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<document id="561D19F2BC7BE6ADDADCD9F35BD6881E" ID-DOI="10.5281/zenodo.197810" ID-GBIF-Dataset="3fed616e-ea51-496b-b522-145b87775b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="197810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460279155734" checkinUser="plazi" docAuthor="Rahmadi, Cahyo, Harvey, Mark S. &amp; Kojima, Jun-Ichi" docDate="2010" docId="AD1687F3FF8DFF900CEB49FA2F74F908" docLanguage="en" docName="zt02612p021.pdf" docOrigin="Zootaxa 2612" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sarax sangkulirangensis Rahmadi, Harvey &amp; Kojima, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="512FFF8BFF87FF9C0C7C4D7D2E59FFA4" masterDocTitle="Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island" masterLastPageNumber="21" masterPageNumber="1" pageNumber="11" updateTime="1698249151004" updateUser="plazi">
<mods:mods id="2F59C1B3FD7FA2B5113FDB1BF9E2D95D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1B2698CA670BE3B60558BD265901C2C2">
<mods:title id="20F3190D159E2C7A1DBBD661B46A2FEF">Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island</mods:title>
</mods:titleInfo>
<mods:name id="BBCF45642F749D5938D1DB64457BD403" type="personal">
<mods:role id="14B745F6BC4E1CB70B4073A68A52714E">
<mods:roleTerm id="BCB80A9D5BD86AFEB8BA8AA26AA35731">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4158E34907CA1907D1657447D0A4592C">Rahmadi, Cahyo</mods:namePart>
</mods:name>
<mods:name id="EF439E18CA3C92042B675F9F7B0F2063" type="personal">
<mods:role id="F837233F33C7AC51C5323EE8A485DF0A">
<mods:roleTerm id="FA9FB70568727ED9FAB340839BE01B91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A70027ACD7A8AE1DC70307A903E7F7D">Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:name id="741DC3A6BD7BA46A653A9E83A2C55444" type="personal">
<mods:role id="33C38E4F2B8F6579A00A70BCC77D9E97">
<mods:roleTerm id="03E456A3C95963008D209F891DB296F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D84B9759171BCC33BA8433492FC49472">Kojima, Jun-Ichi</mods:namePart>
</mods:name>
<mods:typeOfResource id="52F2C51AC0559779117740DCF94E5D08">text</mods:typeOfResource>
<mods:relatedItem id="2209694ACA79E2554F848E20AA1234E1" type="host">
<mods:titleInfo id="BF71AE7766BDB3EBFF2DBBD7DB12F46E">
<mods:title id="6B49610AAB1FE403896F4EBB52BBE341">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1465670D121182AA2696B594841D13EE">
<mods:date id="A2D17A0C636DBED70A1C76A1AC798377">2010</mods:date>
<mods:detail id="C1A97E7494A9F4B0228A02AAD1BF4D0A" type="volume">
<mods:number id="D6F12BF341643E5A2B9B4882EBADD906">2612</mods:number>
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<mods:start id="D4583176FF5A8D653E0A0ED3D6D42553">1</mods:start>
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<mods:classification id="75CD11895B9E0D632F8B04F0F1321607">journal article</mods:classification>
<mods:identifier id="578F65E69D0ADD771125B68760ECCC7B" type="DOI">10.5281/zenodo.197810</mods:identifier>
<mods:identifier id="81AD83C62415C4784B3C0BC0E4B25498" type="GBIF-Dataset">3fed616e-ea51-496b-b522-145b87775b5a</mods:identifier>
<mods:identifier id="63A35BDF9983D59253016B678FB754EB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="324609239AC2E38DFC5412E35FF847F4" type="Zenodo-Dep">197810</mods:identifier>
</mods:mods>
<treatment id="AD1687F3FF8DFF900CEB49FA2F74F908" ID-DOI="http://doi.org/10.5281/zenodo.6207615" ID-GBIF-Taxon="119396509" ID-Zenodo-Dep="6207615" LSID="urn:lsid:plazi:treatment:AD1687F3FF8DFF900CEB49FA2F74F908" httpUri="http://treatment.plazi.org/id/AD1687F3FF8DFF900CEB49FA2F74F908" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="6DA5656EFF8DFF960CEB49FA2FD5FB60" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="250036E5FF8DFF960CEB49FA2C71FB05" blockId="10.[151,552,1159,1220]" box="[151,552,1159,1185]" pageId="10" pageNumber="11">
<heading id="7E488189FF8DFF960CEB49FA2C71FB05" bold="true" box="[151,552,1159,1185]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="17CBEAF7FF8DFF960CEB49FA2C71FB05" bold="true" box="[151,552,1159,1185]" pageId="10" pageNumber="11">
<taxonomicName id="E2BF4D66FF8DFF960CEB49FA2F9AFB05" ID-CoL="6XMF7" box="[151,451,1159,1185]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sangkulirangensis" status="sp. nov.">
<emphasis id="17CBEAF7FF8DFF960CEB49FA2F9AFB05" bold="true" box="[151,451,1159,1185]" italics="true" pageId="10" pageNumber="11">Sarax sangkulirangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CF8578CFF8DFF960DB749FA2C71FB05" box="[459,552,1159,1185]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="250036E5FF8DFF960CEB49D72FD5FB60" blockId="10.[151,552,1159,1220]" box="[151,396,1194,1220]" pageId="10" pageNumber="11">
(
<figureCitation id="BD842A60FF8DFF960CE349D72F74FB60" box="[159,301,1194,1220]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="10" pageNumber="11">Figs. 2125</figureCitation>
,
<figureCitation id="BD842A60FF8DFF960D4649D72FDDFB60" box="[314,388,1194,1220]" captionStart="FIGURES 33 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="10" pageNumber="11">3738</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8DFF960CEB49882B1DF9B0" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="250036E5FF8DFF960CEB49882B1DF9B0" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
<emphasis id="17CBEAF7FF8DFF960CEB49882FCBFAAB" bold="true" box="[151,402,1269,1295]" pageId="10" pageNumber="11">Material examined:</emphasis>
Female
<typeStatus id="FA048847FF8DFF960D8749882C3AFAAB" box="[507,611,1269,1295]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="43AEAE20FF8DFF960E0849882CE0FAAB" box="[628,697,1269,1295]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="11">MZB</collectionCode>
.Ambl.070),
<collectingCountry id="5DA87675FF8DFF960F2C49882DAEFAAB" box="[848,1015,1269,1295]" name="Indonesia" pageId="10" pageNumber="11">INDONESIA</collectingCountry>
:
<emphasis id="17CBEAF7FF8DFF96087E49882A8EFAAA" box="[1026,1239,1269,1294]" italics="true" pageId="10" pageNumber="11">East Kalimantan:</emphasis>
Berau Regency, District Tabalar Ulu, Gua Ke 4, GPS coordinates
<geoCoordinate id="408B5022FF8DFF960E8848672DC5FA90" box="[756,924,1306,1332]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.4703417">1°28'13.23&quot;N</geoCoordinate>
,
<geoCoordinate id="408B5022FF8DFF960FD648672A30FA90" box="[938,1129,1306,1332]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.65782">117°39'28.14&quot;E</geoCoordinate>
(approximate),
<date id="51011025FF8DFF96095148672E8CFAFE" pageId="10" pageNumber="11" value="2004-08-09">9 August 2004</date>
, C. Rahmadi, ovigerous.
<typeStatus id="FA048847FF8DFF960E7E483D2C20FAFE" box="[514,633,1344,1370]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="33B9FD6CFF8DFF960EF9483D2C8CFAFE" box="[645,725,1344,1370]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<collectionCode id="43AEAE20FF8DFF960E9A483D2D72FAFE" box="[742,811,1344,1370]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="11">MZB</collectionCode>
.Ambl.150), Berau Regency, Kelai District, Merapun Village, near Tebo Lake, Gua Danum Tengen, GPS coordinates
<geoCoordinate id="408B5022FF8DFF960FF048182A77FADB" box="[908,1070,1381,1407]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.5245833">1°31'28.50&quot;N</geoCoordinate>
,
<geoCoordinate id="408B5022FF8DFF96084748182ABFFADB" box="[1083,1254,1381,1407]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.36803">117°22'4.90&quot;E</geoCoordinate>
(approximate),
<date id="51011025FF8DFF960CEB48F72F25FA00" box="[151,380,1418,1444]" pageId="10" pageNumber="11" value="2004-09-02">2 September 2004</date>
, Y.R. Suhardjono;
<specimenCount id="33B9FD6CFF8DFF960E1B48F72CBAFA00" box="[615,739,1418,1444]" pageId="10" pageNumber="11" type="juvenile">1 juvenile</specimenCount>
(MNHN-Am.10), Berau Regency, Tabalar District, Gua Louwading, Tabalar Ulu,
<date id="51011025FF8DFF960DB748CD2CD5FA6E" box="[459,652,1456,1482]" pageId="10" pageNumber="11" value="2004-08-11">11 August 2004</date>
, L. Deharveng &amp; A. Bedos (KAL-059);
<specimenCount id="33B9FD6CFF8DFF96080F48CD2A9DFA6E" box="[1139,1220,1456,1482]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(MNHN-Am.11), Kutai Timur Regency, Sangkulirang District, Pengadan Village, Gua Ampanas, GPS coordinates
<geoCoordinate id="408B5022FF8DFF960CEB48872F60F9B0" box="[151,313,1530,1556]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.2004056">1°12'01.46&quot;N</geoCoordinate>
,
<geoCoordinate id="408B5022FF8DFF960D3948872FA7F9B0" box="[325,510,1530,1556]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.73432">117°44'03.58&quot;E</geoCoordinate>
(approximate),
<date id="51011025FF8DFF960ECA48872D28F9B0" box="[694,881,1530,1556]" pageId="10" pageNumber="11" value="2004-08-18">18 August 2004</date>
, L. Deharveng &amp; A. Bedos (KAL-113).
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8DFF960CB94B622F0DF90E" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="250036E5FF8DFF960CB94B622F0DF90E" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
<emphasis id="17CBEAF7FF8DFF960CB94B622F12F99D" bold="true" box="[197,331,1567,1593]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
<taxonomicName id="E2BF4D66FF8DFF960D284B5D2C27F99D" box="[340,638,1568,1593]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sangkulirangensis">
<emphasis id="17CBEAF7FF8DFF960D284B5D2C27F99D" box="[340,638,1568,1593]" italics="true" pageId="10" pageNumber="11">Sarax sangkulirangensis</emphasis>
</taxonomicName>
is relatively small sized (adult body length
<quantity id="E2479B00FF8DFF9608EB4B5D2B77F99E" box="[1175,1326,1567,1594]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.8" metricValueMin="5.9" pageId="10" pageNumber="11" unit="mm" value="7.85" valueMax="9.8" valueMin="5.9">
5.9
<emphasis id="17CBEAF7FF8DFF9608C14B622A95F99D" bold="true" box="[1213,1228,1567,1593]" italics="true" pageId="10" pageNumber="11"></emphasis>
9.8 mm
</quantity>
), lacking sexual dimorphism, the pedipalpal tarsus has two spines with the proximal spine about half as long as distal spine, and tibia of leg IV has 19 trichobothria with
<emphasis id="17CBEAF7FF8DFF960E904B162D5AF920" box="[748,771,1643,1668]" italics="true" pageId="10" pageNumber="11">bt</emphasis>
on the middle of fourth basitibial segment, and
<emphasis id="17CBEAF7FF8DFF96094F4B162B16F920" box="[1331,1359,1643,1668]" italics="true" pageId="10" pageNumber="11">bc</emphasis>
closer to
<emphasis id="17CBEAF7FF8DFF960CC94BED2E95F90D" box="[181,204,1680,1705]" italics="true" pageId="10" pageNumber="11">bf</emphasis>
than to
<emphasis id="17CBEAF7FF8DFF960D574BED2F14F90D" box="[299,333,1680,1705]" italics="true" pageId="10" pageNumber="11">sbf</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8DFF900CB94BC82F9CFB48" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="250036E5FF8DFF960CB94BC82CB8F8BE" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
<emphasis id="17CBEAF7FF8DFF960CB94BC82F3BF96B" bold="true" box="[197,354,1717,1743]" pageId="10" pageNumber="11">Description:</emphasis>
<emphasis id="17CBEAF7FF8DFF960D164BC82F97F96A" box="[362,462,1717,1742]" italics="true" pageId="10" pageNumber="11">Female:</emphasis>
Color in alcohol: Carapace brown, centrally with black marks, flange yellowishbrown; pedipalp yellowish-brown except for tarsus reddish-brown and spine yellow; legs yellowish-brown, but patella dark brown; abdomen greenish-brown.
</paragraph>
<paragraph id="250036E5FF8DFF960CB94A582B73F870" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
Carapace (
<figureCitation id="BD842A60FF8DFF960D3F4A582FC5F89B" box="[323,412,1829,1855]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="10" pageNumber="11">Fig. 21</figureCitation>
): Width about 1.4
<emphasis id="17CBEAF7FF8DFF960E0D4A582CD9F89B" bold="true" box="[625,640,1829,1855]" italics="true" pageId="10" pageNumber="11"></emphasis>
1.7 times its length; surface granulate, without setiferous tubercles; median sulcus deep in posterior one-third of the carapace; 3 pairs lateral sulci present; flange wide and bent upward; anterior margin slightly rounded, with 6 frontal setae. Median eye tubercle black, well-developed and high, without apical setae, slightly emarginate antero-medially to form heart-shape and median eyes facing antero-laterally; lateral eyes close to lateral margin of carapace, well-developed with black tapetum.
</paragraph>
<caption id="71C0666DFF8CFF970CEB49B82FE7FA99" httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12" targetBox="[204,1355,266,1164]" targetPageId="11">
<paragraph id="250036E5FF8CFF970CEB49B82FE7FA99" blockId="11.[151,1436,1221,1341]" pageId="11" pageNumber="12">
<emphasis id="17CBEAF7FF8CFF970CEB49B82F06FB79" bold="true" box="[151,351,1221,1245]" pageId="11" pageNumber="12">FIGURES 2125</emphasis>
.
<taxonomicName id="E2BF4D66FF8CFF970D1249BB2CDCFB79" box="[366,645,1222,1245]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sangkulirangensis" status="sp. nov.">
<emphasis id="17CBEAF7FF8CFF970D1249BB2CDCFB79" box="[366,645,1222,1245]" italics="true" pageId="11" pageNumber="12">Sarax sangkulirangensis</emphasis>
</taxonomicName>
<emphasis id="17CBEAF7FF8CFF970EF049B82CBEFB79" bold="true" box="[652,743,1221,1245]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="0CF8578CFF8CFF970EF049B82CBEFB79" box="[652,743,1221,1245]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB.Ambl.150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB.Ambl.150).
</paragraph>
</caption>
<paragraph id="250036E5FF8CFF970CB948162B72FA51" blockId="11.[151,1437,1387,2011]" pageId="11" pageNumber="12">
Chelicera (
<figureCitation id="BD842A60FF8CFF970D3A48162FC6FA21" box="[326,415,1387,1413]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
): Dorsal surface smooth, with 1 fine frontal seta and several fine setae. Basal segment with 4 teeth: lower-most tooth largest, upper-most tooth bicuspid, with upper cusp larger than lower; inner surface with several setae arranged in vertical row; outer surface with small tooth opposite bicuspid tooth, ventrally with several setae near proximal margin. Movable article with 5 teeth almost equal in size.
</paragraph>
<paragraph id="250036E5FF8CFF970CB94B7C2CA9F9C1" blockId="11.[151,1437,1387,2011]" pageId="11" pageNumber="12">
Sternum (
<figureCitation id="BD842A60FF8CFF970D474B7C2FCCF9BF" box="[315,405,1537,1563]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 23</figureCitation>
): First sternite (= tritosternum) elongate, with paired apical and several setae. Second and third sternites rounded and slightly rounded, respectively with 3 and 2 setae in addition to paired apical setae. Fourth sternite (= metasternum) with 5 setae.
</paragraph>
<paragraph id="250036E5FF8CFF900CB94B0C2A82FEE3" blockId="11.[151,1437,1387,2011]" lastBlockId="12.[151,1437,152,1708]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Pedipalp (
<figureCitation id="BD842A60FF8CFF970D3F4B0C2FFBF92F" box="[323,418,1649,1675]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
): Short and stout. Trochanter on antero-dorsal margin with 4 setiferous tubercles, without median spine; antero-ventral margin with ventro-anterior apophysis equipped with several setiferous tubercles on distal margin of trochanter. Femur: antero-dorsal margin with 4 major spines (length
<emphasis id="17CBEAF7FF8CFF970CEB4B9D2F0DF95E" bold="true" box="[151,340,1760,1786]" pageId="11" pageNumber="12">F1&gt;F2&gt;F3&gt;F4)</emphasis>
, 1 minor spine, several setiferous tubercles and small tubercles; antero-ventral margin with 4 major spines (length
<emphasis id="17CBEAF7FF8CFF970DEE4A782C2AF8BB" bold="true" box="[402,627,1797,1823]" pageId="11" pageNumber="12">FI&gt;FII&gt;FIII&gt;FIV</emphasis>
), 2 minor spines and small tubercles; 1 spine present dorsally of
<emphasis id="17CBEAF7FF8CFF9709024A782BC2F8BB" bold="true" box="[1406,1435,1797,1823]" pageId="11" pageNumber="12">FI</emphasis>
and as long as three-quarters length of
<emphasis id="17CBEAF7FF8CFF970E004A562CFAF8E1" bold="true" box="[636,675,1835,1861]" pageId="11" pageNumber="12">FI,</emphasis>
minor spine present between
<emphasis id="17CBEAF7FF8CFF9708644A562ADAF8E1" bold="true" box="[1048,1155,1835,1861]" pageId="11" pageNumber="12">FII -FIII</emphasis>
. Patella: antero-dorsal margin with 4 major spines (length
<emphasis id="17CBEAF7FF8CFF970E284A2D2D49F8CE" bold="true" box="[596,784,1872,1898]" pageId="11" pageNumber="12">P1&gt;P2&gt;P3&gt;P4</emphasis>
), clumped distally, 1 minor spine, several setiferous tubercles and small tubercles; 1 minor spine as long as half of
<emphasis id="17CBEAF7FF8CFF970FF94A082DFFF82B" bold="true" box="[901,934,1909,1935]" pageId="11" pageNumber="12">P1</emphasis>
presents between
<emphasis id="17CBEAF7FF8CFF9708FF4A082AFAF82B" bold="true" box="[1155,1187,1909,1935]" pageId="11" pageNumber="12">P1</emphasis>
and distal margin of patella, 1 spine presents between
<emphasis id="17CBEAF7FF8CFF970E5E4AE62C1AF811" bold="true" box="[546,579,1947,1973]" pageId="11" pageNumber="12">P4</emphasis>
and proximal margin; antero-ventral margin with 4 major spines (length
<emphasis id="17CBEAF7FF8CFF970CEB4ABD2F2FF87E" bold="true" box="[151,374,1984,2010]" pageId="11" pageNumber="12">PII&gt;PI&gt;PIII&gt;PIV</emphasis>
), several setiferous tubercles and small tubercles present. Tibia with outer surface smooth, with spines as follows: 2 major spines on antero-dorsal margin, proximal spine about half as long as distal one; 1 major spine on antero-ventral margin close to distal margin of tibia. Tarsus completely divided (claw clearly demarcated by articulation), with 2 spines on antero-dorsal margin separated from each other by about three times basal diameter of proximal spine: proximal spine shortest, length of distal spine about twice as long as proximal one; cleaning organ ventrally with about 30 modified hairs; apotele present.
</paragraph>
<paragraph id="250036E5FF8BFF900CB94C2F2F16FDD6" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
Legs (
<figureCitation id="BD842A60FF8BFF900D734C2F2F30FEC8" box="[271,361,338,364]" captionStart="FIGURES 21 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
): Femora of legs I
<emphasis id="17CBEAF7FF8BFF900E384C2F2C0AFEC8" bold="true" box="[580,595,338,364]" italics="true" pageId="12" pageNumber="13"></emphasis>
IV with small tubercles bearing setae. Tibia and tarsus of leg I with 23 and 41 segments, respectively; tibiae of legs
<date id="51011025FF8BFF900ED34C052D6BFE36" box="[687,818,376,402]" pageId="12" pageNumber="13">II and III 2</date>
-segmented; basitibia of leg
<date id="51011025FF8BFF9008F94C052AE5FE36" box="[1157,1212,376,402]" pageId="12" pageNumber="13">IV 4</date>
-segmented, fourth segment with 1 trichobothrium (value in parentheses: ratio of the distance from the trichobothrium to the proximal margin of the segment against the length of the segment),
<emphasis id="17CBEAF7FF8BFF900FB64CBF2DBBFE7F" box="[970,994,450,475]" italics="true" pageId="12" pageNumber="13">bt</emphasis>
(0.52); distitibiae of legs II
<emphasis id="17CBEAF7FF8BFF9009534CBF2B67FE78" bold="true" box="[1327,1342,450,476]" italics="true" pageId="12" pageNumber="13"></emphasis>
IV each with 18 trichobothria
<emphasis id="17CBEAF7FF8BFF900DE94C952FE3FDA5" box="[405,442,488,513]" italics="true" pageId="12" pageNumber="13">bf1</emphasis>
(0.07),
<emphasis id="17CBEAF7FF8BFF900E6B4C952C60FDA5" box="[535,569,488,513]" italics="true" pageId="12" pageNumber="13">sbf</emphasis>
(0.31),
<emphasis id="17CBEAF7FF8BFF900EEA4C952CE8FDA5" box="[662,689,488,513]" italics="true" pageId="12" pageNumber="13">bc</emphasis>
(0.16),
<emphasis id="17CBEAF7FF8BFF900F724C952D7DFDA5" box="[782,804,488,513]" italics="true" pageId="12" pageNumber="13">bt</emphasis>
close to distal margin of basitibial fourth segment,
<emphasis id="17CBEAF7FF8BFF9009FC4C952BC2FDA5" box="[1408,1435,488,513]" italics="true" pageId="12" pageNumber="13">bc</emphasis>
close to
<emphasis id="17CBEAF7FF8BFF900D7C4F702F7DFD82" box="[256,292,525,550]" italics="true" pageId="12" pageNumber="13">sbf</emphasis>
than to
<emphasis id="17CBEAF7FF8BFF900DF04F702FFDFD82" box="[396,420,525,550]" italics="true" pageId="12" pageNumber="13">bf</emphasis>
. Tarsi of legs
<date id="51011025FF8BFF900E244F702CEDFD83" box="[600,692,525,551]" pageId="12" pageNumber="13">
II
<emphasis id="17CBEAF7FF8BFF900E104F702C22FD83" bold="true" box="[620,635,525,551]" italics="true" pageId="12" pageNumber="13"></emphasis>
IV 4
</date>
-segmented; first segment more than length of subsequent 3 segments combined; second segment with light-yellow transverse line; fourth segment without oblique slit; pulvilli present.
</paragraph>
<paragraph id="250036E5FF8BFF900CB94F002C4AFD18" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">Genitalia: Gonopods covered ventrally with genital operculum, with posterior margin concave. Gonopods paired, soft, white and tube-like.</paragraph>
<paragraph id="250036E5FF8BFF900CB94FB52A7BFCF6" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
<emphasis id="17CBEAF7FF8BFF900CB94FB52F57FD45" box="[197,270,712,737]" italics="true" pageId="12" pageNumber="13">Male:</emphasis>
Similar to female pedipalpal femur with 3 major spines (
<emphasis id="17CBEAF7FF8BFF900FA84FBA2A3CFD45" bold="true" box="[980,1125,711,737]" pageId="12" pageNumber="13">F1&gt;F2&gt;F3)</emphasis>
on antero-dorsal margin. Genitalia: covered ventrally by genital operculum, slightly concave on posterior margin; paired pointedprojections posteriorly, lateral lobes with brown band on basal lobe (
<figureCitation id="BD842A60FF8BFF900F9D4E6F2A64FC88" box="[993,1085,786,812]" captionStart="FIGURES 33 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="12" pageNumber="13">Fig. 38</figureCitation>
). In dorsal view, soft, paired brownish band running from anterior to posterior to median section (
<figureCitation id="BD842A60FF8BFF900FC04E452A4DFCF6" box="[956,1044,824,850]" captionStart="FIGURES 33 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="12" pageNumber="13">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="250036E5FF8BFF900CB94E202F9CFB48" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
Measurements (in mm): male (n=2) [female (n=1)]; values for segments of the appendages are their lengths. Body length (excluding chelicera) 5.94
<emphasis id="17CBEAF7FF8BFF900EBD4EFF2C96FC38" bold="true" box="[705,719,898,924]" italics="true" pageId="12" pageNumber="13"></emphasis>
7.01 [9.85]. Carapace: median length 2.352.65 [3.63]; width 3.443.82 [5.28]; median eyes to anterior margin of carapace 0.030.07 [0.07]; distance between lateral eyes 1.651.87 [2.55]; lateral eye to anterior margin of carapace 0.320.45 [0.50]; lateral eye to lateral margin of carapace 0.140.15 [0.37]. Pedipalps: trochanter 0.971.01 [1.32]; femur 1.962.14 [3.44]; patella 2.322.45 [4.00]; tibia 1.111.50 [1.13]; tarsus 1.221.39 [2.01]. Leg I: femur 6.487.62 [12.13]; patella 0.490.55 [0.71]. Leg II: femur 3.384.53 [7.10], patella 0.620.72 [1.03]; basitibia 2.553.67[6.17]; distitibia 1.67 2.30 [3.51]; metatarsus + tarsus 1.131.56 [2.74]. Leg III: femur 4.065.53 [8.35]; patella 0.650.79 [1.16]; basitibia 3.334.83 [7.65]; distitibia 1.872.48 [3.70]; metatarsus+ tarsus 1.592.09 [2.89]. Leg IV: femur 3.604.81 [7.09]; patella 0.660.73 [0.58]; basitibia 3.42
<emphasis id="17CBEAF7FF8BFF900F4B49D02D1FFB63" bold="true" box="[823,838,1197,1223]" italics="true" pageId="12" pageNumber="13"></emphasis>
5.38 [7.52]; distitibia 1.61
<emphasis id="17CBEAF7FF8BFF90080049D02AD2FB63" bold="true" box="[1148,1163,1197,1223]" italics="true" pageId="12" pageNumber="13"></emphasis>
2.10 [3.33]; metatarsus + tarsus 1.80
<emphasis id="17CBEAF7FF8BFF900D5149AF2F65FB48" bold="true" box="[301,316,1234,1260]" italics="true" pageId="12" pageNumber="13"></emphasis>
1.92 [2.97].
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8BFF900CB9498A2A41FAB6" box="[197,1048,1271,1298]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="250036E5FF8BFF900CB9498A2A41FAB6" blockId="12.[151,1437,152,1708]" box="[197,1048,1271,1298]" pageId="12" pageNumber="13">
<emphasis id="17CBEAF7FF8BFF900CB9498A2F0CFAB5" bold="true" box="[197,341,1271,1297]" pageId="12" pageNumber="13">Etymology:</emphasis>
The species name is derived from the name of
<typeStatus id="FA048847FF8BFF900FFC49852DEBFAB6" box="[896,946,1272,1298]" pageId="12" pageNumber="13">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8BFF900CB948602DFAF9B3" pageId="12" pageNumber="13" type="distribution">
<paragraph id="250036E5FF8BFF900CB948602F90FAF8" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
<emphasis id="17CBEAF7FF8BFF900CB948602F37FA93" bold="true" box="[197,366,1309,1335]" pageId="12" pageNumber="13">Distribution:</emphasis>
<taxonomicName id="E2BF4D66FF8BFF900D0B48602CFDFA92" box="[375,676,1309,1334]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
<emphasis id="17CBEAF7FF8BFF900D0B48602CFDFA92" box="[375,676,1309,1334]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
</taxonomicName>
is only known from the Sangkulirang Karst, East Kalimantan (
<collectingCountry id="5DA87675FF8BFF900CE3483F2F4CFAF8" box="[159,277,1346,1372]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
) (
<figureCitation id="BD842A60FF8BFF900D50483F2FE5FAF8" box="[300,444,1346,1372]" captionStart="FIGURES 41 42" captionStartId="16.[151,272,1727,1751]" captionTargetBox="[200,1385,221,1687]" captionTargetId="figure@16.[191,1395,194,1706]" captionTargetPageId="16" captionText="FIGURES 41 42. Map showing the distribution of Sarax species: 41. The distribution in Southeast Asia, except for Sarax species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan." httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="12" pageNumber="13">Figs. 41, 42</figureCitation>
).
</paragraph>
<paragraph id="250036E5FF8BFF900CB9481A2DFAF9B3" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
<emphasis id="17CBEAF7FF8BFF900CB9481A2FC8FA25" bold="true" box="[197,401,1383,1409]" pageId="12" pageNumber="13">Natural history:</emphasis>
This species was collected from caves situated in three different limestone hills on the northern part of the Sangkulirang Karst formation, including Tabalar Ulu which is a small limestone area with several horizontal underground rivers, the Tebo Area where an isolated karst lake is located, and Gua Ampanas.
<taxonomicName id="E2BF4D66FF8BFF900D6948A52C6FFA55" box="[277,566,1496,1521]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
<emphasis id="17CBEAF7FF8BFF900D6948A52C6FFA55" box="[277,566,1496,1521]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
</taxonomicName>
was collected from the same caves as
<taxonomicName id="E2BF4D66FF8BFF90087C48A52F4EF9B3" authority="Golovatch et al. 2009" authorityName="Golovatch et al." authorityYear="2009" class="Diplopoda" family="Glyphiulidae" genus="Plusioglyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="17CBEAF7FF8BFF90087C48A52B43FA55" box="[1024,1306,1496,1521]" italics="true" pageId="12" pageNumber="13">Plusioglyphiulus similis</emphasis>
<bibRefCitation id="412E4B14FF8BFF90095E48A52F4EF9B3" author="Golovatch" pageId="12" pageNumber="13" refString="Golovatch, S. I., Geoffroy, J. J., Mauries, J. P. &amp; Van den Spiegel, D. (2009) Review of the millipede genus Plusioglyphiulus Silvestri, 1923, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema, 31 (1), 71 - 116." type="journal article" year="2009">
Golovatch
<emphasis id="17CBEAF7FF8BFF900CEB48802E93F9B2" box="[151,202,1533,1558]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2009
</bibRefCitation>
</taxonomicName>
, a recently described millipede (
<bibRefCitation id="412E4B14FF8BFF900EED48802DCFF9B3" author="Golovatch" box="[657,918,1533,1559]" pageId="12" pageNumber="13" refString="Golovatch, S. I., Geoffroy, J. J., Mauries, J. P. &amp; Van den Spiegel, D. (2009) Review of the millipede genus Plusioglyphiulus Silvestri, 1923, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema, 31 (1), 71 - 116." type="journal article" year="2009">
Golovatch
<emphasis id="17CBEAF7FF8BFF900F6948802D11F9B2" box="[789,840,1533,1558]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DA5656EFF8BFF900CB94B5F2F74F908" pageId="12" pageNumber="13" type="discussion">
<paragraph id="250036E5FF8BFF900CB94B5F2F74F908" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
<emphasis id="17CBEAF7FF8BFF900CB94B5F2F18F998" bold="true" box="[197,321,1570,1596]" pageId="12" pageNumber="13">Remarks:</emphasis>
<taxonomicName id="E2BF4D66FF8BFF900D354B5E2C34F998" box="[329,621,1571,1596]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
<emphasis id="17CBEAF7FF8BFF900D354B5E2C34F998" box="[329,621,1571,1596]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="E2BF4D66FF8BFF900F774B5E2D9EF998" box="[779,967,1571,1596]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="17CBEAF7FF8BFF900F774B5E2D9EF998" box="[779,967,1571,1596]" italics="true" pageId="12" pageNumber="13">S. sarawakensis</emphasis>
</taxonomicName>
by the number of trichobothria and the size of the spines on the pedipalpal tarsus. It differs from
<taxonomicName id="E2BF4D66FF8BFF900F4F4B352DF3F9C5" box="[819,938,1608,1633]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mardua">
<emphasis id="17CBEAF7FF8BFF900F4F4B352DF3F9C5" box="[819,938,1608,1633]" italics="true" pageId="12" pageNumber="13">S. mardua</emphasis>
</taxonomicName>
in having smaller spines on the pedipalpal tarsus, the lack of distinct sexual dimorphism and can be easily distinguished by the high and black median eye tubercle.
</paragraph>
</subSubSection>
</treatment>
</document>