treatments-xml/data/E2/88/24/E28824C68DB052F68688C5F1D149E573.xml
2024-06-21 12:55:09 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e82644" ID-GBIF-Dataset="1d579114-3554-4e38-bd81-ec33a8c44b9f" ID-Pensoft-Pub="1864-8312-80-279" ID-Pensoft-UUID="42DA6B89695151758680B79C0CF21997" ID-ZooBank="1D57CB0C56C44DA591E1CF7E9BB40E74" ModsDocID="1864-8312-80-279" checkinTime="1658318833579" checkinUser="pensoft" docAuthor="Yin, Zi-Wei, Tihelka, Erik, Lozano-Fernandez, Jesus &amp; Cai, Chen-Yang" docDate="2022" docId="E28824C68DB052F68688C5F1D149E573" docLanguage="en" docName="ArthropodSystPhyl 80: 279-294" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-07-19" docSource="http://dx.doi.org/10.3897/asp.80.e82644" docTitle="Europharinodes Yin &amp; Cai 2022, gen. nov." docType="treatment" docUuid="0B631F8C-D490-4174-89F9-EE6274592F92" docUuidSource="ZooBank" docVersion="2" id="42DA6B89695151758680B79C0CF21997" lastPageNumber="279" masterDocId="42DA6B89695151758680B79C0CF21997" masterDocTitle="The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications" masterLastPageNumber="294" masterPageNumber="279" pageNumber="279" updateTime="1658319438294" updateUser="ExternalLinkService">
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<mods:title>The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6659-9448</mods:nameIdentifier>
<mods:affiliation>Laboratory of Systematic Entomology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai, 200234, China</mods:affiliation>
<mods:nameIdentifier type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:namePart>Tihelka, Erik</mods:namePart>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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<mods:namePart>Lozano-Fernandez, Jesus</mods:namePart>
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<mods:affiliation>Department of Genetics, Microbiology and Statistics, Biodiversity Research Institute (IRBio), University of Barcelona, Avd. Diagonal 643, 08028</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cai, Chen-Yang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9283-8323</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-07-19</mods:number>
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<treatment LSID="urn:lsid:zoobank.org:act:0B631F8C-D490-4174-89F9-EE6274592F92" httpUri="http://treatment.plazi.org/id/E28824C68DB052F68688C5F1D149E573" lastPageNumber="279" pageId="0" pageNumber="279">
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<paragraph pageId="0" pageNumber="279">
<taxonomicName LSID="http://zoobank.org/0B631F8C-D490-4174-89F9-EE6274592F92" authority="Yin &amp; Cai" authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Europharinodes" order="Coleoptera" pageId="0" pageNumber="279" rank="genus" status="gen. nov.">Europharinodes Yin &amp; Cai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="279">gen. nov.</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="279" type="description">
<paragraph pageId="0" pageNumber="279">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="type species">
<paragraph pageId="0" pageNumber="279">Type species.</paragraph>
<paragraph pageId="0" pageNumber="279">
<taxonomicName authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi">
<emphasis italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="279">sp. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="diagnosis">
<paragraph pageId="0" pageNumber="279">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="279">Most of body covered with squamous setae; body generally compact; head, pronotum and femora coarsely punctate. Head roundly-triangular, with short and narrow frontal rostrum; large vertexal and frontal foveae present; antennal insertions close, antennomeres slightly elongate to sub-moniliform, clubs formed by apical three enlarged antennomeres; lacking ocular-mandibular carinae; clypeus sharply sloping, anterior margin carinate and rounded, lateral margins straight to eyes; maxillary palpi small, four-segmented, with short apical palpal cone. Pronotum with median and lateral antebasal foveae obscured by squamous setae; with antero-hypomeral foveae, lacking hypomeral carinae. Elytra each with two large basal foveae and two distinct discal striae. Each tarsus plesiomorphically with two subequal claws. Abdomen with tergite 1 (IV) slightly longer than 2 (V), tergites 1-4 (IV-VII) broadly sulcate at bases; paratergites moderately broad and laterally protruding. Aedeagus symmetrical, basal capsule enlarged, paired parameres elongate.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="description">
<paragraph pageId="0" pageNumber="279">Description.</paragraph>
<paragraph pageId="0" pageNumber="279">
Habitus (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2B-D</figureCitation>
;
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3A, B</figureCitation>
) stout, compact; most of body with squamous setae. Head (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
) roundly triangular; vertex coarsely punctate, vertexal foveae (= dorsal tentorial pits; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">vf</emphasis>
) relatively large, obscured by squamous setae; frontal fovea (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ff</emphasis>
) distinct, rostrum short and narrow, antennal insertions located at ventral surface of rostrum, tubercles barely raised; compound eyes (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ce</emphasis>
) large, prominent; lacking ocular-mandibular carina; clypeus (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">cl</emphasis>
) sharply sloping, with broadly rounded, ridged anterior margin; labrum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">la</emphasis>
) transverse, subtrapezoidal; mandible (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ma</emphasis>
) with single distinct apical and preapical teeth; maxillary palpus small (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H, I</figureCitation>
), four-segmented, with palpomere 1 small, indistinct, 2 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">p2</emphasis>
) pedunculate basally and broadened toward apex, 3 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">p3</emphasis>
) roundly triangular, 4 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">p4</emphasis>
) sub-fusiform, with short palpal cone at apex. Venter with small, widely separated gular foveae (= posterior tentorial pits; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3F</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">gf</emphasis>
) in shared transverse impression, gular moderately raised along middle, weakly impressed admesally; neck region broad. Paired tentorial arms (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ta</emphasis>
) V-shaped, each branched at base and extending anteriorly to reach inner wall of clypeus (right arm broken at apex in holotype). Pharynx (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ph</emphasis>
) about 2/3 of head length, posteriorly broadened. Antenna 11-segmented, moderately elongate, extending to approximately half of elytral length when extending posteriorly, antennomeres each slightly elongate to sub-moniliform, club (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3F</figureCitation>
) formed by enlarged apical three antennomeres.
</paragraph>
<caption doi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="279">
<emphasis bold="true" pageId="0" pageNumber="279">Figure 2.</emphasis>
Amber piece (A) containing
<taxonomicName authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi">
<emphasis italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
, holotype, SNUC-Paleo-0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction).
<emphasis bold="true" pageId="0" pageNumber="279">A</emphasis>
Original amber piece containing the holotype before cutting and polishing.
<emphasis bold="true" pageId="0" pageNumber="279">B</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">C</emphasis>
Left lateral habitus.
<emphasis bold="true" pageId="0" pageNumber="279">D</emphasis>
Right lateral habitus.
<emphasis bold="true" pageId="0" pageNumber="279">E</emphasis>
Head in lateral view, show tentorial arms and pharynx.
<emphasis bold="true" pageId="0" pageNumber="279">F</emphasis>
Right antennal club.
<emphasis bold="true" pageId="0" pageNumber="279">G</emphasis>
Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F.
</paragraph>
</caption>
<caption doi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="279">
<emphasis bold="true" pageId="0" pageNumber="279">Figure 3.</emphasis>
Morphology of
<taxonomicName authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi">
<emphasis italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
, holotype, SNUC-Paleo-0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction).
<emphasis bold="true" pageId="0" pageNumber="279">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">B</emphasis>
Dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="279">C</emphasis>
Left half of pronotum, showing squamous setae.
<emphasis bold="true" pageId="0" pageNumber="279">D</emphasis>
Lateral portion of the abdominal base, showing squamous setae.
<emphasis bold="true" pageId="0" pageNumber="279">E</emphasis>
Head dorsum.
<emphasis bold="true" pageId="0" pageNumber="279">F</emphasis>
Head venter.
<emphasis bold="true" pageId="0" pageNumber="279">G</emphasis>
Head, in antero-lateral view.
<emphasis bold="true" pageId="0" pageNumber="279">H</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">I</emphasis>
Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p2-4, maxillary palpomere 2-4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="279">
Pronotum (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4A</figureCitation>
) approximately as long as broad, sides rounded, broadest at middle, anteriorly and posteriorly narrowed, surface with dense squamous setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3C</figureCitation>
); disc coarsely punctate, with squamous setae all over surface, covering median and lateral antebasal foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4A, B</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">maf</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="279">laf</emphasis>
), lacking sulcus or carina. Hypomeron extended, lacking carina, with anterior hypomeral fovea (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4B, C</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ahf</emphasis>
). Prosternum (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4C</figureCitation>
) with widely separated lateral procoxal foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4C</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">lpcf</emphasis>
), anterior part shorter than coxal part, margins of coxal cavity moderately carinate.
</paragraph>
<caption doi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="279">
<emphasis bold="true" pageId="0" pageNumber="279">Figure 4.</emphasis>
X-ray microtomographic reconstruction of
<taxonomicName authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi">
<emphasis italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
, holotype, SNUC-Paleo-0102.
<emphasis bold="true" pageId="0" pageNumber="279">A</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="279">C</emphasis>
: prothorax, dorsal (A), lateral (B) and ventral (C).
<emphasis bold="true" pageId="0" pageNumber="279">D</emphasis>
Right elytron.
<emphasis bold="true" pageId="0" pageNumber="279">E</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">F</emphasis>
Meso- and metathorax, ventral (E) and lateral (F).
<emphasis bold="true" pageId="0" pageNumber="279">G</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="279">I</emphasis>
Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt1-3, paratergite 1-3; s2-6, sternite 2-6 (IV-VIII); t1-5, tergite 1-5 (IV-VIII). Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="279">
Elytra broadly truncate at bases, each elytron with two large basal foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4D</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">bef</emphasis>
), with two long discal striae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ds</emphasis>
) extending from foveae to posterior elytral margin, lacking subhumeral fovea or marginal sulcus. Metathoracic (hind) wings (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2D</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">hw</emphasis>
) fully developed.
</paragraph>
<paragraph pageId="0" pageNumber="279">
Mesoventrite with median foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">mmsf</emphasis>
) originating from shared transverse impression, moderately separated, with pairs of large lateral and distinct anterolateral mesoventral foveae (not shown in figure); metaventrite with large, setose lateral coxal foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">lmcf</emphasis>
), with single median metaventral fovea (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">mmtf</emphasis>
), posterior margin with small, narrow split at middle (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">ms</emphasis>
); metacoxae broadly separated. Marginal carina of meso- and metaventrite complete (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">mc</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="279">
Legs (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5A-C</figureCitation>
) moderately elongate; femora roughly punctate; with short tarsomere 1 and long tarsomere 2 and 3, 2 slightly longer than 3, each tarsus with two subequal pretarsal claws (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2G</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="279">
<emphasis bold="true" pageId="0" pageNumber="279">Figure 5.</emphasis>
Morphology of
<taxonomicName authorityName="Yin &amp; Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi">
<emphasis italics="true" pageId="0" pageNumber="279">Europharinodes schaufussi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="279">gen. et sp. nov.</emphasis>
, holotype, SNUC-Paleo-0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic).
<emphasis bold="true" pageId="0" pageNumber="279">A</emphasis>
Right foreleg.
<emphasis bold="true" pageId="0" pageNumber="279">B</emphasis>
Right middle leg.
<emphasis bold="true" pageId="0" pageNumber="279">C</emphasis>
Left hindleg.
<emphasis bold="true" pageId="0" pageNumber="279">D</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="279">F</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">I</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="279">K</emphasis>
Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K).
<emphasis bold="true" pageId="0" pageNumber="279">G</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="279">H</emphasis>
endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="279">
Abdomen constricted at base, with dense squamous setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3D</figureCitation>
); tergite 1 (IV) slightly longer than 2 (V), 2-4 (V-VII) subequal in length, tergites 1-4 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G, H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">t1-4</emphasis>
) each deeply and broadly sulcate at base, at lease with one pair of basolateral foveae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">blf</emphasis>
), tergite 5 (VIII) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">t5</emphasis>
) roundly triangular, transverse, medially roundly emarginate at posterior margin; sternite 2-5 (IV-VII) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H, I</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">s2-5</emphasis>
) each broadly sulcate at base, with one pair of basolateral foveae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3H</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">blf</emphasis>
), 6 (VIII), sternite 6 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4H, I</figureCitation>
;
<emphasis italics="true" pageId="0" pageNumber="279">s6</emphasis>
) transverse.
</paragraph>
<paragraph pageId="0" pageNumber="279">
Males have modified sternites 4 and 5 (VI and VII). Aedeagus (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5D-F, I-K</figureCitation>
) dorso-ventrally symmetrical.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="etymology">
<paragraph pageId="0" pageNumber="279">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="279">
The new generic name is a combination of Latin '
<emphasis italics="true" pageId="0" pageNumber="279">
<normalizedToken originalValue="Eurōpa">Eurōpa</normalizedToken>
</emphasis>
(Europe)' and genus
<taxonomicName authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Apharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apharinodes" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Apharinodes</emphasis>
</taxonomicName>
, referring to the origin of the fossil in Baltic amber and its affinity with
<taxonomicName authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Apharinodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apharinodes" order="Coleoptera" pageId="0" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Apharinodes</emphasis>
</taxonomicName>
. The gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>