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<document id="F828DDBF59ECF5178C55CF33D9023B4D" ID-CLB-Dataset="22365" ID-DOI="10.5852/ejt.2018.396" ID-GBIF-Dataset="2b95a851-e1bf-4bdd-9cce-441ddc4135d2" ID-Zenodo-Dep="3787049" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1588696404769" checkinUser="valdenar" docAuthor="Camfield, Rebecca &amp; Hughes, Mark" docDate="2018" docId="0397A96F2F63FFC603BFFB11E3BB2FBC" docLanguage="en" docName="ejt-396_camfield_hughes.pdf.imf" docOrigin="European Journal of Taxonomy 396" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia adscendens C. B. Clarke (Clarke 1890" docType="treatment" docVersion="11" lastPageNumber="18" masterDocId="FFAED1172F6DFFD7025FFFB1E00A2E5C" masterDocTitle="A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India" masterLastPageNumber="116" masterPageNumber="1" pageNumber="15" updateTime="1737107973296" updateUser="diego">
<mods:mods id="8E860E873D783068783241DB8D59844C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C69DE01ABA9F83DEBF7194851F2991EC">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
</mods:titleInfo>
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<mods:namePart id="9BBC22B873D9117119E699BF2B07805B">Camfield, Rebecca</mods:namePart>
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<mods:namePart id="59C46432ED74715307810E874C8029D9">Hughes, Mark</mods:namePart>
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<mods:title id="5519C41619A4D6B53D5F4B98FDDA0089">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="101862BAE8BD985BB63B90AF753A2B33">2018</mods:date>
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<mods:number id="ED9BE7AADC04119AE3E87C9661AB6A97">2018-01-19</mods:number>
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<mods:detail id="4475FB847BF1E19D21802CF1E6D5FFD2" type="volume">
<mods:number id="7AEBEE63B2DF997576DE529EEF606B9F">396</mods:number>
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<mods:identifier id="947176DACC5CA22031B80B374C8B3D53" type="CLB-Dataset">22365</mods:identifier>
<mods:identifier id="8BAB28DEA3380D88E06F1C51EBE54911" type="DOI">10.5852/ejt.2018.396</mods:identifier>
<mods:identifier id="755F40F6CC5C9E41B95DEB5642E59D7B" type="GBIF-Dataset">2b95a851-e1bf-4bdd-9cce-441ddc4135d2</mods:identifier>
<mods:identifier id="2AA3375DF9334DFAD2C51BDD89921127" type="Zenodo-Dep">3787049</mods:identifier>
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<treatment id="0397A96F2F63FFC603BFFB11E3BB2FBC" ID-DOI="http://doi.org/10.5281/zenodo.3794315" ID-GBIF-Taxon="163963846" ID-Zenodo-Dep="3794315" LSID="urn:lsid:plazi:treatment:0397A96F2F63FFC603BFFB11E3BB2FBC" httpUri="http://treatment.plazi.org/id/0397A96F2F63FFC603BFFB11E3BB2FBC" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" scope_class="Equisetopsida" scope_family="Begoniaceae" scope_order="Cucurbitales" scope_subClass="Magnoliidae" scope_superOrder="Rosanae">
<paragraph id="8B8118792F63FFD903BFFB11E45E2AE7" blockId="14.[480,1108,1184,1246]" box="[480,1108,1184,1211]" pageId="14" pageNumber="15">
<heading id="D0C9AF152F63FFD903BFFB11E45E2AE7" box="[480,1108,1184,1211]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C3E63FA2F63FFD903BFFB11E3682AE6" ID-CoL="7SDMF" authority="C. B. Clarke" authorityName="C. B. Clarke (Clarke" authorityYear="1890" box="[480,866,1184,1211]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F63FFD903BFFB11E2DD2AE7" bold="true" box="[480,727,1184,1211]" italics="true" pageId="14" pageNumber="15">Begonia adscendens</emphasis>
C.B.Clarke
</taxonomicName>
[sect.
<taxonomicName id="4C3E63FA2F63FFD901F0FB11E4402AE6" box="[943,1098,1184,1210]" class="Magnoliopsida" family="Begoniaceae" genus="Diploclinium" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F63FFD901F0FB11E4402AE6" box="[943,1098,1184,1210]" italics="true" pageId="14" pageNumber="15">Diploclinium</emphasis>
</taxonomicName>
]
</heading>
</paragraph>
<paragraph id="8B8118792F63FFD900BBFB72E35A2A81" blockId="14.[480,1108,1184,1246]" box="[740,848,1219,1246]" pageId="14" pageNumber="15">
<figureCitation id="130504FC2F63FFD900BBFB72E35A2A81" box="[740,848,1219,1246]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1991,2017]" captionStartId-1="16.[189,232,1957,1983]" captionTargetBox-0="[425,1160,1217,1950]" captionTargetBox-1="[295,1292,262,1918]" captionTargetId-0="figure@15.[416,1172,1209,1962]" captionTargetId-1="figure@16.[295,1292,262,1918]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 8. Map showing the location of B. adscendens C.B.Clarke specimens." captionText-1="Fig. 9. Illustration of B. adscendens C.B.Clarke by J. Allen (Clarke 1890). a. Dissected fruit capsule. b. Fruit cross-section diagram. Image courtesy of RBGE library and Lynsey Wilson." figureDoi-0="http://doi.org/10.5281/zenodo.3787063" figureDoi-1="http://doi.org/10.5281/zenodo.3787065" httpUri-0="https://zenodo.org/record/3787063/files/figure.png" httpUri-1="https://zenodo.org/record/3787065/files/figure.png" pageId="14" pageNumber="15">Figs 8–9</figureCitation>
</paragraph>
<paragraph id="8B8118792F63FFD902E2FAB4E3A02B43" blockId="14.[189,1399,1285,1347]" box="[189,938,1285,1312]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2E3F572F63FFD902E2FAB4E3A02B43" box="[189,938,1285,1312]" pageId="14" pageNumber="15">
<emphasis id="B94AC46B2F63FFD902E2FAB4E29D2B43" box="[189,663,1285,1311]" italics="true" pageId="14" pageNumber="15">Journal of the Linnean Society, Botany</emphasis>
25: 26
<treatmentCitation id="0A9F3E682F63FFD900A4FAB4E3AD2B43" box="[763,935,1285,1311]" pageId="14" pageNumber="15">
(
<bibRefCitation id="EFAF65882F63FFD9015CFAB4E3972B43" author="Clarke C. B." box="[771,925,1285,1311]" pageId="14" pageNumber="15" pagination="1 - 107" refId="ref53104" refString="Clarke C. B. 1890. On the plants of Kohima and Muneypore. Journal of the Linnean Society, Botany 25: 1 - 107. Available from http: // biodiversitylibrary. org / page / 180617 [accessed May 2009]." type="journal article" year="1890">Clarke 1890</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8118792F63FFD901EAFAB4E5222B1F" blockId="14.[189,1399,1285,1347]" pageId="14" pageNumber="15">
–
<materialsCitation id="3B5612242F63FFD90191FAB4E43B2B1E" collectingDate="1885-10-25" collectionCode="K" collectorName="Clarke" country="India" location="Jakpho" municipality="Naga Hills" pageId="14" pageNumber="15" specimenCode="K000634623" specimenCount="1" stateProvince="Nagaland" typeStatus="lectotype">
<typeStatus id="5485A6DB2F63FFD90191FAB4E4072B43" box="[974,1037,1285,1311]" pageId="14" pageNumber="15">Type</typeStatus>
:
<collectingCountry id="F32958E92F63FFD90643FAB4E4512B43" box="[1052,1115,1285,1311]" name="India" pageId="14" pageNumber="15">India</collectingCountry>
,
<collectingRegion id="49FAD69B2F63FFD90635FAB4E4D72B43" box="[1130,1245,1285,1311]" country="India" name="Nagaland" pageId="14" pageNumber="15">Nagaland</collectingRegion>
,
<collectingMunicipality id="6BE582032F63FFD906B2FAB4E5782B43" box="[1261,1394,1285,1311]" pageId="14" pageNumber="15">Naga Hills</collectingMunicipality>
,
<location id="8EE14EA22F63FFD902E2FA99E1192B1E" LSID="urn:lsid:plazi:treatment:0397A96F2F63FFC603BFFB11E3BB2FBC:8EE14EA22F63FFD902E2FA99E1192B1E" box="[189,275,1320,1346]" country="India" municipality="Naga Hills" name="Jakpho" pageId="14" pageNumber="15" stateProvince="Nagaland">Jakpho</location>
,
<date id="FF803EB92F63FFD90340FA99E1B32B1F" box="[287,441,1320,1347]" pageId="14" pageNumber="15" value="1885-10-25">
<collectingDate id="EFC4C7512F63FFD90340FA99E1B32B1F" box="[287,441,1320,1347]" pageId="14" pageNumber="15" value="1885-10-25">25 Oct. 1885</collectingDate>
</date>
,
<emphasis id="B94AC46B2F63FFD90399FA99E26C2B1E" box="[454,614,1320,1346]" italics="true" pageId="14" pageNumber="15">
<collectorName id="26CB7DAF2F63FFD90399FA99E21C2B1E" box="[454,534,1320,1346]" pageId="14" pageNumber="15">Clarke</collectorName>
41240
</emphasis>
(lecto-:
<specimenCode id="DB98B0022F63FFD90095FA99E3682B1E" box="[714,866,1320,1346]" pageId="14" pageNumber="15">K000634623</specimenCode>
, here designated;
</materialsCitation>
<materialsCitation id="3B5612242F63FFD90667FA99E52E2B1F" box="[1080,1316,1320,1347]" collectingDate="1885-10-25" collectionCode="K" collectorName="Clarke" country="India" location="Jakpho" municipality="Naga Hills" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Nagaland" typeStatus="isolectotype">isolecto-: CAL n.v.)</materialsCitation>
.
</paragraph>
<paragraph id="8B8118792F63FFD902E2FADBE4FD2BD8" blockId="14.[189,1271,1385,1412]" box="[189,1271,1385,1412]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2E3F572F63FFD902E2FADBE4FD2BD8" box="[189,1271,1385,1412]" pageId="14" pageNumber="15">
<emphasis id="B94AC46B2F63FFD902E2FADBE1FD2BDF" box="[189,503,1385,1411]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3E63FA2F63FFD902E2FADBE1CA2BDF" box="[189,448,1385,1411]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="subSpecies" species="obversa" subSpecies="auct">Begonia obversa auct</taxonomicName>
. non
</emphasis>
C.B.Clarke: Chauhan in
<emphasis id="B94AC46B2F63FFD9017FFAD8E3E52BDF" box="[800,1007,1385,1412]" italics="true" pageId="14" pageNumber="15">
Flora of
<collectingRegion id="49FAD69B2F63FFD901D7FADBE3E52BDF" box="[904,1007,1386,1411]" country="India" name="Manipur" pageId="14" pageNumber="15">Manipur</collectingRegion>
</emphasis>
: 425
<treatmentCitation id="0A9F3E682F63FFD9066DFAD8E4F92BD8" box="[1074,1267,1385,1412]" pageId="14" pageNumber="15">
(
<bibRefCitation id="EFAF65882F63FFD90665FAD8E4E12BD8" author="Chauhan A. S." box="[1082,1259,1385,1412]" pageId="14" pageNumber="15" pagination="424 - 429" refId="ref52957" refString="Chauhan A. S. 2000. Begoniaceae. In: Singh N. P, Chauhan A. S. &amp; Mondal M. S. (eds) Flora of Manipur: 424 - 429. Botanical Survey of India, Calcutta." type="book chapter" year="2000">Chauhan 2000</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8118792F63FFD902E2FA1AE2332B99" blockId="14.[189,569,1451,1477]" box="[189,569,1451,1477]" pageId="14" pageNumber="15">
<heading id="D0C9AF152F63FFD902E2FA1AE2332B99" bold="true" box="[189,569,1451,1477]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B94AC46B2F63FFD902E2FA1AE2332B99" bold="true" box="[189,569,1451,1477]" pageId="14" pageNumber="15">Citations in other publications</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F63FFD902E2FA6BE47C2BA8" blockId="14.[189,1142,1497,1524]" box="[189,1142,1497,1524]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB2E3F572F63FFD902E2FA6BE47C2BA8" box="[189,1142,1497,1524]" pageId="14" pageNumber="15">
As
<taxonomicName id="4C3E63FA2F63FFD902BAFA6BE1862BAF" ID-CoL="7SDMF" authority=": Uddin (2007: 591)" authorityName=": Uddin" authorityPageNumber="591" authorityYear="2007" box="[229,396,1497,1523]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F63FFD902BAFA6BE1862BAF" box="[229,396,1497,1523]" italics="true" pageId="14" pageNumber="15">B. adscendens</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9F3E682F63FFD903C3FA68E2732BA8" author="Uddin A." box="[412,633,1497,1524]" page=")" pageId="14" pageNumber="15" year="2007">
<bibRefCitation id="EFAF65882F63FFD903C3FA68E2732BA8" author="Uddin A." box="[412,633,1497,1524]" pageId="14" pageNumber="15" pagination="591 - 597" refId="ref57753" refString="Uddin A. 2007. Distribution and status of India Begonia species. Journal of Economic and Taxonomic Botany 31 (3): 591 - 597." type="journal article" year="2007">Uddin (2007: 591)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F63FFD900DAFA68E35E2BA8" author="Hughes M." box="[645,852,1497,1524]" page="3" pageId="14" pageNumber="15" year="2008">
<bibRefCitation id="EFAF65882F63FFD900DAFA68E35E2BA8" author="Hughes M." box="[645,852,1497,1524]" pageId="14" pageNumber="15" refId="ref54915" refString="Hughes M. 2008. An annotated checklist of Southeast Asian Begonia. Royal Botanic Garden Edinburgh." type="book" year="2008">Hughes (2008: 3)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F63FFD9013EFA68E4782BA8" author="Uddin A. &amp; Gogoi R. &amp; Mao A. A. &amp; Phukan S." box="[865,1138,1497,1524]" page="53" pageId="14" pageNumber="15" year="2008">
<bibRefCitation id="EFAF65882F63FFD9013EFA68E4782BA8" author="Uddin A. &amp; Gogoi R. &amp; Mao A. A. &amp; Phukan S." box="[865,1138,1497,1524]" pageId="14" pageNumber="15" pagination="53 - 55" refId="ref57781" refString="Uddin A., Gogoi R., Mao A. A. &amp; Phukan S. 2008. Rediscovery of Begonia adscendens C. B. Clarke. Rheedea 18 (1): 53 - 55." type="journal article" year="2008">
Uddin
<emphasis id="B94AC46B2F63FFD901EDFA6BE3E72BAF" box="[946,1005,1497,1523]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(2008: 53)
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8118792F63FFD902E2F9AAE1732869" blockId="14.[189,377,1563,1589]" box="[189,377,1563,1589]" pageId="14" pageNumber="15">
<heading id="D0C9AF152F63FFD902E2F9AAE1732869" bold="true" box="[189,377,1563,1589]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B94AC46B2F63FFD902E2F9AAE1732869" bold="true" box="[189,377,1563,1589]" pageId="14" pageNumber="15">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F63FFD902E2F9FBE24628DB" blockId="14.[189,1399,1609,1671]" pageId="14" pageNumber="15">
<materialsCitation id="3B5612242F63FFD902E2F9FBE4682838" ID-GBIF-Occurrence="2610421308" box="[189,1122,1609,1636]" pageId="14" pageNumber="15">
<collectingCountry id="F32958E92F63FFD902E2F9FBE11F2838" box="[189,277,1610,1636]" name="India" pageId="14" pageNumber="15">INDIA</collectingCountry>
:
<emphasis id="B94AC46B2F63FFD90340F9F8E185283F" bold="true" box="[287,399,1609,1635]" pageId="14" pageNumber="15">
<collectingRegion id="49FAD69B2F63FFD90340F9F8E185283F" box="[287,399,1609,1635]" country="India" name="Manipur" pageId="14" pageNumber="15">Manipur</collectingRegion>
</emphasis>
: Esii Hill,
<date id="FF803EB92F63FFD90050F9F8E297283F" box="[527,669,1609,1635]" pageId="14" pageNumber="15" value="2006-09-01">1 Sep. 2006</date>
,
<emphasis id="B94AC46B2F63FFD900F7F9FBE3A12838" box="[680,939,1610,1636]" italics="true" pageId="14" pageNumber="15">Mao &amp; Gogoi 109408</emphasis>
(
<collectingRegion id="49FAD69B2F63FFD901E6F9F8E42A283F" box="[953,1056,1609,1635]" country="India" name="Assam" pageId="14" pageNumber="15">ASSAM</collectingRegion>
n.v.).
</materialsCitation>
<materialsCitation id="3B5612242F63FFD90635F9FBE24628DB" ID-GBIF-Occurrence="2610421308" pageId="14" pageNumber="15">
<emphasis id="B94AC46B2F63FFD90635F9FBE4E92838" bold="true" box="[1130,1251,1610,1636]" pageId="14" pageNumber="15">
<collectingRegion id="49FAD69B2F63FFD90635F9FBE4E92838" box="[1130,1251,1610,1636]" country="India" name="Nagaland" pageId="14" pageNumber="15">Nagaland</collectingRegion>
</emphasis>
: Naga Hills, Jakpho,
<emphasis id="B94AC46B2F63FFD90340F9DCE1B528DB" box="[287,447,1645,1671]" italics="true" pageId="14" pageNumber="15">Clarke 41247</emphasis>
(
<collectionCode id="ED2F80BC2F63FFD90391F9DCE20028DB" box="[462,522,1645,1671]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="14" pageNumber="15" type="Herbarium">CAL</collectionCode>
n.v.).
</materialsCitation>
</paragraph>
<paragraph id="8B8118792F63FFD902E2F91FE1442894" blockId="14.[189,334,1710,1736]" box="[189,334,1710,1736]" pageId="14" pageNumber="15">
<heading id="D0C9AF152F63FFD902E2F91FE1442894" bold="true" box="[189,334,1710,1736]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B94AC46B2F63FFD902E2F91FE1442894" bold="true" box="[189,334,1710,1736]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F63FFD802E2F96DE2292CDA" blockId="14.[189,1399,1756,1994]" lastBlockId="15.[189,1399,267,646]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Tuberous, monoecious herb,
<quantity id="4CC6B59C2F63FFD9004FF96CE28028AB" box="[528,650,1756,1783]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="14" pageNumber="15" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 cm</quantity>
high. Stipules: ovate, 2 ×
<quantity id="4CC6B59C2F63FFD901E3F96CE42928AB" box="[956,1059,1757,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="14" pageNumber="15" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
, glabrous, caduceus. Leaves: petiole
<quantity id="4CC6B59C2F63FFD90349F8B1E18A2946" box="[278,384,1792,1819]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="14" pageNumber="15" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
long, glabrous; lamina ovate to broadly ovate, basifixed, base cordate with lobes not overlapping, 4–16 ×
<quantity id="4CC6B59C2F63FFD903EDF892E22A2961" box="[434,544,1827,1853]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="2.2" metricValueMin="0.2" pageId="14" pageNumber="15" unit="cm" value="12.0" valueMax="22.0" valueMin="2.0">2–22 cm</quantity>
, slightly asymmetric, upper surface green, glabrous, underside pale green or pale green with red markings, puberulous on veins mostly, venation palmate; midrib
<quantity id="4CC6B59C2F63FFD906ABF8F7E57D293D" box="[1268,1399,1862,1889]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="13.0" metricValueMin="3.5" pageId="14" pageNumber="15" unit="cm" value="8.25" valueMax="13.0" valueMin="3.5">3.5–13 cm</quantity>
long; margin toothed at the end of the veins with smaller teeth between, with sparse hairs; apex shortly acuminate. Inflorescence: cymose, terminal, few; peduncle glabrous, branching 1–2 times, primary
<quantity id="4CC6B59C2F63FFD902E2F801E13F2996" box="[189,309,1968,1994]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="14" pageNumber="15" unit="cm" value="14.0" valueMax="18.0" valueMin="10.0">10–18 cm</quantity>
long, secondary
<quantity id="4CC6B59C2F63FFD903A2F801E25D2996" box="[509,599,1968,1994]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="14" pageNumber="15" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 cm</quantity>
long, with 2–
<specimenCount id="9D38D3F02F63FFD900A1F801E36F2996" box="[766,869,1968,1994]" pageId="14" pageNumber="15" type="female">4 female</specimenCount>
and 2–
<specimenCount id="9D38D3F02F63FFD9019FF801E41A2996" box="[960,1040,1968,1994]" pageId="14" pageNumber="15" type="male">6 male</specimenCount>
flowers; bracts not seen. Male flower: pedicel
<quantity id="4CC6B59C2F62FFD80330FEBAE1EA2F7A" box="[367,480,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="15" pageNumber="16" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
long, glabrous; tepals 4; outer tepals orbicular to ovate, 10–18 × (5–)
<quantity id="4CC6B59C2F62FFD806A3FEBAE5782F7A" box="[1276,1394,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="15" pageNumber="16" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">9–12 mm</quantity>
, white to pink, sparsely pale puberulous on reverse, near the base, margin entire; inner tepals narrowly elliptic, 5–7 ×
<quantity id="4CC6B59C2F62FFD80339FEE3E1C12F30" box="[358,459,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="15" pageNumber="16" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
, white to pink, glabrous; androecium with 16–20 stamens, asymmetric; filaments
<quantity id="4CC6B59C2F62FFD802E2FEC4E13F2FD3" box="[189,309,373,400]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="15" pageNumber="16" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, unequal, fused at base into a short column; anther oblong elliptic,
<quantity id="4CC6B59C2F62FFD806D0FEC4E4D92FD3" box="[1167,1235,373,399]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
long, slightly hooded, connective not extended. Female flower: pedicel
<quantity id="4CC6B59C2F62FFD80133FE29E3EB2FEF" box="[876,993,408,435]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="15" pageNumber="16" unit="mm" value="13.0" valueMax="18.0" valueMin="8.0">8–18 mm</quantity>
long, glabrous; bracteoles absent; tepals 4–5, unequal, ovate, outer tepals, 8–15 ×
<quantity id="4CC6B59C2F62FFD80150FE0AE3702F8A" box="[783,890,443,470]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="15" pageNumber="16" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
, white tinted pink, sparsely puberulent on reverse, margin entire, inner tepals as in male; ovary 3-locular, placentae bifid; capsule globose-ellipsoid, glabrous, with one long triangular wing and two short triangular wings; styles 3, convoluted with slightly twisted ends, persistent. Fruit pendulous; capsule globose-ellipsoid 6–10 ×
<quantity id="4CC6B59C2F62FFD8061EFD94E4A72C63" box="[1089,1197,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="15" pageNumber="16" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, glabrous; wings extending along the pedicel slightly, unequal; longest wing rounded oblong, 26–28 ×
<quantity id="4CC6B59C2F62FFD806F6FDF9E5052C3F" box="[1193,1295,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="15" pageNumber="16" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
; shortest wings triangular, 3 ×
<quantity id="4CC6B59C2F62FFD803E7FDDAE2152CDA" box="[440,543,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="15" pageNumber="16" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
.
</paragraph>
<paragraph id="8B8118792F62FFD802E2FD1DE21F2C9B" blockId="15.[189,533,684,711]" box="[189,533,684,711]" pageId="15" pageNumber="16">
<heading id="D0C9AF152F62FFD802E2FD1DE21F2C9B" bold="true" box="[189,533,684,711]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B94AC46B2F62FFD802E2FD1DE21F2C9B" bold="true" box="[189,533,684,711]" pageId="15" pageNumber="16">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F62FFD802E2FD6AE1832D44" blockId="15.[189,1398,731,792]" pageId="15" pageNumber="16">
Manipur and Nagaland; also in
<collectingCountry id="F32958E92F62FFD8006BFD6DE2AC2CAA" box="[564,678,732,758]" name="Myanmar" pageId="15" pageNumber="16">Myanmar</collectingCountry>
and
<collectingCountry id="F32958E92F62FFD800BEFD6DE3142CAA" box="[737,798,732,758]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
;
<quantity id="4CC6B59C2F62FFD80175FD6AE3DC2CAA" box="[810,982,731,758]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="15" pageNumber="16" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">2400–2600 m</quantity>
. Flowering: September to October; fruiting: October.
</paragraph>
<paragraph id="8B8118792F62FFD802E2FCF1E1BF2D06" blockId="15.[189,437,832,858]" box="[189,437,832,858]" pageId="15" pageNumber="16">
<heading id="D0C9AF152F62FFD802E2FCF1E1BF2D06" bold="true" box="[189,437,832,858]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B94AC46B2F62FFD802E2FCF1E1BF2D06" bold="true" box="[189,437,832,858]" pageId="15" pageNumber="16">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F62FFD802E2FCDEE3B32A49" blockId="15.[189,1399,877,1046]" pageId="15" pageNumber="16">
Least Concern. Current data for
<taxonomicName id="4C3E63FA2F62FFD80061FCDEE2ED2DD4" authorityName=": Uddin" authorityYear="2007" box="[574,743,878,904]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F62FFD80061FCDEE2ED2DD4" box="[574,743,878,904]" italics="true" pageId="15" pageNumber="16">B. adscendens</emphasis>
</taxonomicName>
gives an EOO of
<quantity id="4CC6B59C2F62FFD8019FFCDFE4412DD4" box="[960,1099,878,905]" metricMagnitude="8" metricUnit="m" metricValue="1.13895" pageId="15" pageNumber="16" unit="km" value="113895.0">113,895 km</quantity>
<superScript id="7C4BB5312F62FFD80613FCDCE45F2D27" attach="left" box="[1100,1109,877,891]" fontSize="6" pageId="15" pageNumber="16">2</superScript>
and an AOO of
<quantity id="4CC6B59C2F62FFD80744FCDFE56C2DD4" box="[1307,1382,878,904]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="16" unit="km" value="32.0">32 km</quantity>
<superScript id="7C4BB5312F62FFD80738FCDCE57A2D27" attach="right" box="[1383,1392,877,891]" fontSize="6" pageId="15" pageNumber="16">2</superScript>
. This sample size is low, but the specimens are widespread. The data would suggest this species is vulnerable but due to the distribution and supply of suitable habitat this species is classed as least concern. This species is found throughout the
<collectingRegion id="49FAD69B2F62FFD800B6FC69E3352DAE" box="[745,831,984,1010]" country="Myanmar" name="Rakhine" pageId="15" pageNumber="16">Arakan</collectingRegion>
mountains and into the mountains of Northern
<collectingCountry id="F32958E92F62FFD802E2FC4DE13B2A4A" box="[189,305,1020,1046]" name="Myanmar" pageId="15" pageNumber="16">Myanmar</collectingCountry>
, both of which have extensive areas of suitable habitat.
</paragraph>
<caption id="DF4148F12F62FFD802E2F876E42D29BD" ID-DOI="http://doi.org/10.5281/zenodo.3787063" ID-Zenodo-Dep="3787063" box="[189,1063,1991,2017]" httpUri="https://zenodo.org/record/3787063/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1991,2017]" targetBox="[425,1160,1217,1950]" targetPageId="15">
<paragraph id="8B8118792F62FFD802E2F876E42D29BD" blockId="15.[189,1063,1991,2017]" box="[189,1063,1991,2017]" pageId="15" pageNumber="16">
<emphasis id="B94AC46B2F62FFD802E2F876E10029BD" bold="true" box="[189,266,1991,2017]" pageId="15" pageNumber="16">Fig. 8.</emphasis>
Map showing the location of
<taxonomicName id="4C3E63FA2F62FFD80035F879E39429BD" authority="C. B. Clarke" authorityName="C. B. Clarke (Clarke" authorityYear="1890" box="[618,926,1991,2017]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F62FFD80035F879E31829BD" box="[618,786,1991,2017]" italics="true" pageId="15" pageNumber="16">B. adscendens</emphasis>
C.B.Clarke
</taxonomicName>
specimens.
</paragraph>
</caption>
<caption id="DF4148F12F7DFFC702E2F814E49329BE" ID-DOI="http://doi.org/10.5281/zenodo.3787065" ID-Zenodo-Dep="3787065" httpUri="https://zenodo.org/record/3787065/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1957,1983]" targetBox="[295,1292,262,1918]" targetPageId="16">
<paragraph id="8B8118792F7DFFC702E2F814E49329BE" blockId="16.[189,1399,1957,2019]" pageId="16" pageNumber="17">
<emphasis id="B94AC46B2F7DFFC702E2F814E10629E3" bold="true" box="[189,268,1957,1983]" pageId="16" pageNumber="17">Fig. 9.</emphasis>
Illustration of
<taxonomicName id="4C3E63FA2F7DFFC7039EF817E2F229E3" authority="C. B. Clarke" authorityName="C. B. Clarke (Clarke" authorityYear="1890" box="[449,760,1957,1983]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F7DFFC7039EF817E26129E3" box="[449,619,1957,1983]" italics="true" pageId="16" pageNumber="17">B. adscendens</emphasis>
C.B.Clarke
</taxonomicName>
by J. Allen (
<bibRefCitation id="EFAF65882F7DFFC701CBF814E42729E3" author="Clarke C. B." box="[916,1069,1957,1983]" pageId="16" pageNumber="17" pagination="1 - 107" refId="ref53104" refString="Clarke C. B. 1890. On the plants of Kohima and Muneypore. Journal of the Linnean Society, Botany 25: 1 - 107. Available from http: // biodiversitylibrary. org / page / 180617 [accessed May 2009]." type="journal article" year="1890">Clarke 1890</bibRefCitation>
). a. Dissected fruit capsule. b. Fruit cross-section diagram. Image courtesy of RBGE library and Lynsey Wilson.
</paragraph>
</caption>
<paragraph id="8B8118792F7CFFC602E2FEBAE1252F79" blockId="17.[189,303,267,293]" box="[189,303,267,293]" pageId="17" pageNumber="18">
<heading id="D0C9AF152F7CFFC602E2FEBAE1252F79" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B94AC46B2F7CFFC602E2FEBAE1252F79" bold="true" box="[189,303,267,293]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F7CFFC602E2FE88E3BB2FBC" blockId="17.[189,1399,313,481]" pageId="17" pageNumber="18">
This species is similar to
<taxonomicName id="4C3E63FA2F7CFFC603B4FE8BE2652F0F" authorityName="H. Lev. (Leveille" authorityYear="1904" box="[491,623,313,339]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="labordei">
<emphasis id="B94AC46B2F7CFFC603B4FE8BE2652F0F" box="[491,623,313,339]" italics="true" pageId="17" pageNumber="18">B. labordei</emphasis>
</taxonomicName>
which differs in having hairy leaves and dark purple hairs on the exterior of the male flowers that are visible without a microscope; the hairs are pale and near the base of the tepals in
<taxonomicName id="4C3E63FA2F7CFFC6032BFE30E2162FC6" authorityName="C. B. Clarke (Clarke" authorityYear="1890" box="[372,540,384,410]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F7CFFC6032BFE30E2162FC6" box="[372,540,384,410]" italics="true" pageId="17" pageNumber="18">B. adscendens</emphasis>
</taxonomicName>
. The two species also differ in fruit wing shape; the longest wing curves adaxially on
<taxonomicName id="4C3E63FA2F7CFFC60304FE15E1EB2FE1" authorityName="H. Lev. (Leveille" authorityYear="1904" box="[347,481,419,445]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="labordei">
<emphasis id="B94AC46B2F7CFFC60304FE15E1EB2FE1" box="[347,481,419,445]" italics="true" pageId="17" pageNumber="18">B. labordei</emphasis>
</taxonomicName>
but on
<taxonomicName id="4C3E63FA2F7CFFC6001BFE15E2E52FE1" authorityName="C. B. Clarke (Clarke" authorityYear="1890" box="[580,751,419,445]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="adscendens">
<emphasis id="B94AC46B2F7CFFC6001BFE15E2E52FE1" box="[580,751,419,445]" italics="true" pageId="17" pageNumber="18">B. adscendens</emphasis>
</taxonomicName>
it reaches up past the apex of the fruit capsule. This species is one of the highest altitude
<taxonomicName id="4C3E63FA2F7CFFC60033FE76E2C42FBC" box="[620,718,455,480]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F7CFFC60033FE76E2C42FBC" box="[620,718,455,480]" italics="true" pageId="17" pageNumber="18">Begonia</emphasis>
</taxonomicName>
in Northeast
<collectingCountry id="F32958E92F7CFFC60132FE77E3A72FBC" box="[877,941,454,480]" name="India" pageId="17" pageNumber="18">India</collectingCountry>
.
</paragraph>
</treatment>
</document>
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