. VI. Coleopteres carabiques da la region Malgache (Premiere Partie). Museum National d'Histoire Naturelle; Paris. 372 p." type="book" year="1946">Jeannel (1946)</bibRefCitation>
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treated these two taxa as separate species. Based on the specimens examined for this study, I concur with Jeannel’s assessment. The differences in color pattern between
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<taxonomicName id="04A4F884FFB6FF9DFA0FFC5E3A4529BE" authorityName="Alluaud" authorityYear="1902" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antatsima">
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<emphasis id="F1D05F15FFB6FF9DFA0FFC5E3A4529BE" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
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</taxonomicName>
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(
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<figureCitation id="5B9F9F82FFB6FF9DFEC5FC393AB129BE" box="[332,458,963,987]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="5" pageNumber="4">Figure 1-2</figureCitation>
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) and
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<taxonomicName id="04A4F884FFB6FF9DFD9DFC3E39EA29BE" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[532,657,964,987]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
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<emphasis id="F1D05F15FFB6FF9DFD9DFC3E39EA29BE" box="[532,657,964,987]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="5B9F9F82FFB6FF9DFD16FC39385B29BE" box="[671,800,963,987]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="5" pageNumber="4">Figure 3-4</figureCitation>
|
||
), although subtle, appear to be consistent among the specimens available for study. I was unable to find any specimens that represented intermediate forms that might be expected if subspecific variation was involved. I confirm that
|
||
<taxonomicName id="04A4F884FFB6FF9DFBE2FBFE3EFF2E7E" box="[1131,1412,1027,1051]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="variety" species="antatsima" variety="fotsy">
|
||
<emphasis id="F1D05F15FFB6FF9DFBE2FBFE3E772E7E" box="[1131,1292,1028,1051]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
|
||
var.
|
||
<emphasis id="F1D05F15FFB6FF9DFAC2FBF93EFF2E7E" box="[1355,1412,1027,1051]" italics="true" pageId="5" pageNumber="4">fotsy</emphasis>
|
||
</taxonomicName>
|
||
is conspecific with
|
||
<taxonomicName id="04A4F884FFB6FF9DFE0AFBDE39662E5E" authorityName="Alluaud" authorityYear="1902" box="[387,541,1060,1083]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antatsima">
|
||
<emphasis id="F1D05F15FFB6FF9DFE0AFBDE39662E5E" box="[387,541,1060,1083]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
|
||
</taxonomicName>
|
||
based on examination of the relevant
|
||
<typeStatus id="1C1F3DA5FFB6FF9DFC57FBDE3F682E59" box="[990,1043,1060,1084]" pageId="5" pageNumber="4">type</typeStatus>
|
||
specimens. Specimens identified as
|
||
<taxonomicName id="04A4F884FFB6FF9DFF69FBBE3A942E3E" box="[224,495,1091,1115]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="variety" species="antatsima" variety="fotsy">
|
||
<emphasis id="F1D05F15FFB6FF9DFF69FBBE3A072E3E" box="[224,380,1092,1115]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
|
||
var.
|
||
<emphasis id="F1D05F15FFB6FF9DFE3EFBB93A942E3E" box="[439,495,1091,1115]" italics="true" pageId="5" pageNumber="4">fotsy</emphasis>
|
||
</taxonomicName>
|
||
in collections are individuals of
|
||
<taxonomicName id="04A4F884FFB6FF9DFCE2FBBE3F7C2E3E" authorityName="Alluaud" authorityYear="1902" box="[875,1031,1092,1115]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antatsima">
|
||
<emphasis id="F1D05F15FFB6FF9DFCE2FBBE3F7C2E3E" box="[875,1031,1092,1115]" italics="true" pageId="5" pageNumber="4">C. antatsima</emphasis>
|
||
</taxonomicName>
|
||
in which the dark markings of the elytra are especially reduced.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8BBED08CFFB6FF9DFF49FB593AE72F5E" pageId="5" pageNumber="4" type="materials_examined">
|
||
<paragraph id="C31B8307FFB6FF9DFF49FB593AE72F5E" blockId="5.[192,1440,1187,1339]" pageId="5" pageNumber="4">
|
||
<emphasis id="F1D05F15FFB6FF9DFF49FB593AB32EDE" bold="true" box="[192,456,1187,1211]" pageId="5" pageNumber="4">Material examined.</emphasis>
|
||
<materialsCitation id="73CC895AFFB6FF9DFE44FB59388A2EDE" box="[461,1009,1187,1211]" collectionCode="DEIC" country="Madagascar" location="Ambositra" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1">
|
||
<collectingCountry id="BBB3C397FFB6FF9DFE44FB5939EC2EDE" box="[461,663,1187,1211]" name="Madagascar" pageId="5" pageNumber="4">MADAGASCAR</collectingCountry>
|
||
:
|
||
<location id="C67BD5DCFFB6FF9DFD29FB59385B2EDE" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFD29FB59385B2EDE" box="[672,800,1187,1211]" country="Madagascar" name="Ambositra" pageId="5" pageNumber="4">Ambositra</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFCB9FB5938EC2EDE" box="[816,919,1187,1211]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFC29FB5938922EDE" box="[928,1001,1187,1211]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFC72FB593A692EBE" collectionCode="DEIC" country="Madagascar" location="Ampandrandava" pageId="5" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="C67BD5DCFFB6FF9DFC72FB593FB32EDE" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFC72FB593FB32EDE" box="[1019,1224,1187,1211]" country="Madagascar" name="Ampandrandava" pageId="5" pageNumber="4">Ampandrandava</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFB50FB593E522EDE" box="[1241,1321,1187,1211]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="D5A2488EFFB6FF9DFABAFB593EE72EDE" box="[1331,1436,1187,1211]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFF49FB393A712EBE" box="[192,266,1219,1243]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFE97FB3939282EBE" box="[286,595,1219,1243]" collectionCode="DEIC" country="Madagascar" location="Beroroha" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1">
|
||
<location id="C67BD5DCFFB6FF9DFE97FB393AEA2EBE" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFE97FB393AEA2EBE" box="[286,401,1219,1243]" country="Madagascar" name="Beroroha" pageId="5" pageNumber="4">Beroroha</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFE2AFB393A8D2EBE" box="[419,502,1219,1243]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFD88FB3939302EBE" box="[513,587,1219,1243]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFDD6FB3E3F402EBE" box="[607,1083,1219,1243]" collectionCode="DEIC" country="Madagascar" location="Manampatra" pageId="5" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="C67BD5DCFFB6FF9DFDD6FB3E38792EBE" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFDD6FB3E38792EBE" box="[607,770,1220,1243]" country="Madagascar" name="Manampatra" pageId="5" pageNumber="4">Manampatra</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFC9DFB39381C2EBE" box="[788,871,1219,1243]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="D5A2488EFFB6FF9DFCFBFB3938A52EBE" box="[882,990,1219,1243]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFC60FB393F482EBE" box="[1001,1075,1219,1243]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFBCEFB3939392E9E" collectingDate="1933-01" collectionCode="DEIC" country="Madagascar" location="Mandrare" pageId="5" pageNumber="4" specimenCount="2" specimenCount-female="2">
|
||
<location id="C67BD5DCFFB6FF9DFBCEFB393FBD2EBE" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFBCEFB393FBD2EBE" box="[1095,1222,1219,1243]" country="Madagascar" name="Mandrare" pageId="5" pageNumber="4">Mandrare</location>
|
||
,
|
||
<date id="B71AA5C7FFB6FF9DFB5BFB393E5B2EBE" box="[1234,1312,1219,1243]" pageId="5" pageNumber="4" value="1933-01">
|
||
<collectingDate id="A75E5C2FFFB6FF9DFB5BFB393E5B2EBE" box="[1234,1312,1219,1243]" pageId="5" pageNumber="4" value="1933-01">I.1933</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFABBFB393EE02EBE" box="[1330,1435,1219,1243]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFF49FB193A712E9E" box="[192,266,1251,1275]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
), 1933 (
|
||
<specimenCount id="D5A2488EFFB6FF9DFEFAFB193A9B2E9E" box="[371,480,1251,1275]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFE79FB1939412E9E" box="[496,570,1251,1275]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFDD8FB193F422E9E" box="[593,1081,1251,1275]" collectingDate="1931-03" collectionCode="DEIC" country="Madagascar" location="Morondava" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1">
|
||
<location id="C67BD5DCFFB6FF9DFDD8FB19399B2E9E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFDD8FB19399B2E9E" box="[593,736,1251,1275]" country="Madagascar" name="Morondava" pageId="5" pageNumber="4">Morondava</location>
|
||
,
|
||
<date id="B71AA5C7FFB6FF9DFD79FB19382F2E9E" box="[752,852,1251,1275]" pageId="5" pageNumber="4" value="1931-03">
|
||
<collectingDate id="A75E5C2FFFB6FF9DFD79FB19382F2E9E" box="[752,852,1251,1275]" pageId="5" pageNumber="4" value="1931-03">III.1931</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFCE3FB1938AC2E9E" box="[874,983,1251,1275]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFC6FFB193F4B2E9E" box="[998,1072,1251,1275]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFBC1FB1939672F7E" collectionCode="DEIC" country="Madagascar" location="Plateau de l'Andray - Reg. d'Ambovombe" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1">
|
||
<location id="C67BD5DCFFB6FF9DFBC1FB193A1D2F7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFBC1FB193A1D2F7E" country="Madagascar" name="Plateau de l'Andray - Reg. d'Ambovombe" pageId="5" pageNumber="4">Plateau de l’Andray – Reg. d’Ambovombe</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFEFFFAF93AB82F7E" box="[374,451,1283,1307]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFE44FAF9396F2F7E" box="[461,532,1283,1307]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFDACFAFE389C2F7E" box="[549,999,1283,1307]" collectingDate="1920-04-09" collectionCode="DEIC" country="Madagascar" location="Tamatave" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1">
|
||
<location id="C67BD5DCFFB6FF9DFDACFAFE39DB2F7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFDACFAFE39DB2F7E" box="[549,672,1284,1307]" country="Madagascar" name="Tamatave" pageId="5" pageNumber="4">Tamatave</location>
|
||
,
|
||
<date id="B71AA5C7FFB6FF9DFD20FAF938672F7E" box="[681,796,1283,1307]" pageId="5" pageNumber="4" value="1920-04-09">
|
||
<collectingDate id="A75E5C2FFFB6FF9DFD20FAF938672F7E" box="[681,796,1283,1307]" pageId="5" pageNumber="4" value="1920-04-09">9.IV.1920</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFCA2FAF938F42F7E" box="[811,911,1283,1307]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFC1EFAF938A52F7E" box="[919,990,1283,1307]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="73CC895AFFB6FF9DFC79FAF93AE32F5E" collectingDate="1919-12" collectionCode="DEIC, CMNH" country="Madagascar" location="Tananarive" pageId="5" pageNumber="4" specimenCount="3" specimenCount-male="3">
|
||
<location id="C67BD5DCFFB6FF9DFC79FAF93F012F7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FE393AE72F5E:C67BD5DCFFB6FF9DFC79FAF93F012F7E" box="[1008,1146,1283,1307]" country="Madagascar" name="Tananarive" pageId="5" pageNumber="4">Tananarive</location>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFB00FAF93FAC2F7E" box="[1161,1239,1283,1307]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFB69FAF93E5C2F7E" box="[1248,1319,1283,1307]" pageId="5" pageNumber="4">DEIC</collectionCode>
|
||
),
|
||
<date id="B71AA5C7FFB6FF9DFAB1FAF93EDB2F7E" box="[1336,1440,1283,1307]" pageId="5" pageNumber="4" value="1919-12">
|
||
<collectingDate id="A75E5C2FFFB6FF9DFAB1FAF93EDB2F7E" box="[1336,1440,1283,1307]" pageId="5" pageNumber="4" value="1919-12">XII.1919</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="D5A2488EFFB6FF9DFF40FAD93A532F5E" box="[201,296,1315,1339]" pageId="5" pageNumber="4" type="male">2 males</specimenCount>
|
||
,
|
||
<collectionCode id="A5B51BC2FFB6FF9DFEBAFAD93AEB2F5E" box="[307,400,1315,1339]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="5" pageNumber="4" type="Museum">CMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |