196 lines
33 KiB
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196 lines
33 KiB
XML
<document id="CD1EF2C19DAA98CA1907A804E4A35358" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. & Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFB3076A54C1FC7E7CC9F80D" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Holthuija Ng & Forges 2015, n. gen." docType="key" docUuid="8DB52511-0A93-47A6-9382-6DFAF32A962C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="160" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="160" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F052B557F2758F7741C311ECC9A3F1B2">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="A8D7B3A11501CE45D079F8D4326EED07">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="40E525AFD02F393F5895676965EF67BB">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="0D73CE058D881E1A42DBB44D5A285D40">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="8DDF7FE304F13A00438DA5FA9D3B0F07">2015</mods:date>
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<mods:number id="47673D243838A73055A2CACF8C63F58A">63</mods:number>
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<heading id="D0F76331FFB3076A54C1FC7E7C50FBB6" bold="true" box="[934,1330,1021,1045]" centered="true" fontSize="10" level="2" pageId="50" pageNumber="160" reason="2">
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<emphasis id="B974084FFFB3076A54C1FC7E7C50FBB6" bold="true" box="[934,1330,1021,1045]" pageId="50" pageNumber="160">
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Key to species of
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<taxonomicName id="4C00AFDEFFB3076A5313FC7E7DBDFBB6" ID-CoL="7PKL9" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1140,1247,1021,1045]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B974084FFFB3076A5313FC7E7DBDFBB6" bold="true" box="[1140,1247,1021,1045]" italics="true" pageId="50" pageNumber="160">Holthuija</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B534FFB3076A5382FC7E7C50FBB6" box="[1253,1330,1021,1045]" pageId="50" pageNumber="160" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A87D6FFB3076A544AFBBA7CC9F80D" pageId="50" pageNumber="160" type="key">
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<keyStep id="30F4CCF8FFB3076A544AFBBA7CC8FBC8" pageId="50" pageNumber="160">
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<paragraph id="8BBFD45DFFB3076A544AFBBA7CC8FBED" blockId="50.[813,1450,1081,1131]" box="[813,1450,1081,1103]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFBBA7CC8FBED" box="[813,1450,1081,1103]" pageId="50" pageNumber="160">
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1. Carapace with 2 median gastric spines (e.g.,
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<figureCitation id="133BC8D8FFB3076A526BFBBA7C07FBEC" box="[1292,1381,1081,1103]" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Fig. 26A</figureCitation>
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)........2
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</keyLead>
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</paragraph>
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<paragraph id="8BBFD45DFFB3076A544AFBD57CC8FBC8" blockId="50.[813,1450,1081,1131]" box="[813,1450,1109,1131]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFBD57CC8FBC8" box="[813,1450,1109,1131]" pageId="50" pageNumber="160">
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– Carapace with single median gastric spine (e.g.,
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<figureCitation id="133BC8D8FFB3076A5257FBD67CE5FBC8" box="[1328,1415,1109,1131]" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Fig. 26C</figureCitation>
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) ..3
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep id="30F4CCF8FFB3076A544AFB0C7CC9FA96" pageId="50" pageNumber="160">
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||
<paragraph id="8BBFD45DFFB3076A544AFB0C7CC8FB7C" blockId="50.[813,1452,1167,1333]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFB0C7CC8FB7C" pageId="50" pageNumber="160">
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2. Carapace with relatively small lateral spines, 1 cardiac spine (
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<figureCitation id="133BC8D8FFB3076A543DFB2F7AD5FB61" box="[858,951,1196,1218]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Fig. 24E</figureCitation>
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); pseudorostral spines directed horizontally (
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<figureCitation id="133BC8D8FFB3076A52E4FB2F7A1DFB7C" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Fig. 26A</figureCitation>
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);
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<collectingCountry id="F31794CDFFB3076A54F5FB4A7A9CFB7C" box="[914,1022,1225,1247]" name="Philippines" pageId="50" pageNumber="160">Philippines</collectingCountry>
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............
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<taxonomicName id="4C00AFDEFFB3076A5320FB4A7CC8FB7C" ID-CoL="6LYQD" authority="(Rathbun, 1916)" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[1095,1450,1225,1247]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="suluensis">
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<emphasis id="B974084FFFB3076A5320FB4A7C61FB7D" box="[1095,1283,1225,1246]" italics="true" pageId="50" pageNumber="160">Holthuija suluensis</emphasis>
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(
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<bibRefCitation id="EF91A9ACFFB3076A5276FB4A7CC1FB7C" author="Rathbun MJ" box="[1297,1443,1225,1247]" pageId="50" pageNumber="160" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer " Albatross, " 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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<paragraph id="8BBFD45DFFB3076A544AFB657CC9FA96" blockId="50.[813,1452,1167,1333]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFB657CC9FA96" pageId="50" pageNumber="160">
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– Carapace with relatively long lateral spines, 2 or 3 cardiac spines (
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<figureCitation id="133BC8D8FFB3076A543FFA817AADFABB" box="[856,975,1282,1304]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">Fig. 25B–D</figureCitation>
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); pseudorostral spines gently curved downwards (
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<figureCitation id="133BC8D8FFB3076A543DFA9C7AD7FA96" box="[858,949,1311,1333]" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">Fig. 26D</figureCitation>
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); Eastern Indian Ocean........
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<taxonomicName id="4C00AFDEFFB3076A53AAFA9C7C0FFA96" ID-CoL="3MBWP" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1229,1389,1311,1333]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="aussie" status="sp. nov.">
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<emphasis id="B974084FFFB3076A53AAFA9C7C0FFA96" box="[1229,1389,1311,1333]" italics="true" pageId="50" pageNumber="160">Holthuija aussie</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B534FFB3076A5212FAA37CC9FA96" box="[1397,1451,1312,1333]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30F4CCF8FFB3076A544AFADA7CC8F9B8" pageId="50" pageNumber="160">
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<paragraph id="8BBFD45DFFB3076A544AFADA7CC8FA67" blockId="50.[813,1452,1369,1563]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFADA7CC8FA67" pageId="50" pageNumber="160">
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3. Carapace with relatively short lateral spines, pseudorostral spines almost straight or gently curved outward, appears level from lateral view and not prominently curved upward (
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<figureCitation id="133BC8D8FFB3076A521CFA117AF9FA67" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24A–D</figureCitation>
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,
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<figureCitation id="133BC8D8FFB3076A54C0FA2C7AB3FA66" box="[935,977,1455,1477]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25A</figureCitation>
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) ...........................................................................4
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</keyLead>
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</paragraph>
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<paragraph id="8BBFD45DFFB3076A544AFA4F7CC8F9B8" blockId="50.[813,1452,1369,1563]" pageId="50" pageNumber="160">
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<keyLead id="30F17768FFB3076A544AFA4F7CC8F9B8" pageId="50" pageNumber="160">
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– Carapace with relatively long lateral spines, pseudorostral spines strongly curved outward and usually distinctly bent or curved upward (
|
||
<figureCitation id="133BC8D8FFB3076A5493F9867D36F9B8" box="[1012,1108,1541,1563]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Figs. 24F</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A5307F9867DC1F9B8" box="[1120,1187,1541,1563]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">25E, F</figureCitation>
|
||
,
|
||
<figureCitation id="133BC8D8FFB3076A53C8F9867C6DF9B8" box="[1199,1295,1541,1563]" captionStart="Fig" captionStartId="56.[113,148,1208,1230]" captionTargetBox="[113,1451,190,1192]" captionTargetId="figure-15@56.[113,1451,190,1193]" captionTargetPageId="56" captionText="Fig. 26. Lateral views of carapaces, Holthuija species. A, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; B, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; C, H. pauli n. sp., paratype male (35.0 × 27.0 mm) (ZRC 2013.1378), Philippines; D, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea [laterally inverted to match earlier figures]; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea laterally inverted to match earlier figures]." figureDoi="http://doi.org/10.5281/zenodo.5384648" httpUri="https://zenodo.org/record/5384648/files/figure.png" pageId="50" pageNumber="160">26B, E, F</figureCitation>
|
||
)......................5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30F4CCF8FFB3076A544AF9BC7CC9F946" pageId="50" pageNumber="160">
|
||
<paragraph id="8BBFD45DFFB3076A544AF9BC7CC8F92C" blockId="50.[813,1451,1599,1765]" pageId="50" pageNumber="160">
|
||
<keyLead id="30F17768FFB3076A544AF9BC7CC8F92C" pageId="50" pageNumber="160">
|
||
4. Pseudorostral spines gently curved outward (
|
||
<figureCitation id="133BC8D8FFB3076A5245F9BC7CC2F9F6" box="[1314,1440,1599,1621]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Fig. 24A–D</figureCitation>
|
||
); G1 strongly curved with small flap distally (
|
||
<figureCitation id="133BC8D8FFB3076A5245F9DF7CC2F9D1" box="[1314,1440,1628,1650]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. A–C, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; F–H, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; I–K, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; L–O, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [A–F, I, L]; 0.5 mm [D, G, H, J, K, M–O]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="50" pageNumber="160">Fig. 27A–C</figureCitation>
|
||
);
|
||
<collectingCountry id="F31794CDFFB3076A5434F9FA7AD6F92C" box="[851,948,1657,1679]" name="Singapore" pageId="50" pageNumber="160">Singapore</collectingCountry>
|
||
.............................
|
||
<taxonomicName id="4C00AFDEFFB3076A5300F9FA7CC8F92C" ID-CoL="3MBWR" authority="(Walker, 1887)" baseAuthorityName="Walker" baseAuthorityYear="1887" box="[1127,1450,1657,1679]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="miersii">
|
||
<emphasis id="B974084FFFB3076A5300F9FA7C6CF92D" box="[1127,1294,1657,1678]" italics="true" pageId="50" pageNumber="160">Holthuija miersii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF91A9ACFFB3076A527BF9FA7CC1F92C" author="Walker AO" box="[1308,1443,1657,1679]" pageId="50" pageNumber="160" pagination="107 - 117" refId="ref89616" refString="Walker AO (1887) Notes on a collection of Crustacea from Singapore. Journal of the Linnean Society of London, 20 (118): 107 - 117, pls. 6 - 9." type="journal article" year="1887">Walker, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BBFD45DFFB3076A544AF9157CC9F946" blockId="50.[813,1451,1599,1765]" pageId="50" pageNumber="160">
|
||
<keyLead id="30F17768FFB3076A544AF9157CC9F946" pageId="50" pageNumber="160">
|
||
– Pseudorostral spines almost straight (
|
||
<figureCitation id="133BC8D8FFB3076A53A8F9167C4CF908" box="[1231,1326,1685,1707]" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">Fig. 25A</figureCitation>
|
||
); G1 almost straight with distal part dilated (
|
||
<figureCitation id="133BC8D8FFB3076A53E9F9317D9CF96B" box="[1166,1278,1714,1736]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. A–C, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; F–H, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; I–K, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; L–O, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [A–F, I, L]; 0.5 mm [D, G, H, J, K, M–O]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="50" pageNumber="160">Fig. 27I–K</figureCitation>
|
||
);
|
||
<collectingCountry id="F31794CDFFB3076A5277F9307C25F96B" box="[1296,1351,1715,1736]" name="Japan" pageId="50" pageNumber="160">Japan</collectingCountry>
|
||
................ ...........................................................
|
||
<taxonomicName id="4C00AFDEFFB3076A53DBF94C7C0FF946" ID-CoL="3MBWQ" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1212,1389,1743,1765]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="cognata" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A53DBF94C7C0FF946" box="[1212,1389,1743,1765]" italics="true" pageId="50" pageNumber="160">Holthuija cognata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A5212F9537CC9F946" box="[1397,1451,1744,1765]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30F4CCF8FFB3076A544AF88A7CC9F80D" pageId="50" pageNumber="160">
|
||
<paragraph id="8BBFD45DFFB3076A544AF88A7CC9F8FB" blockId="50.[813,1452,1801,1967]" pageId="50" pageNumber="160">
|
||
<keyLead id="30F17768FFB3076A544AF88A7CC9F8FB" pageId="50" pageNumber="160">
|
||
5. Dorsal surface of carapace with low granules (
|
||
<figureCitation id="133BC8D8FFB3076A5245F88A7C19F8BC" box="[1314,1403,1801,1823]" captionStart="Fig" captionStartId="54.[113,148,2005,2027]" captionTargetBox="[113,1451,561,1990]" captionTargetId="figure-15@54.[113,1451,561,1990]" captionTargetPageId="54" captionText="Fig. 24. General habitus, Holthuija species. A, Holthuija miersi (as Maia miersii, after Walker, 1887: pl. 6, fig. 1); B, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; C, H. miersii, female (16.5 × 12.2 mm) (ZRC 1995.311), Singapore; D, H. miersii, female (21.4 × 16.7 mm) (NHM 1900.10.22.13), Singapore; E, H. suluensis, holotype female (32.4 × 41.2 mm) (USNM 48224a), Philippines; F, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384644" httpUri="https://zenodo.org/record/5384644/files/figure.png" pageId="50" pageNumber="160">Fig. 24F</figureCitation>
|
||
); G1 almost straight with distal part slightly dilated (
|
||
<figureCitation id="133BC8D8FFB3076A524CF8A67CC2F89F" box="[1323,1440,1829,1852]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. A–C, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; F–H, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; I–K, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; L–O, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [A–F, I, L]; 0.5 mm [D, G, H, J, K, M–O]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="50" pageNumber="160">Fig. 27F–H</figureCitation>
|
||
);
|
||
<collectingCountry id="F31794CDFFB3076A5435F8C17AA3F8FB" box="[850,961,1858,1880]" name="Philippines" pageId="50" pageNumber="160">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31794CDFFB3076A54ABF8C17D76F8FB" box="[972,1044,1858,1880]" name="Taiwan" pageId="50" pageNumber="160">Taiwan</collectingCountry>
|
||
................................
|
||
<taxonomicName id="4C00AFDEFFB3076A53BFF8C17C0FF8F4" ID-CoL="3MBWS" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1240,1389,1858,1879]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="pauli" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A53BFF8C17C0FF8F4" box="[1240,1389,1858,1879]" italics="true" pageId="50" pageNumber="160">Holthuija pauli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A5212F8C07CC9F8FB" box="[1397,1451,1859,1880]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BBFD45DFFB3076A544AF8E37CC9F80D" blockId="50.[813,1452,1801,1967]" pageId="50" pageNumber="160">
|
||
<keyLead id="30F17768FFB3076A544AF8E37CC9F80D" pageId="50" pageNumber="160">
|
||
– Dorsal surface of carapace with distinct rounded granules (
|
||
<figureCitation id="133BC8D8FFB3076A52E2F8DC7AF5F832" captionStart="Fig" captionStartId="55.[136,171,1961,1983]" captionTargetBox="[136,1474,234,1945]" captionTargetId="figure-8@55.[136,1474,232,1947]" captionTargetPageId="55" captionText="Fig. 25. General habitus, Holthuija species. A, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; B, H. aussie n. sp., holotype ovigerous female (42.1 × 34.4 mm) (NMV J63752), Arafura Sea; C, H. aussie n. sp., paratype juvenile female (11.9 × 9.1 mm) (NMV J53958), Australia; D, H. aussie n. sp., juvenile female (14.3 × 11.9 mm) (NMV J61058), Australia; E, H. poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea; F, H. poorei n. sp., paratype female (27.0 × 22.5 mm) (NMV J63751), Arafura Sea." figureDoi="http://doi.org/10.5281/zenodo.5384646" httpUri="https://zenodo.org/record/5384646/files/figure.png" pageId="50" pageNumber="160">Fig. 25E, F</figureCitation>
|
||
); G1 curved, distal part simple (
|
||
<figureCitation id="133BC8D8FFB3076A53BEF8FF7C32F831" box="[1241,1360,1916,1938]" captionStart="Fig" captionStartId="57.[135,170,1644,1666]" captionTargetBox="[146,1470,190,1627]" captionTargetId="figure-8@57.[140,1479,186,1634]" captionTargetPageId="57" captionText="Fig. 27. Left G1s, Holthuija species. A–C, H. miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; D, E, H. suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; F–H, H. pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; I–K, H. cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; L–O, H. poorei n. sp., paratype male (26.1 × 20.1 mm) (NMV J63169), Timor Sea. Scale bars = 1.0 mm [A–F, I, L]; 0.5 mm [D, G, H, J, K, M–O]; 0.25 mm [E]." figureDoi="http://doi.org/10.5281/zenodo.5384650" httpUri="https://zenodo.org/record/5384650/files/figure.png" pageId="50" pageNumber="160">Fig. 27L–O</figureCitation>
|
||
); Eastern Indian Ocean.........................................
|
||
<taxonomicName id="4C00AFDEFFB3076A53ACF81A7C0FF80D" ID-CoL="3MBWT" authority="Ng & Forges, 2015" authorityName="Ng & Forges" authorityYear="2015" box="[1227,1389,1945,1966]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="50" pageNumber="160" phylum="Arthropoda" rank="species" species="poorei" status="sp. nov.">
|
||
<emphasis id="B974084FFFB3076A53ACF81A7C0FF80D" box="[1227,1389,1945,1966]" italics="true" pageId="50" pageNumber="160">Holthuija poorei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B534FFB3076A5212F81A7CC9F80D" box="[1397,1451,1945,1966]" pageId="50" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |