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<document id="757261BCE15D7260B159D0F00C69513F" ID-DOI="10.5281/zenodo.5161214" ID-ISSN="1942-1354" ID-Zenodo-Dep="5161214" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628116654208" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2011" docId="4B0D3211FFB6FF9DFF49FA98397B2D3E" docLanguage="en" docName="InsectaMundi.0191.1-13.pdf" docOrigin="Insecta Mundi 2011 (191)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Chaetodera perrieri" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="B7344A69FFB3FF98FF89FFFA3B7B2A65" masterDocTitle="Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1699035013485" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="BCF1295F53373A091920C61BD1620750">Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="A624AF7DA062C42D4FAB42E6C5B38DF0">Mawdsley, Jonathan R.</mods:namePart>
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<mods:date id="8D862C4245E712D7200FEB61A6888AF4">2011</mods:date>
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<mods:number id="E848E982854F13F98E016E4C37ECDF13">2011-09-02</mods:number>
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<mods:number id="4107D3DBD58993BCE974D6DB2F1DB724">191</mods:number>
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<mods:identifier id="3A683AD85CD2CE76B626447CBF71DE00" type="ISSN">1942-1354</mods:identifier>
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<treatment id="4B0D3211FFB6FF9DFF49FA98397B2D3E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E" httpUri="http://treatment.plazi.org/id/4B0D3211FFB6FF9DFF49FA98397B2D3E" lastPageNumber="4" pageId="5" pageNumber="4">
<subSubSection id="8BBED08CFFB6FF9DFF49FA9839C12F1E" box="[192,698,1378,1403]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="C31B8307FFB6FF9DFF49FA9839C12F1E" blockId="5.[192,698,1378,1435]" box="[192,698,1378,1403]" pageId="5" pageNumber="4">
<heading id="9853346BFFB6FF9DFF49FA9839C12F1E" bold="true" box="[192,698,1378,1403]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<taxonomicName id="04A4F884FFB6FF9DFF49FA9839C12F1E" ID-CoL="TBPH" authority="(Fairmaire 1897)" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[192,698,1378,1403]" class="Insecta" family="Carabidae" genus="Chaetodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="F1D05F15FFB6FF9DFF49FA9839C12F1E" bold="true" box="[192,698,1378,1403]" pageId="5" pageNumber="4">
<emphasis id="F1D05F15FFB6FF9DFF49FA983AAB2F1E" bold="true" box="[192,464,1378,1403]" italics="true" pageId="5" pageNumber="4">Chaetodera perrieri</emphasis>
(
<bibRefCitation id="A735FEF6FFB6FF9DFE56FA9839CA2F1E" author="Fairmaire, L." box="[479,689,1378,1403]" pageId="5" pageNumber="4" pagination="363 - 406" refId="ref9789" refString="Fairmaire, L. 1897. Materiaux pour la faune coleopterique de la region Malgache (4 e note). Annales de la Societe entomologique de Belgique. 41: 363 - 406." type="journal article" year="1897">Fairmaire 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFB6FF9DFF49FA793A052FFE" box="[192,382,1411,1435]" pageId="5" pageNumber="4" type="description">
<paragraph id="C31B8307FFB6FF9DFF49FA793A052FFE" blockId="5.[192,698,1378,1435]" box="[192,382,1411,1435]" pageId="5" pageNumber="4">
<figureCitation id="5B9F9F82FFB6FF9DFF49FA793A2F2FFE" box="[192,340,1411,1435]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="5" pageNumber="4">Figures 3, 4</figureCitation>
,
<figureCitation id="5B9F9F82FFB6FF9DFEE9FA793A052FFE" box="[352,382,1411,1435]" captionStart="Figures 16-17" captionStartId="7.[192,289,891,913]" captionTargetBox="[244,1359,193,856]" captionTargetId="figure-234@7.[192,1440,192,876]" captionTargetPageId="7" captionText="Figures 16-17. Distribution of Chaetodera species in Africa and Madagascar. 16) Distribution of C. antatsima (dark circles), C. perrieri (dark triangles), and C. blanchardi (dark squares). 17) Distribution of C. andriana (open circles), C. maheva (open stars), and C. regalis (dark stars)." figureDoi="http://doi.org/10.5281/zenodo.5161218" httpUri="https://zenodo.org/record/5161218/files/figure.png" pageId="5" pageNumber="4">16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFB6FF9DFF49FA3939D92FBE" box="[192,674,1475,1499]" pageId="5" pageNumber="4" type="reference_group">
<paragraph id="C31B8307FFB6FF9DFF49FA3939D92FBE" blockId="5.[192,674,1475,1499]" box="[192,674,1475,1499]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="E3B4A429FFB6FF9DFF49FA3939D92FBE" box="[192,674,1475,1499]" pageId="5" pageNumber="4">
<treatmentCitation id="4205A516FFB6FF9DFF49FA3939D92FBE" author="Fairmaire, L." box="[192,674,1475,1499]" page=")" pageId="5" pageNumber="4" year="1897">
<taxonomicName id="04A4F884FFB6FF9DFF49FA3939D92FBE" ID-CoL="V9ZK" authority="Fairmaire (1897: 364)" authorityName="Fairmaire" authorityPageNumber="364" authorityYear="1897" box="[192,674,1475,1499]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="F1D05F15FFB6FF9DFF49FA393AED2FBE" box="[192,406,1475,1499]" italics="true" pageId="5" pageNumber="4">Cicindela perrieri</emphasis>
<bibRefCitation id="A735FEF6FFB6FF9DFE12FA3939D92FBE" author="Fairmaire, L." box="[411,674,1475,1499]" pageId="5" pageNumber="4" pagination="363 - 406" refId="ref9789" refString="Fairmaire, L. 1897. Materiaux pour la faune coleopterique de la region Malgache (4 e note). Annales de la Societe entomologique de Belgique. 41: 363 - 406." type="journal article" year="1897">Fairmaire (1897: 364)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFB6FF9DFF49F9F93E152C5E" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="C31B8307FFB6FF9DFF49F9F93E152C5E" blockId="5.[192,1440,1539,1595]" pageId="5" pageNumber="4">
<emphasis id="F1D05F15FFB6FF9DFF49F9F93AFF2C7E" bold="true" box="[192,388,1539,1563]" pageId="5" pageNumber="4">Type material.</emphasis>
<typeStatus id="1C1F3DA5FFB6FF9DFE01F9F93A902C7E" box="[392,491,1539,1563]" pageId="5" pageNumber="4" type="syntype">Syntype</typeStatus>
male of
<taxonomicName id="04A4F884FFB6FF9DFDC2F9FE39BE2C7E" authorityName="Fairmaire" authorityYear="1897" box="[587,709,1540,1563]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="F1D05F15FFB6FF9DFDC2F9FE39BE2C7E" box="[587,709,1540,1563]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
</taxonomicName>
, labeled “Fairmaire/
<collectingCountry id="BBB3C397FFB6FF9DFC31F9F93F312C7E" box="[952,1098,1539,1563]" name="Madagascar" pageId="5" pageNumber="4">Madagascar</collectingCountry>
” (
<collectionCode id="A5B51BC2FFB6FF9DFBEBF9F93FD12C7E" box="[1122,1194,1539,1563]" pageId="5" pageNumber="4">DEIC</collectionCode>
, examined);
<typeStatus id="1C1F3DA5FFB6FF9DFAB4F9F93EDB2C7E" box="[1341,1440,1539,1563]" pageId="5" pageNumber="4" type="syntype">Syntype</typeStatus>
female of
<taxonomicName id="04A4F884FFB6FF9DFEBAF9DE3AD52C5E" authorityName="Fairmaire" authorityYear="1897" box="[307,430,1572,1595]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="F1D05F15FFB6FF9DFEBAF9DE3AD52C5E" box="[307,430,1572,1595]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
</taxonomicName>
labeled “
<taxonomicName id="04A4F884FFB6FF9DFD92F9D939EB2C5E" authorityName="Linnaeus" authorityYear="1758" box="[539,656,1571,1595]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Cicindela</taxonomicName>
/perrieri/Madag.” and “Fairmaire/Perrier” (
<collectionCode id="A5B51BC2FFB6FF9DFB1CF9D93FA52C5E" box="[1173,1246,1571,1595]" pageId="5" pageNumber="4">DEIC</collectionCode>
, examined).
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFB6FF9DFF49F9993F4B2CBE" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="C31B8307FFB6FF9DFF49F9993F4B2CBE" blockId="5.[192,1440,1635,1755]" pageId="5" pageNumber="4">
<emphasis id="F1D05F15FFB6FF9DFF49F9993A2A2C1E" bold="true" box="[192,337,1635,1659]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
Length 10.0-13.0 mm. Dorsal coloration as shown in
<figureCitation id="5B9F9F82FFB6FF9DFC64F9993FF82C1E" box="[1005,1155,1635,1659]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="5" pageNumber="4">Figures 3, 4</figureCitation>
. This species is distinguished by its obliquely banded ivory-white and black elytral coloration. The other Madagascan species of
<taxonomicName id="04A4F884FFB6FF9DFF49F9593A352CDE" authorityName="Jeannel. Characters" authorityYear="1946" box="[192,334,1699,1723]" class="Insecta" family="Carabidae" genus="Chaetodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="F1D05F15FFB6FF9DFF49F9593A352CDE" box="[192,334,1699,1723]" italics="true" pageId="5" pageNumber="4">Chaetodera</emphasis>
</taxonomicName>
with strongly oblique elytral markings is
<taxonomicName id="04A4F884FFB6FF9DFCD9F95E38A92CDE" baseAuthorityName="Kunckel d'Herculais" baseAuthorityYear="1887" box="[848,978,1699,1723]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="maheva">
<emphasis id="F1D05F15FFB6FF9DFCD9F95E38A92CDE" box="[848,978,1699,1723]" italics="true" pageId="5" pageNumber="4">C. maheva</emphasis>
</taxonomicName>
, which has blue or violet tibiae and is much larger (15.0-18.5 mm in length vs. 10.0-13.0 mm) than
<taxonomicName id="04A4F884FFB6FF9DFC23F93E3F532CBE" authorityName="Fairmaire" authorityYear="1897" box="[938,1064,1732,1755]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="F1D05F15FFB6FF9DFC23F93E3F532CBE" box="[938,1064,1732,1755]" italics="true" pageId="5" pageNumber="4">C. perrieri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFB6FF9DFF49F8F9397B2D3E" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="C31B8307FFB6FF9DFF49F8F9397B2D3E" blockId="5.[192,1440,1795,1883]" pageId="5" pageNumber="4">
<emphasis id="F1D05F15FFB6FF9DFF49F8F93ABE2D7E" bold="true" box="[192,453,1795,1819]" pageId="5" pageNumber="4">Material examined.</emphasis>
<materialsCitation id="73CC895AFFB6FF9DFE43F8F93F252D7E" box="[458,1118,1795,1819]" collectionCode="V, DEIC" location="Ambobaka" pageId="5" pageNumber="4" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
MADAGSCAR:
<location id="C67BD5DCFFB6FF9DFD0EF8F938702D7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E:C67BD5DCFFB6FF9DFD0EF8F938702D7E" box="[647,779,1795,1819]" name="Ambobaka" pageId="5" pageNumber="4">Ambobaka</location>
,
<collectionCode id="A5B51BC2FFB6FF9DFC9CF8FE38532D7E" box="[789,808,1796,1819]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
(
<specimenCount id="D5A2488EFFB6FF9DFCBCF8F938E82D7E" box="[821,915,1795,1819]" pageId="5" pageNumber="4" type="male">2 males</specimenCount>
,
<specimenCount id="D5A2488EFFB6FF9DFC14F8F93F7F2D7E" box="[925,1028,1795,1819]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFB6FF9DFB84F8F93F2E2D7E" box="[1037,1109,1795,1819]" pageId="5" pageNumber="4">DEIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFB6FF9DFBEEF8F93EE02D7E" box="[1127,1435,1795,1819]" collectionCode="DEIC" location="Analalava" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1">
<location id="C67BD5DCFFB6FF9DFBEEF8F93F992D7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E:C67BD5DCFFB6FF9DFBEEF8F93F992D7E" box="[1127,1250,1795,1819]" name="Analalava" pageId="5" pageNumber="4">Analalava</location>
(
<specimenCount id="D5A2488EFFB6FF9DFB7AF8F93E392D7E" box="[1267,1346,1795,1819]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFB6FF9DFAC2F8F93EE82D7E" box="[1355,1427,1795,1819]" pageId="5" pageNumber="4">DEIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFB6FF9DFF49F8D9393E2D5E" box="[192,581,1827,1851]" collectionCode="CMNH" location="Ile de Berafia" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1">
<location id="C67BD5DCFFB6FF9DFF49F8D93A1A2D5E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E:C67BD5DCFFB6FF9DFF49F8D93A1A2D5E" box="[192,353,1827,1851]" name="Ile de Berafia" pageId="5" pageNumber="4">Ile de Berafia</location>
(
<specimenCount id="D5A2488EFFB6FF9DFEFBF8D93AAC2D5E" box="[370,471,1827,1851]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFB6FF9DFE68F8D939472D5E" box="[481,572,1827,1851]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="5" pageNumber="4" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFB6FF9DFDC7F8D9389C2D5E" box="[590,999,1827,1851]" collectionCode="DEIC" location="Majunga" pageId="5" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location id="C67BD5DCFFB6FF9DFDC7F8D939C02D5E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E:C67BD5DCFFB6FF9DFDC7F8D939C02D5E" box="[590,699,1827,1851]" name="Majunga" pageId="5" pageNumber="4">Majunga</location>
(
<specimenCount id="D5A2488EFFB6FF9DFD42F8D938612D5E" box="[715,794,1827,1851]" pageId="5" pageNumber="4" type="male">1 male</specimenCount>
,
<specimenCount id="D5A2488EFFB6FF9DFCADF8D938F72D5E" box="[804,908,1827,1851]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
,
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<materialsCitation id="73CC895AFFB6FF9DFC78F8DE3EE02D5E" box="[1009,1435,1827,1851]" collectingDate="1929-01" collectionCode="DEIC" location="Mt. d'Ambre" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1">
<location id="C67BD5DCFFB6FF9DFC78F8DE3FF02D5E" LSID="urn:lsid:plazi:treatment:4B0D3211FFB6FF9DFF49FA98397B2D3E:C67BD5DCFFB6FF9DFC78F8DE3FF02D5E" box="[1009,1163,1827,1851]" name="Mt. d'Ambre" pageId="5" pageNumber="4">Mt. dAmbre</location>
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(
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<materialsCitation id="73CC895AFFB6FF9DFF49F8B93A872D3E" box="[192,508,1859,1883]" collectionCode="DEIC" location="Suberb Ile" pageId="5" pageNumber="4" specimenCount="1" specimenCount-male="1">
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)
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.
</paragraph>
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