314 lines
38 KiB
XML
314 lines
38 KiB
XML
<document id="561D19F2BC7BE6ADDADCD9F35BD6881E" ID-DOI="10.5281/zenodo.197810" ID-GBIF-Dataset="3fed616e-ea51-496b-b522-145b87775b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="197810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460279155734" checkinUser="plazi" docAuthor="Rahmadi, Cahyo, Harvey, Mark S. & Kojima, Jun-Ichi" docDate="2010" docId="AD1687F3FF8DFF900CEB49FA2F74F908" docLanguage="en" docName="zt02612p021.pdf" docOrigin="Zootaxa 2612" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sarax sangkulirangensis Rahmadi, Harvey & Kojima, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="512FFF8BFF87FF9C0C7C4D7D2E59FFA4" masterDocTitle="Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island" masterLastPageNumber="21" masterPageNumber="1" pageNumber="11" updateTime="1698249151004" updateUser="plazi">
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<mods:mods id="2F59C1B3FD7FA2B5113FDB1BF9E2D95D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1B2698CA670BE3B60558BD265901C2C2">
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<mods:title id="20F3190D159E2C7A1DBBD661B46A2FEF">Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island</mods:title>
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</mods:titleInfo>
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<mods:name id="BBCF45642F749D5938D1DB64457BD403" type="personal">
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<mods:role id="14B745F6BC4E1CB70B4073A68A52714E">
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<mods:roleTerm id="BCB80A9D5BD86AFEB8BA8AA26AA35731">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="4158E34907CA1907D1657447D0A4592C">Rahmadi, Cahyo</mods:namePart>
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</mods:name>
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<mods:name id="EF439E18CA3C92042B675F9F7B0F2063" type="personal">
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<mods:role id="F837233F33C7AC51C5323EE8A485DF0A">
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<mods:roleTerm id="FA9FB70568727ED9FAB340839BE01B91">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9A70027ACD7A8AE1DC70307A903E7F7D">Harvey, Mark S.</mods:namePart>
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</mods:name>
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<mods:name id="741DC3A6BD7BA46A653A9E83A2C55444" type="personal">
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<mods:role id="33C38E4F2B8F6579A00A70BCC77D9E97">
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<mods:roleTerm id="03E456A3C95963008D209F891DB296F5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D84B9759171BCC33BA8433492FC49472">Kojima, Jun-Ichi</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="52F2C51AC0559779117740DCF94E5D08">text</mods:typeOfResource>
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<mods:relatedItem id="2209694ACA79E2554F848E20AA1234E1" type="host">
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<mods:titleInfo id="BF71AE7766BDB3EBFF2DBBD7DB12F46E">
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<mods:title id="6B49610AAB1FE403896F4EBB52BBE341">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1465670D121182AA2696B594841D13EE">
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<mods:date id="A2D17A0C636DBED70A1C76A1AC798377">2010</mods:date>
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||
<mods:detail id="C1A97E7494A9F4B0228A02AAD1BF4D0A" type="volume">
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||
<mods:number id="D6F12BF341643E5A2B9B4882EBADD906">2612</mods:number>
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</mods:detail>
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<mods:extent id="5140DAB098C89C6A9E1207E430B30C94" unit="page">
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<mods:start id="D4583176FF5A8D653E0A0ED3D6D42553">1</mods:start>
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<mods:end id="17022112302CB5E8A5AC010598101AD9">21</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:classification id="75CD11895B9E0D632F8B04F0F1321607">journal article</mods:classification>
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<mods:identifier id="578F65E69D0ADD771125B68760ECCC7B" type="DOI">10.5281/zenodo.197810</mods:identifier>
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<mods:identifier id="81AD83C62415C4784B3C0BC0E4B25498" type="GBIF-Dataset">3fed616e-ea51-496b-b522-145b87775b5a</mods:identifier>
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<mods:identifier id="63A35BDF9983D59253016B678FB754EB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="324609239AC2E38DFC5412E35FF847F4" type="Zenodo-Dep">197810</mods:identifier>
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</mods:mods>
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<treatment id="AD1687F3FF8DFF900CEB49FA2F74F908" ID-DOI="http://doi.org/10.5281/zenodo.6207615" ID-GBIF-Taxon="119396509" ID-Zenodo-Dep="6207615" LSID="urn:lsid:plazi:treatment:AD1687F3FF8DFF900CEB49FA2F74F908" httpUri="http://treatment.plazi.org/id/AD1687F3FF8DFF900CEB49FA2F74F908" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection id="6DA5656EFF8DFF960CEB49FA2FD5FB60" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="250036E5FF8DFF960CEB49FA2C71FB05" blockId="10.[151,552,1159,1220]" box="[151,552,1159,1185]" pageId="10" pageNumber="11">
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<heading id="7E488189FF8DFF960CEB49FA2C71FB05" bold="true" box="[151,552,1159,1185]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="17CBEAF7FF8DFF960CEB49FA2C71FB05" bold="true" box="[151,552,1159,1185]" pageId="10" pageNumber="11">
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<taxonomicName id="E2BF4D66FF8DFF960CEB49FA2F9AFB05" ID-CoL="6XMF7" box="[151,451,1159,1185]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sangkulirangensis" status="sp. nov.">
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<emphasis id="17CBEAF7FF8DFF960CEB49FA2F9AFB05" bold="true" box="[151,451,1159,1185]" italics="true" pageId="10" pageNumber="11">Sarax sangkulirangensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0CF8578CFF8DFF960DB749FA2C71FB05" box="[459,552,1159,1185]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="250036E5FF8DFF960CEB49D72FD5FB60" blockId="10.[151,552,1159,1220]" box="[151,396,1194,1220]" pageId="10" pageNumber="11">
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(
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<figureCitation id="BD842A60FF8DFF960CE349D72F74FB60" box="[159,301,1194,1220]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="10" pageNumber="11">Figs. 21–25</figureCitation>
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,
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<figureCitation id="BD842A60FF8DFF960D4649D72FDDFB60" box="[314,388,1194,1220]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="10" pageNumber="11">37–38</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF8DFF960CEB49882B1DF9B0" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="250036E5FF8DFF960CEB49882B1DF9B0" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
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<emphasis id="17CBEAF7FF8DFF960CEB49882FCBFAAB" bold="true" box="[151,402,1269,1295]" pageId="10" pageNumber="11">Material examined:</emphasis>
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Female
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<typeStatus id="FA048847FF8DFF960D8749882C3AFAAB" box="[507,611,1269,1295]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="43AEAE20FF8DFF960E0849882CE0FAAB" box="[628,697,1269,1295]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="11">MZB</collectionCode>
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.Ambl.070),
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<collectingCountry id="5DA87675FF8DFF960F2C49882DAEFAAB" box="[848,1015,1269,1295]" name="Indonesia" pageId="10" pageNumber="11">INDONESIA</collectingCountry>
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:
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<emphasis id="17CBEAF7FF8DFF96087E49882A8EFAAA" box="[1026,1239,1269,1294]" italics="true" pageId="10" pageNumber="11">East Kalimantan:</emphasis>
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Berau Regency, District Tabalar Ulu, Gua Ke 4, GPS coordinates
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<geoCoordinate id="408B5022FF8DFF960E8848672DC5FA90" box="[756,924,1306,1332]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.4703417">1°28'13.23"N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF8DFF960FD648672A30FA90" box="[938,1129,1306,1332]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.65782">117°39'28.14"E</geoCoordinate>
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(approximate),
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<date id="51011025FF8DFF96095148672E8CFAFE" pageId="10" pageNumber="11" value="2004-08-09">9 August 2004</date>
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, C. Rahmadi, ovigerous.
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<typeStatus id="FA048847FF8DFF960E7E483D2C20FAFE" box="[514,633,1344,1370]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="33B9FD6CFF8DFF960EF9483D2C8CFAFE" box="[645,725,1344,1370]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
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(
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<collectionCode id="43AEAE20FF8DFF960E9A483D2D72FAFE" box="[742,811,1344,1370]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="11">MZB</collectionCode>
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.Ambl.150), Berau Regency, Kelai District, Merapun Village, near Tebo Lake, Gua Danum Tengen, GPS coordinates
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<geoCoordinate id="408B5022FF8DFF960FF048182A77FADB" box="[908,1070,1381,1407]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.5245833">1°31'28.50"N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF8DFF96084748182ABFFADB" box="[1083,1254,1381,1407]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.36803">117°22'4.90"E</geoCoordinate>
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(approximate),
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<date id="51011025FF8DFF960CEB48F72F25FA00" box="[151,380,1418,1444]" pageId="10" pageNumber="11" value="2004-09-02">2 September 2004</date>
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, Y.R. Suhardjono;
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<specimenCount id="33B9FD6CFF8DFF960E1B48F72CBAFA00" box="[615,739,1418,1444]" pageId="10" pageNumber="11" type="juvenile">1 juvenile</specimenCount>
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(MNHN-Am.10), Berau Regency, Tabalar District, Gua Louwading, Tabalar Ulu,
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<date id="51011025FF8DFF960DB748CD2CD5FA6E" box="[459,652,1456,1482]" pageId="10" pageNumber="11" value="2004-08-11">11 August 2004</date>
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, L. Deharveng & A. Bedos (KAL-059);
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<specimenCount id="33B9FD6CFF8DFF96080F48CD2A9DFA6E" box="[1139,1220,1456,1482]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
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(MNHN-Am.11), Kutai Timur Regency, Sangkulirang District, Pengadan Village, Gua Ampanas, GPS coordinates
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<geoCoordinate id="408B5022FF8DFF960CEB48872F60F9B0" box="[151,313,1530,1556]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="1.2004056">1°12'01.46"N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF8DFF960D3948872FA7F9B0" box="[325,510,1530,1556]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="117.73432">117°44'03.58"E</geoCoordinate>
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(approximate),
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<date id="51011025FF8DFF960ECA48872D28F9B0" box="[694,881,1530,1556]" pageId="10" pageNumber="11" value="2004-08-18">18 August 2004</date>
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, L. Deharveng & A. Bedos (KAL-113).
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF8DFF960CB94B622F0DF90E" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="250036E5FF8DFF960CB94B622F0DF90E" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
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<emphasis id="17CBEAF7FF8DFF960CB94B622F12F99D" bold="true" box="[197,331,1567,1593]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
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<taxonomicName id="E2BF4D66FF8DFF960D284B5D2C27F99D" box="[340,638,1568,1593]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sangkulirangensis">
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<emphasis id="17CBEAF7FF8DFF960D284B5D2C27F99D" box="[340,638,1568,1593]" italics="true" pageId="10" pageNumber="11">Sarax sangkulirangensis</emphasis>
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</taxonomicName>
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is relatively small sized (adult body length
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<quantity id="E2479B00FF8DFF9608EB4B5D2B77F99E" box="[1175,1326,1567,1594]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.8" metricValueMin="5.9" pageId="10" pageNumber="11" unit="mm" value="7.85" valueMax="9.8" valueMin="5.9">
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5.9
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<emphasis id="17CBEAF7FF8DFF9608C14B622A95F99D" bold="true" box="[1213,1228,1567,1593]" italics="true" pageId="10" pageNumber="11">–</emphasis>
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9.8 mm
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</quantity>
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), lacking sexual dimorphism, the pedipalpal tarsus has two spines with the proximal spine about half as long as distal spine, and tibia of leg IV has 19 trichobothria with
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<emphasis id="17CBEAF7FF8DFF960E904B162D5AF920" box="[748,771,1643,1668]" italics="true" pageId="10" pageNumber="11">bt</emphasis>
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on the middle of fourth basitibial segment, and
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<emphasis id="17CBEAF7FF8DFF96094F4B162B16F920" box="[1331,1359,1643,1668]" italics="true" pageId="10" pageNumber="11">bc</emphasis>
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closer to
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<emphasis id="17CBEAF7FF8DFF960CC94BED2E95F90D" box="[181,204,1680,1705]" italics="true" pageId="10" pageNumber="11">bf</emphasis>
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than to
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<emphasis id="17CBEAF7FF8DFF960D574BED2F14F90D" box="[299,333,1680,1705]" italics="true" pageId="10" pageNumber="11">sbf</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF8DFF900CB94BC82F9CFB48" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
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<paragraph id="250036E5FF8DFF960CB94BC82CB8F8BE" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
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<emphasis id="17CBEAF7FF8DFF960CB94BC82F3BF96B" bold="true" box="[197,354,1717,1743]" pageId="10" pageNumber="11">Description:</emphasis>
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<emphasis id="17CBEAF7FF8DFF960D164BC82F97F96A" box="[362,462,1717,1742]" italics="true" pageId="10" pageNumber="11">Female:</emphasis>
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Color in alcohol: Carapace brown, centrally with black marks, flange yellowishbrown; pedipalp yellowish-brown except for tarsus reddish-brown and spine yellow; legs yellowish-brown, but patella dark brown; abdomen greenish-brown.
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</paragraph>
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<paragraph id="250036E5FF8DFF960CB94A582B73F870" blockId="10.[151,1438,1269,2004]" pageId="10" pageNumber="11">
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Carapace (
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<figureCitation id="BD842A60FF8DFF960D3F4A582FC5F89B" box="[323,412,1829,1855]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="10" pageNumber="11">Fig. 21</figureCitation>
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): Width about 1.4
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<emphasis id="17CBEAF7FF8DFF960E0D4A582CD9F89B" bold="true" box="[625,640,1829,1855]" italics="true" pageId="10" pageNumber="11">–</emphasis>
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1.7 times its length; surface granulate, without setiferous tubercles; median sulcus deep in posterior one-third of the carapace; 3 pairs lateral sulci present; flange wide and bent upward; anterior margin slightly rounded, with 6 frontal setae. Median eye tubercle black, well-developed and high, without apical setae, slightly emarginate antero-medially to form heart-shape and median eyes facing antero-laterally; lateral eyes close to lateral margin of carapace, well-developed with black tapetum.
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</paragraph>
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<caption id="71C0666DFF8CFF970CEB49B82FE7FA99" httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12" targetBox="[204,1355,266,1164]" targetPageId="11">
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<paragraph id="250036E5FF8CFF970CEB49B82FE7FA99" blockId="11.[151,1436,1221,1341]" pageId="11" pageNumber="12">
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<emphasis id="17CBEAF7FF8CFF970CEB49B82F06FB79" bold="true" box="[151,351,1221,1245]" pageId="11" pageNumber="12">FIGURES 21–25</emphasis>
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.
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<taxonomicName id="E2BF4D66FF8CFF970D1249BB2CDCFB79" box="[366,645,1222,1245]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sangkulirangensis" status="sp. nov.">
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<emphasis id="17CBEAF7FF8CFF970D1249BB2CDCFB79" box="[366,645,1222,1245]" italics="true" pageId="11" pageNumber="12">Sarax sangkulirangensis</emphasis>
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</taxonomicName>
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<emphasis id="17CBEAF7FF8CFF970EF049B82CBEFB79" bold="true" box="[652,743,1221,1245]" pageId="11" pageNumber="12">
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<taxonomicNameLabel id="0CF8578CFF8CFF970EF049B82CBEFB79" box="[652,743,1221,1245]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB.Ambl.150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB.Ambl.150).
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</paragraph>
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</caption>
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<paragraph id="250036E5FF8CFF970CB948162B72FA51" blockId="11.[151,1437,1387,2011]" pageId="11" pageNumber="12">
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Chelicera (
|
||
<figureCitation id="BD842A60FF8CFF970D3A48162FC6FA21" box="[326,415,1387,1413]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
|
||
): Dorsal surface smooth, with 1 fine frontal seta and several fine setae. Basal segment with 4 teeth: lower-most tooth largest, upper-most tooth bicuspid, with upper cusp larger than lower; inner surface with several setae arranged in vertical row; outer surface with small tooth opposite bicuspid tooth, ventrally with several setae near proximal margin. Movable article with 5 teeth almost equal in size.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8CFF970CB94B7C2CA9F9C1" blockId="11.[151,1437,1387,2011]" pageId="11" pageNumber="12">
|
||
Sternum (
|
||
<figureCitation id="BD842A60FF8CFF970D474B7C2FCCF9BF" box="[315,405,1537,1563]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 23</figureCitation>
|
||
): First sternite (= tritosternum) elongate, with paired apical and several setae. Second and third sternites rounded and slightly rounded, respectively with 3 and 2 setae in addition to paired apical setae. Fourth sternite (= metasternum) with 5 setae.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8CFF900CB94B0C2A82FEE3" blockId="11.[151,1437,1387,2011]" lastBlockId="12.[151,1437,152,1708]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Pedipalp (
|
||
<figureCitation id="BD842A60FF8CFF970D3F4B0C2FFBF92F" box="[323,418,1649,1675]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
|
||
): Short and stout. Trochanter on antero-dorsal margin with 4 setiferous tubercles, without median spine; antero-ventral margin with ventro-anterior apophysis equipped with several setiferous tubercles on distal margin of trochanter. Femur: antero-dorsal margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF8CFF970CEB4B9D2F0DF95E" bold="true" box="[151,340,1760,1786]" pageId="11" pageNumber="12">F1>F2>F3>F4)</emphasis>
|
||
, 1 minor spine, several setiferous tubercles and small tubercles; antero-ventral margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF8CFF970DEE4A782C2AF8BB" bold="true" box="[402,627,1797,1823]" pageId="11" pageNumber="12">FI>FII>FIII>FIV</emphasis>
|
||
), 2 minor spines and small tubercles; 1 spine present dorsally of
|
||
<emphasis id="17CBEAF7FF8CFF9709024A782BC2F8BB" bold="true" box="[1406,1435,1797,1823]" pageId="11" pageNumber="12">FI</emphasis>
|
||
and as long as three-quarters length of
|
||
<emphasis id="17CBEAF7FF8CFF970E004A562CFAF8E1" bold="true" box="[636,675,1835,1861]" pageId="11" pageNumber="12">FI,</emphasis>
|
||
minor spine present between
|
||
<emphasis id="17CBEAF7FF8CFF9708644A562ADAF8E1" bold="true" box="[1048,1155,1835,1861]" pageId="11" pageNumber="12">FII -FIII</emphasis>
|
||
. Patella: antero-dorsal margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF8CFF970E284A2D2D49F8CE" bold="true" box="[596,784,1872,1898]" pageId="11" pageNumber="12">P1>P2>P3>P4</emphasis>
|
||
), clumped distally, 1 minor spine, several setiferous tubercles and small tubercles; 1 minor spine as long as half of
|
||
<emphasis id="17CBEAF7FF8CFF970FF94A082DFFF82B" bold="true" box="[901,934,1909,1935]" pageId="11" pageNumber="12">P1</emphasis>
|
||
presents between
|
||
<emphasis id="17CBEAF7FF8CFF9708FF4A082AFAF82B" bold="true" box="[1155,1187,1909,1935]" pageId="11" pageNumber="12">P1</emphasis>
|
||
and distal margin of patella, 1 spine presents between
|
||
<emphasis id="17CBEAF7FF8CFF970E5E4AE62C1AF811" bold="true" box="[546,579,1947,1973]" pageId="11" pageNumber="12">P4</emphasis>
|
||
and proximal margin; antero-ventral margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF8CFF970CEB4ABD2F2FF87E" bold="true" box="[151,374,1984,2010]" pageId="11" pageNumber="12">PII>PI>PIII>PIV</emphasis>
|
||
), several setiferous tubercles and small tubercles present. Tibia with outer surface smooth, with spines as follows: 2 major spines on antero-dorsal margin, proximal spine about half as long as distal one; 1 major spine on antero-ventral margin close to distal margin of tibia. Tarsus completely divided (claw clearly demarcated by articulation), with 2 spines on antero-dorsal margin separated from each other by about three times basal diameter of proximal spine: proximal spine shortest, length of distal spine about twice as long as proximal one; cleaning organ ventrally with about 30 modified hairs; apotele present.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8BFF900CB94C2F2F16FDD6" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
Legs (
|
||
<figureCitation id="BD842A60FF8BFF900D734C2F2F30FEC8" box="[271,361,338,364]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
|
||
): Femora of legs I
|
||
<emphasis id="17CBEAF7FF8BFF900E384C2F2C0AFEC8" bold="true" box="[580,595,338,364]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
IV with small tubercles bearing setae. Tibia and tarsus of leg I with 23 and 41 segments, respectively; tibiae of legs
|
||
<date id="51011025FF8BFF900ED34C052D6BFE36" box="[687,818,376,402]" pageId="12" pageNumber="13">II and III 2</date>
|
||
-segmented; basitibia of leg
|
||
<date id="51011025FF8BFF9008F94C052AE5FE36" box="[1157,1212,376,402]" pageId="12" pageNumber="13">IV 4</date>
|
||
-segmented, fourth segment with 1 trichobothrium (value in parentheses: ratio of the distance from the trichobothrium to the proximal margin of the segment against the length of the segment),
|
||
<emphasis id="17CBEAF7FF8BFF900FB64CBF2DBBFE7F" box="[970,994,450,475]" italics="true" pageId="12" pageNumber="13">bt</emphasis>
|
||
(0.52); distitibiae of legs II
|
||
<emphasis id="17CBEAF7FF8BFF9009534CBF2B67FE78" bold="true" box="[1327,1342,450,476]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
IV each with 18 trichobothria
|
||
<emphasis id="17CBEAF7FF8BFF900DE94C952FE3FDA5" box="[405,442,488,513]" italics="true" pageId="12" pageNumber="13">bf1</emphasis>
|
||
(0.07),
|
||
<emphasis id="17CBEAF7FF8BFF900E6B4C952C60FDA5" box="[535,569,488,513]" italics="true" pageId="12" pageNumber="13">sbf</emphasis>
|
||
(0.31),
|
||
<emphasis id="17CBEAF7FF8BFF900EEA4C952CE8FDA5" box="[662,689,488,513]" italics="true" pageId="12" pageNumber="13">bc</emphasis>
|
||
(0.16),
|
||
<emphasis id="17CBEAF7FF8BFF900F724C952D7DFDA5" box="[782,804,488,513]" italics="true" pageId="12" pageNumber="13">bt</emphasis>
|
||
close to distal margin of basitibial fourth segment,
|
||
<emphasis id="17CBEAF7FF8BFF9009FC4C952BC2FDA5" box="[1408,1435,488,513]" italics="true" pageId="12" pageNumber="13">bc</emphasis>
|
||
close to
|
||
<emphasis id="17CBEAF7FF8BFF900D7C4F702F7DFD82" box="[256,292,525,550]" italics="true" pageId="12" pageNumber="13">sbf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF8BFF900DF04F702FFDFD82" box="[396,420,525,550]" italics="true" pageId="12" pageNumber="13">bf</emphasis>
|
||
. Tarsi of legs
|
||
<date id="51011025FF8BFF900E244F702CEDFD83" box="[600,692,525,551]" pageId="12" pageNumber="13">
|
||
II
|
||
<emphasis id="17CBEAF7FF8BFF900E104F702C22FD83" bold="true" box="[620,635,525,551]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
IV 4
|
||
</date>
|
||
-segmented; first segment more than length of subsequent 3 segments combined; second segment with light-yellow transverse line; fourth segment without oblique slit; pulvilli present.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8BFF900CB94F002C4AFD18" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">Genitalia: Gonopods covered ventrally with genital operculum, with posterior margin concave. Gonopods paired, soft, white and tube-like.</paragraph>
|
||
<paragraph id="250036E5FF8BFF900CB94FB52A7BFCF6" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
<emphasis id="17CBEAF7FF8BFF900CB94FB52F57FD45" box="[197,270,712,737]" italics="true" pageId="12" pageNumber="13">Male:</emphasis>
|
||
Similar to female pedipalpal femur with 3 major spines (
|
||
<emphasis id="17CBEAF7FF8BFF900FA84FBA2A3CFD45" bold="true" box="[980,1125,711,737]" pageId="12" pageNumber="13">F1>F2>F3)</emphasis>
|
||
on antero-dorsal margin. Genitalia: covered ventrally by genital operculum, slightly concave on posterior margin; paired pointedprojections posteriorly, lateral lobes with brown band on basal lobe (
|
||
<figureCitation id="BD842A60FF8BFF900F9D4E6F2A64FC88" box="[993,1085,786,812]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="12" pageNumber="13">Fig. 38</figureCitation>
|
||
). In dorsal view, soft, paired brownish band running from anterior to posterior to median section (
|
||
<figureCitation id="BD842A60FF8BFF900FC04E452A4DFCF6" box="[956,1044,824,850]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="12" pageNumber="13">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8BFF900CB94E202F9CFB48" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
Measurements (in mm): male (n=2) [female (n=1)]; values for segments of the appendages are their lengths. Body length (excluding chelicera) 5.94
|
||
<emphasis id="17CBEAF7FF8BFF900EBD4EFF2C96FC38" bold="true" box="[705,719,898,924]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
7.01 [9.85]. Carapace: median length 2.35–2.65 [3.63]; width 3.44–3.82 [5.28]; median eyes to anterior margin of carapace 0.03–0.07 [0.07]; distance between lateral eyes 1.65–1.87 [2.55]; lateral eye to anterior margin of carapace 0.32–0.45 [0.50]; lateral eye to lateral margin of carapace 0.14–0.15 [0.37]. Pedipalps: trochanter 0.97–1.01 [1.32]; femur 1.96–2.14 [3.44]; patella 2.32–2.45 [4.00]; tibia 1.11–1.50 [1.13]; tarsus 1.22–1.39 [2.01]. Leg I: femur 6.48–7.62 [12.13]; patella 0.49–0.55 [0.71]. Leg II: femur 3.38–4.53 [7.10], patella 0.62–0.72 [1.03]; basitibia 2.55–3.67[6.17]; distitibia 1.67– 2.30 [3.51]; metatarsus + tarsus 1.13–1.56 [2.74]. Leg III: femur 4.06–5.53 [8.35]; patella 0.65–0.79 [1.16]; basitibia 3.33–4.83 [7.65]; distitibia 1.87–2.48 [3.70]; metatarsus+ tarsus 1.59–2.09 [2.89]. Leg IV: femur 3.60–4.81 [7.09]; patella 0.66–0.73 [0.58]; basitibia 3.42
|
||
<emphasis id="17CBEAF7FF8BFF900F4B49D02D1FFB63" bold="true" box="[823,838,1197,1223]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
5.38 [7.52]; distitibia 1.61
|
||
<emphasis id="17CBEAF7FF8BFF90080049D02AD2FB63" bold="true" box="[1148,1163,1197,1223]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
2.10 [3.33]; metatarsus + tarsus 1.80
|
||
<emphasis id="17CBEAF7FF8BFF900D5149AF2F65FB48" bold="true" box="[301,316,1234,1260]" italics="true" pageId="12" pageNumber="13">–</emphasis>
|
||
1.92 [2.97].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8BFF900CB9498A2A41FAB6" box="[197,1048,1271,1298]" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="250036E5FF8BFF900CB9498A2A41FAB6" blockId="12.[151,1437,152,1708]" box="[197,1048,1271,1298]" pageId="12" pageNumber="13">
|
||
<emphasis id="17CBEAF7FF8BFF900CB9498A2F0CFAB5" bold="true" box="[197,341,1271,1297]" pageId="12" pageNumber="13">Etymology:</emphasis>
|
||
The species name is derived from the name of
|
||
<typeStatus id="FA048847FF8BFF900FFC49852DEBFAB6" box="[896,946,1272,1298]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8BFF900CB948602DFAF9B3" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="250036E5FF8BFF900CB948602F90FAF8" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
<emphasis id="17CBEAF7FF8BFF900CB948602F37FA93" bold="true" box="[197,366,1309,1335]" pageId="12" pageNumber="13">Distribution:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF8BFF900D0B48602CFDFA92" box="[375,676,1309,1334]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF8BFF900D0B48602CFDFA92" box="[375,676,1309,1334]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Sangkulirang Karst, East Kalimantan (
|
||
<collectingCountry id="5DA87675FF8BFF900CE3483F2F4CFAF8" box="[159,277,1346,1372]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
|
||
) (
|
||
<figureCitation id="BD842A60FF8BFF900D50483F2FE5FAF8" box="[300,444,1346,1372]" captionStart="FIGURES 41 – 42" captionStartId="16.[151,272,1727,1751]" captionTargetBox="[200,1385,221,1687]" captionTargetId="figure@16.[191,1395,194,1706]" captionTargetPageId="16" captionText="FIGURES 41 – 42. Map showing the distribution of Sarax species: 41. The distribution in Southeast Asia, except for Sarax species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan." httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="12" pageNumber="13">Figs. 41, 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF8BFF900CB9481A2DFAF9B3" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
<emphasis id="17CBEAF7FF8BFF900CB9481A2FC8FA25" bold="true" box="[197,401,1383,1409]" pageId="12" pageNumber="13">Natural history:</emphasis>
|
||
This species was collected from caves situated in three different limestone hills on the northern part of the Sangkulirang Karst formation, including Tabalar Ulu which is a small limestone area with several horizontal underground rivers, the Tebo Area where an isolated karst lake is located, and Gua Ampanas.
|
||
<taxonomicName id="E2BF4D66FF8BFF900D6948A52C6FFA55" box="[277,566,1496,1521]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF8BFF900D6948A52C6FFA55" box="[277,566,1496,1521]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
was collected from the same caves as
|
||
<taxonomicName id="E2BF4D66FF8BFF90087C48A52F4EF9B3" authority="Golovatch et al. 2009" authorityName="Golovatch et al." authorityYear="2009" class="Diplopoda" family="Glyphiulidae" genus="Plusioglyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="17CBEAF7FF8BFF90087C48A52B43FA55" box="[1024,1306,1496,1521]" italics="true" pageId="12" pageNumber="13">Plusioglyphiulus similis</emphasis>
|
||
<bibRefCitation id="412E4B14FF8BFF90095E48A52F4EF9B3" author="Golovatch" pageId="12" pageNumber="13" refString="Golovatch, S. I., Geoffroy, J. J., Mauries, J. P. & Van den Spiegel, D. (2009) Review of the millipede genus Plusioglyphiulus Silvestri, 1923, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema, 31 (1), 71 - 116." type="journal article" year="2009">
|
||
Golovatch
|
||
<emphasis id="17CBEAF7FF8BFF900CEB48802E93F9B2" box="[151,202,1533,1558]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, a recently described millipede (
|
||
<bibRefCitation id="412E4B14FF8BFF900EED48802DCFF9B3" author="Golovatch" box="[657,918,1533,1559]" pageId="12" pageNumber="13" refString="Golovatch, S. I., Geoffroy, J. J., Mauries, J. P. & Van den Spiegel, D. (2009) Review of the millipede genus Plusioglyphiulus Silvestri, 1923, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema, 31 (1), 71 - 116." type="journal article" year="2009">
|
||
Golovatch
|
||
<emphasis id="17CBEAF7FF8BFF900F6948802D11F9B2" box="[789,840,1533,1558]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF8BFF900CB94B5F2F74F908" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="250036E5FF8BFF900CB94B5F2F74F908" blockId="12.[151,1437,152,1708]" pageId="12" pageNumber="13">
|
||
<emphasis id="17CBEAF7FF8BFF900CB94B5F2F18F998" bold="true" box="[197,321,1570,1596]" pageId="12" pageNumber="13">Remarks:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF8BFF900D354B5E2C34F998" box="[329,621,1571,1596]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF8BFF900D354B5E2C34F998" box="[329,621,1571,1596]" italics="true" pageId="12" pageNumber="13">Sarax sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="E2BF4D66FF8BFF900F774B5E2D9EF998" box="[779,967,1571,1596]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="17CBEAF7FF8BFF900F774B5E2D9EF998" box="[779,967,1571,1596]" italics="true" pageId="12" pageNumber="13">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
by the number of trichobothria and the size of the spines on the pedipalpal tarsus. It differs from
|
||
<taxonomicName id="E2BF4D66FF8BFF900F4F4B352DF3F9C5" box="[819,938,1608,1633]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mardua">
|
||
<emphasis id="17CBEAF7FF8BFF900F4F4B352DF3F9C5" box="[819,938,1608,1633]" italics="true" pageId="12" pageNumber="13">S. mardua</emphasis>
|
||
</taxonomicName>
|
||
in having smaller spines on the pedipalpal tarsus, the lack of distinct sexual dimorphism and can be easily distinguished by the high and black median eye tubercle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |