372 lines
39 KiB
XML
372 lines
39 KiB
XML
<document ID-DOI="10.1111/zoj.12333" ID-ISSN="0024-4082" ID-Zenodo-Dep="4720622" checkinTime="1619444012732" checkinUser="carolina" docAuthor="Prantoni, Alessandro Lívio, Wit, Pierre De & Erséus, Christer" docDate="2016" docId="7B5287F0DE49E46DE0843BA50618CA32" docLanguage="en" docName="ZoolJLinnSoc.176.485-510.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (3)" docSource="http://dx.doi.org/10.1111/zoj.12333" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Grania chilensis Prantoni & Wit & Erséus 2016, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="500" masterDocId="876BFF88DE44E462E349395E0740C819" masterDocTitle="First reports of Grania (Clitellata: Enchytraeidae) from Africa and South America: molecular phylogeny and descriptions of nine new species" masterLastPageNumber="510" masterPageNumber="485" pageNumber="498" updateTime="1673465143273" updateUser="felipe">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>First reports of Grania (Clitellata: Enchytraeidae) from Africa and South America: molecular phylogeny and descriptions of nine new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prantoni, Alessandro Lívio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wit, Pierre De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erséus, Christer</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>Zoological Journal of the Linnean Society</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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<mods:date>2016</mods:date>
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||
<mods:detail type="pubDate">
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<mods:number>2016-02-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>176</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>3</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>485</mods:start>
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<mods:end>510</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.1111/zoj.12333</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/zoj.12333</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4720622</mods:identifier>
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||
</mods:mods>
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||
<treatment LSID="urn:lsid:plazi:treatment:7B5287F0DE49E46DE0843BA50618CA32" httpUri="http://treatment.plazi.org/id/7B5287F0DE49E46DE0843BA50618CA32" lastPageId="15" lastPageNumber="500" pageId="13" pageNumber="498">
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||
<subSubSection box="[973,1294,763,784]" pageId="13" pageNumber="498" type="nomenclature">
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||
<paragraph blockId="13.[888,1378,763,848]" box="[973,1294,763,784]" pageId="13" pageNumber="498">
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<heading box="[973,1294,763,784]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="498" reason="2">
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<taxonomicName ID-CoL="3H7YP" ID-ENA="1702335" authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[973,1196,763,784]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="13" pageNumber="498" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">GRANIA CHILENSIS</taxonomicName>
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<taxonomicNameLabel box="[1203,1287,763,784]" pageId="13" pageNumber="498" rank="species">
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<emphasis bold="true" box="[1203,1235,763,784]" pageId="13" pageNumber="498">SP.</emphasis>
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<emphasis bold="true" box="[1242,1294,763,784]" pageId="13" pageNumber="498">NOV.</emphasis>
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="500" pageId="13" pageNumber="498" type="description">
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<paragraph blockId="13.[888,1378,763,848]" pageId="13" pageNumber="498">
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<figureCitation captionStart="Figure 8" captionStartId="14.[144,225,783,802]" captionTargetBox="[147,1421,198,750]" captionTargetId="figure-572@14.[144,1424,195,753]" captionTargetPageId="14" captionText="Figure 8. Grania chilensis sp. nov. A, chaetae. B, spermatheca. C, penial bulb and sperm funnel. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4720640" httpUri="https://zenodo.org/record/4720640/files/figure.png" pageId="13" pageNumber="498">
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FIGURE 8
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<taxonomicName authority="SP." authorityName="SP." box="[888,975,827,848]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="13" pageNumber="498" phylum="Annelida" rank="genus">
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<emphasis box="[888,975,827,848]" italics="true" pageId="13" pageNumber="498">GRANIA</emphasis>
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</taxonomicName>
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||
</figureCitation>
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||
SP.
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<collectingCountry box="[1020,1091,827,848]" name="Chile" pageId="13" pageNumber="498">CHILE</collectingCountry>
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2; DE WIT
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<emphasis box="[1225,1287,830,847]" italics="true" pageId="13" pageNumber="498">ET AL</emphasis>
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., 2011B
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</paragraph>
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||
<paragraph blockId="13.[826,1442,865,1071]" box="[826,925,865,886]" pageId="13" pageNumber="498">
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||
<emphasis box="[826,925,865,886]" italics="true" pageId="13" pageNumber="498">
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||
<typeStatus box="[826,925,865,886]" pageId="13" pageNumber="498">Holotype</typeStatus>
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||
</emphasis>
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||
</paragraph>
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||
<paragraph blockId="13.[826,1442,865,1071]" pageId="13" pageNumber="498">
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<materialsCitation accessionNumber="GU902190" collectingDate="2009-02-09" collectionCode="ZUEC" collectorName="Lower & P. De Wit & GenBank" country="Chile" latitude="-39.824165" location="Punta Loncoyen" longLatPrecision="19" longitude="-73.406944" municipality="Valdivia" pageId="13" pageNumber="498" specimenCode="CLI 13, PDW193" specimenCount="1" typeStatus="holotype">
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<collectionCode box="[826,899,896,917]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">ZUEC</collectionCode>
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||
<specimenCode box="[907,995,896,917]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 13</specimenCode>
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,
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<specimenCode box="[1014,1121,896,917]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW193</specimenCode>
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||
, whole-mounted, sexually mature specimen, with some segments amputated, from
|
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<location LSID="urn:lsid:plazi:treatment:7B5287F0DE49E46DE0843BA50618CA32:F624603DDE49E46FE0733AE304ABCBCB" box="[826,1003,957,978]" country="Chile" latitude="-39.824165" longLatPrecision="19" longitude="-73.406944" municipality="Valdivia" name="Punta Loncoyen" pageId="13" pageNumber="498">Punta Loncoyen</location>
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||
,
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<collectingMunicipality box="[1014,1104,957,979]" pageId="13" pageNumber="498">Valdivia</collectingMunicipality>
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||
,
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||
<collectingCountry box="[1114,1173,957,979]" name="Chile" pageId="13" pageNumber="498">Chile</collectingCountry>
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||
,
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||
<geoCoordinate box="[1184,1301,956,978]" degrees="39" direction="south" minutes="49" orientation="latitude" pageId="13" pageNumber="498" precision="15" seconds="27" value="-39.824165">39°49′27″S</geoCoordinate>
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||
,
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<geoCoordinate box="[1313,1439,956,979]" degrees="73" direction="west" minutes="24" orientation="longitude" pageId="13" pageNumber="498" precision="15" seconds="25" value="-73.406944">73°24′25″W</geoCoordinate>
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.
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<collectorName box="[826,896,988,1009]" pageId="13" pageNumber="498">Lower</collectorName>
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intertidal, sand among rocks. Collected by
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<collectorName pageId="13" pageNumber="498">P. De Wit</collectorName>
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||
,
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||
<date box="[880,1066,1018,1040]" pageId="13" pageNumber="498" value="2009-02-09">
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<collectingDate box="[880,1066,1018,1040]" pageId="13" pageNumber="498" value="2009-02-09">9 February 2009</collectingDate>
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</date>
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.
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||
<emphasis box="[1076,1122,1018,1039]" italics="true" pageId="13" pageNumber="498">COI</emphasis>
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||
barcode sequence,
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<collectorName box="[1336,1441,1018,1040]" pageId="13" pageNumber="498">GenBank</collectorName>
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||
acc. no.
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<accessionNumber box="[922,1046,1049,1070]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/GU902190" pageId="13" pageNumber="498">GU902190</accessionNumber>
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||
; for other sequences, see
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<tableCitation box="[1348,1433,1049,1071]" captionStart="Table 1" captionStartId="3.[31,95,393,414]" captionTargetPageId="3" captionText="Table 1. Specimens of the new species used for genetic analysis, with places of origin, voucher numbers, and GenBank accession numbers" pageId="13" pageNumber="498">Table 1</tableCitation>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph blockId="13.[826,1027,1111,1164]" box="[826,945,1111,1132]" pageId="13" pageNumber="498">
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||
<emphasis box="[826,945,1111,1132]" italics="true" pageId="13" pageNumber="498">Etymology</emphasis>
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||
</paragraph>
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||
<paragraph blockId="13.[826,1027,1111,1164]" box="[826,1027,1142,1164]" pageId="13" pageNumber="498">
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Named for
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<collectingCountry box="[959,1023,1142,1164]" name="Chile" pageId="13" pageNumber="498">Chile</collectingCountry>
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||
.
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||
</paragraph>
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||
<paragraph blockId="13.[826,1442,1206,1625]" box="[826,939,1206,1227]" pageId="13" pageNumber="498">
|
||
<emphasis box="[826,939,1206,1227]" italics="true" pageId="13" pageNumber="498">
|
||
<typeStatus box="[826,939,1206,1227]" pageId="13" pageNumber="498">Paratypes</typeStatus>
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||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="13.[826,1442,1206,1625]" pageId="13" pageNumber="498">
|
||
<materialsCitation collectingDate="2009-02-06" collectingDateMax="2009-02-16" collectingDateMin="2009-02-06" collectionCode="ZUEC" collectorName="Six & P. De Wit. & Puerto Aldea & Elqui" country="Chile" county="Concepcion" latitude="-36.633335" location="Caleta Tumbes" longLatPrecision="19" longitude="-73.090836" municipality="Talcahuano" pageId="13" pageNumber="498" specimenCode="CLI 14, CLI 16, PDW190, PDW191, PDW194, CLI 17, PDW185, CLI 18, CLI 19, PDW198, PDW199" specimenCount="1" stateProvince="Coquimbo" typeStatus="paratype">
|
||
<collectorName box="[826,862,1236,1258]" pageId="13" pageNumber="498">Six</collectorName>
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||
whole-mounted, sexually mature specimens, collected by
|
||
<collectorName box="[938,1056,1266,1288]" pageId="13" pageNumber="498">P. De Wit.</collectorName>
|
||
<collectionCode box="[1065,1136,1266,1287]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">ZUEC</collectionCode>
|
||
<specimenCode box="[1142,1221,1266,1287]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 14</specimenCode>
|
||
–
|
||
<specimenCode box="[1235,1319,1266,1288]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 16</specimenCode>
|
||
,
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||
<specimenCode box="[1333,1437,1266,1287]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW190</specimenCode>
|
||
,
|
||
<specimenCode box="[826,930,1297,1318]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW191</specimenCode>
|
||
,
|
||
<specimenCode box="[944,1047,1297,1318]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW194</specimenCode>
|
||
, with some segments amputated, from type locality.
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||
<collectionCode box="[1038,1108,1327,1348]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">ZUEC</collectionCode>
|
||
<specimenCode box="[1115,1196,1327,1348]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 17</specimenCode>
|
||
,
|
||
<specimenCode box="[1209,1313,1327,1349]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW185</specimenCode>
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||
, with some segments amputated, from
|
||
<collectorName box="[1150,1304,1358,1380]" pageId="13" pageNumber="498">Puerto Aldea</collectorName>
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||
,
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<collectingRegion box="[1320,1437,1358,1380]" country="Chile" name="Coquimbo" pageId="13" pageNumber="498">Coquimbo</collectingRegion>
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,
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||
<collectorName box="[826,891,1388,1410]" pageId="13" pageNumber="498">Elqui</collectorName>
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||
,
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||
<collectingCountry box="[903,966,1388,1410]" name="Chile" pageId="13" pageNumber="498">Chile</collectingCountry>
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||
,
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||
<geoCoordinate box="[979,1104,1388,1410]" degrees="30" direction="south" minutes="18" orientation="latitude" pageId="13" pageNumber="498" precision="15" seconds="19" value="-30.305277">30°18′19″S</geoCoordinate>
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||
,
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||
<geoCoordinate box="[1118,1251,1388,1410]" degrees="71" direction="west" minutes="39" orientation="longitude" pageId="13" pageNumber="498" precision="15" seconds="33" value="-71.659164">71°39′33″W</geoCoordinate>
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||
, intertidal, sand among rocks,
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||
<date box="[984,1176,1419,1441]" pageId="13" pageNumber="498" value="2009-02-06">
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||
<collectingDate box="[984,1176,1419,1441]" pageId="13" pageNumber="498" value="2009-02-06">6 February 2009</collectingDate>
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||
</date>
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||
.
|
||
<collectionCode box="[1187,1257,1419,1440]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">ZUEC</collectionCode>
|
||
<specimenCode box="[1263,1339,1419,1440]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 18</specimenCode>
|
||
–
|
||
<specimenCode box="[1354,1436,1419,1440]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">CLI 19</specimenCode>
|
||
,
|
||
<specimenCode box="[826,930,1450,1471]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW198</specimenCode>
|
||
,
|
||
<specimenCode box="[944,1048,1450,1471]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="13" pageNumber="498">PDW199</specimenCode>
|
||
, with some segments amputated, from
|
||
<location LSID="urn:lsid:plazi:treatment:7B5287F0DE49E46DE0843BA50618CA32:F624603DDE49E46FE0303C96036ACDC7" box="[889,1066,1480,1502]" country="Chile" county="Concepcion" latitude="-36.633335" longLatPrecision="19" longitude="-73.090836" municipality="Talcahuano" name="Caleta Tumbes" pageId="13" pageNumber="498" stateProvince="Coquimbo">Caleta Tumbes</location>
|
||
,
|
||
<collectingMunicipality box="[1080,1215,1480,1502]" pageId="13" pageNumber="498">Talcahuano</collectingMunicipality>
|
||
,
|
||
<collectingCounty box="[1228,1359,1480,1502]" pageId="13" pageNumber="498">Concepción</collectingCounty>
|
||
,
|
||
<collectingCountry box="[1374,1437,1480,1502]" name="Chile" pageId="13" pageNumber="498">Chile</collectingCountry>
|
||
,
|
||
<geoCoordinate box="[826,960,1510,1533]" degrees="36" direction="south" minutes="38" orientation="latitude" pageId="13" pageNumber="498" precision="15" seconds="00" value="-36.633335">36°38′00″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[979,1120,1510,1533]" degrees="73" direction="west" minutes="05" orientation="longitude" pageId="13" pageNumber="498" precision="15" seconds="27" value="-73.090836">73°05′27″W</geoCoordinate>
|
||
, lower intertidal, heterogeneous sand with organic material between boulders,
|
||
<date box="[887,1086,1572,1594]" pageId="13" pageNumber="498" value="2009-02-16">
|
||
<collectingDate box="[887,1086,1572,1594]" pageId="13" pageNumber="498" value="2009-02-16">16 February 2009</collectingDate>
|
||
</date>
|
||
. For
|
||
<emphasis box="[1143,1189,1572,1593]" italics="true" pageId="13" pageNumber="498">COI</emphasis>
|
||
barcodes of
|
||
<typeStatus box="[1326,1437,1573,1594]" pageId="13" pageNumber="498">paratypes</typeStatus>
|
||
, see
|
||
<tableCitation box="[872,956,1603,1625]" captionStart="Table 1" captionStartId="3.[31,95,393,414]" captionTargetPageId="3" captionText="Table 1. Specimens of the new species used for genetic analysis, with places of origin, voucher numbers, and GenBank accession numbers" pageId="13" pageNumber="498">Table 1</tableCitation>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="13.[826,1441,1666,1902]" box="[826,955,1666,1687]" pageId="13" pageNumber="498">
|
||
<emphasis box="[826,955,1666,1687]" italics="true" pageId="13" pageNumber="498">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="13.[826,1441,1666,1902]" lastBlockId="14.[144,759,901,1902]" lastPageId="14" lastPageNumber="499" pageId="13" pageNumber="498">
|
||
Body>
|
||
<quantity box="[914,1075,1696,1718]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="7.45" metricValueMin="3.85" pageId="13" pageNumber="498" unit="mm" value="5.65" valueMax="7.45" valueMin="3.85">3.85–7.45 mm</quantity>
|
||
long,> 26–40 segments (
|
||
<emphasis box="[1375,1390,1697,1718]" italics="true" pageId="13" pageNumber="498">n</emphasis>
|
||
= 6) (posterior ends used for DNA extractions),
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="13" pageNumber="498" unit="mm" value="0.15" valueMax="0.17" valueMin="0.13">0.13– 0.17 mm</quantity>
|
||
wide at segment V,
|
||
<quantity box="[1224,1397,1757,1779]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="13" pageNumber="498" unit="mm" value="0.15" valueMax="0.19" valueMin="0.11">0.11–0.19 mm</quantity>
|
||
at segment XII (
|
||
<emphasis box="[973,988,1789,1810]" italics="true" pageId="13" pageNumber="498">n</emphasis>
|
||
= 6). Prostomium conical, 55–80 μm long, 60–80 μm wide, epidermis not reduced at front tip, 7 μm thick (
|
||
<emphasis box="[899,914,1850,1871]" italics="true" pageId="13" pageNumber="498">n</emphasis>
|
||
= 6). Ventral chaetae from segment IV, lateral chaetae beginning in segment XVI (
|
||
<emphasis box="[1317,1332,1880,1901]" italics="true" pageId="13" pageNumber="498">n</emphasis>
|
||
= 1), in segment XVII (
|
||
<emphasis box="[328,343,901,922]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 3), in segment XVIII (
|
||
<emphasis box="[647,662,901,922]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 1), or segment XIX (
|
||
<emphasis box="[303,318,932,953]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 1). Chaetae (
|
||
<figureCitation box="[489,570,931,953]" captionStart="Figure 8" captionStartId="14.[144,225,783,802]" captionTargetBox="[147,1421,198,750]" captionTargetId="figure-572@14.[144,1424,195,753]" captionTargetPageId="14" captionText="Figure 8. Grania chilensis sp. nov. A, chaetae. B, spermatheca. C, penial bulb and sperm funnel. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4720640" httpUri="https://zenodo.org/record/4720640/files/figure.png" pageId="14" pageNumber="499">Fig. 8A</figureCitation>
|
||
) of uniform size, 44–59 μm long, shaft straight, 3.7–5.0 μm thick at midpoint, L-shaped, proximally bent into a foot with low instep, and with distinct heel only in preclitellar segments. Chaetal index (
|
||
<bibRefCitation author="Erseus C & Rota E" box="[394,625,1054,1076]" pageId="14" pageNumber="499" pagination="892 - 900" refId="ref17321" refString="Erseus C, Rota E. 2003. New findings and an overview of the oligochaetous Clitellata (Annelida) of the North Atlantic deep sea. Proceedings of the Biological Society of Washington 116: 892 - 900." type="journal article" year="2003">Rota & Erséus, 2003</bibRefCitation>
|
||
) 4.03 ± 0.52 (
|
||
<emphasis box="[152,167,1085,1106]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 5). Epidermal gland cells inconspicuous. Clitellum maximally 10–12 μm thick (
|
||
<emphasis box="[488,503,1115,1136]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 4), extending from segment XII to anterior half of segment XIII, consisting of transverse rows of granular gland cells interspersed with hyaline cells; with the latter, however, absent ventrally. Spermathecal pores in lateral lines, just posterior to 4/5. Male pores ventrolateral in middle of segment XII. Female pores not observed.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4720640" ID-Zenodo-Dep="4720640" httpUri="https://zenodo.org/record/4720640/files/figure.png" pageId="14" pageNumber="499" startId="14.[144,225,783,802]" targetBox="[147,1421,198,750]" targetPageId="14">
|
||
<paragraph blockId="14.[144,1421,782,831]" pageId="14" pageNumber="499">
|
||
<emphasis bold="true" box="[144,564,782,802]" pageId="14" pageNumber="499">
|
||
Figure 8.
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[263,467,782,802]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[263,467,782,802]" italics="true" pageId="14" pageNumber="499">Grania chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[475,564,783,802]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, chaetae. B, spermatheca. C, penial bulb and sperm funnel. See text for abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="14.[144,759,901,1902]" lastBlockId="14.[806,1422,900,1014]" pageId="14" pageNumber="499">
|
||
Brain posteriorly indented. Head organ (
|
||
<emphasis box="[635,698,1330,1351]" italics="true" pageId="14" pageNumber="499">sensu</emphasis>
|
||
<bibRefCitation author="Rota E & Erseus C" pageId="14" pageNumber="499" pagination="169 - 183" refId="ref18340" refString="Rota E, Erseus C. 1996. Six new species of Grania (Oligochaeta, Enchytraeidae) from the Ross Sea, Antarctica. Antarctic Science 2: 169 - 183." type="journal article" year="1996">Rota & Erséus, 1996</bibRefCitation>
|
||
) absent. Pharyngeal glands in segments IV–VI, dorsal lobes in segment IV (one pair), in segment V (one pair), and in segment VI (one pair), ventral lobes present in segment IV (one pair), in segment V (two pairs), and in segment VI (two pairs); glands not connected dorsally. Nephridia not observed. Chloragogen cells not observed. Dorsal blood vessel commencing in segments XVI–XVIII. Coelomocytes not observed. Sperm sac extending to segments XIII–XV, egg sac extending to segments XV– XVII. Sperm funnels about 1.5–2.5 times longer than wide (
|
||
<figureCitation box="[209,289,1696,1718]" captionStart="Figure 8" captionStartId="14.[144,225,783,802]" captionTargetBox="[147,1421,198,750]" captionTargetId="figure-572@14.[144,1424,195,753]" captionTargetPageId="14" captionText="Figure 8. Grania chilensis sp. nov. A, chaetae. B, spermatheca. C, penial bulb and sperm funnel. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4720640" httpUri="https://zenodo.org/record/4720640/files/figure.png" pageId="14" pageNumber="499">Fig. 8C</figureCitation>
|
||
). Vasa deferentia observed in segments XI– XIII, internally ciliated, coiled, 10 μm wide. Penial apparatus
|
||
<typeStatus box="[244,293,1758,1779]" pageId="14" pageNumber="499">type</typeStatus>
|
||
1 (
|
||
<emphasis box="[333,396,1758,1779]" italics="true" pageId="14" pageNumber="499">sensu</emphasis>
|
||
<bibRefCitation author="Coates KA" box="[407,561,1757,1779]" pageId="14" pageNumber="499" pagination="45 - 50" refId="ref16484" refString="Coates KA. 1984. Specific criteria in Grania (Oligochaeta, Enchytraeidae). Hydrobiologia 115: 45 - 50." type="journal article" year="1984">Coates, 1984</bibRefCitation>
|
||
), small, compact glandular bulb, 35–55 μm long, 37–50 μm wide (
|
||
<emphasis box="[688,703,1789,1810]" italics="true" pageId="14" pageNumber="499">n</emphasis>
|
||
= 6); stylet absent. Midventral copulatory gland (in segment XIV) present. Spermatheca attached to oesophagus in posterior half of segment V by narrow ental duct. Ampulla oval, 42–65 μm long, 30–42 μm wide (
|
||
<figureCitation box="[1327,1409,900,923]" captionStart="Figure 8" captionStartId="14.[144,225,783,802]" captionTargetBox="[147,1421,198,750]" captionTargetId="figure-572@14.[144,1424,195,753]" captionTargetPageId="14" captionText="Figure 8. Grania chilensis sp. nov. A, chaetae. B, spermatheca. C, penial bulb and sperm funnel. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4720640" httpUri="https://zenodo.org/record/4720640/files/figure.png" pageId="14" pageNumber="499">Fig. 8B</figureCitation>
|
||
). Sperm rings maximally 17 μm wide, but few. Ectal duct, 54–95 μm long, 13–30 μm wide at midcourse. No gland at spermathecal pore.
|
||
</paragraph>
|
||
<paragraph blockId="14.[806,1422,1053,1901]" box="[806,907,1053,1074]" pageId="14" pageNumber="499">
|
||
<emphasis box="[806,907,1053,1074]" italics="true" pageId="14" pageNumber="499">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="14.[806,1422,1053,1901]" pageId="14" pageNumber="499">
|
||
The compact and small penial bulb, the sperm funnels being 1.5–2.5 times longer than wide, the absence of penial stylets, the presence of a midventral copulatory gland in segment XIV, and the distribution of the lateral chaetae, starting from segments XVI–XIX, make
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[806,933,1237,1258]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[806,933,1237,1258]" italics="true" pageId="14" pageNumber="499">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[940,1022,1237,1258]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
reminiscent of
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[1195,1327,1237,1258]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="hinojosai" status="sp. nov.">
|
||
<emphasis box="[1195,1327,1237,1258]" italics="true" pageId="14" pageNumber="499">G. hinojosai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1333,1418,1237,1258]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; however,
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[924,1069,1268,1289]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="hinojosai" status="sp. nov.">
|
||
<emphasis box="[924,1069,1268,1289]" italics="true" pageId="14" pageNumber="499">G. hinojosai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1077,1166,1268,1289]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a spermathecal ampulla that is heart-shaped, a distinct chaetal size distribution (chaetae becoming larger towards the posterior end), and possesses a gland at the spermathecal pore, whereas in
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[1015,1146,1390,1411]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[1015,1146,1390,1411]" italics="true" pageId="14" pageNumber="499">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1153,1237,1390,1411]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the ampulla is ovoid, the chaetal size is uniform, and there are no glands at the spermathecal pore.
|
||
</paragraph>
|
||
<paragraph blockId="14.[806,1422,1053,1901]" lastBlockId="15.[164,779,197,555]" lastPageId="15" lastPageNumber="500" pageId="14" pageNumber="499">
|
||
Three species from South Pacific,
|
||
<taxonomicName authority="De Wit & Erseus, 2007" authorityName="De Wit & Erseus" authorityYear="2007" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis box="[1210,1384,1482,1503]" italics="true" pageId="14" pageNumber="499">Grania galbina</emphasis>
|
||
<bibRefCitation author="De Wit P & Erseus C" pageId="14" pageNumber="499" pagination="27 - 50" refId="ref16899" refString="De Wit P, Erseus C. 2007. Seven new species of Grania (Annelida: Clitellata: Enchytraeidae) from New Caledonia, South Pacific Ocean. Zootaxa 50: 27 - 50." type="journal article" year="2007">De Wit & Erséus, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="De Wit, Rota & Erseus, 2009" authorityName="De Wit, Rota & Erseus" authorityYear="2009" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="breviductus">
|
||
<emphasis box="[1045,1266,1512,1534]" italics="true" pageId="14" pageNumber="499">Grania breviductus</emphasis>
|
||
<bibRefCitation author="De Wit P & Rota E & Erseus C" pageId="14" pageNumber="499" pagination="16 - 38" refId="ref17021" refString="De Wit P, Rota E, Erseus C. 2009. Grania (Annelida: Clitellata: Enchytraeidae) of the Great Barrier Reef, Australia, including four new species and a re-description of Grania trichaeta Jamieson, 1977. Zootaxa 38: 16 - 38." type="journal article" year="2009">De Wit, Rota & Erséus, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="De Wit & Erseus" authorityYear="2007" box="[1045,1241,1543,1564]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="novacaledonia">
|
||
<emphasis box="[1045,1241,1543,1564]" italics="true" pageId="14" pageNumber="499">G. novacaledonia</emphasis>
|
||
</taxonomicName>
|
||
share a similar distribution of the lateral chaetae to that in
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[806,933,1604,1625]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[806,933,1604,1625]" italics="true" pageId="14" pageNumber="499">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[939,1021,1605,1626]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Moreover, as in
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[1201,1327,1604,1625]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[1201,1327,1604,1625]" italics="true" pageId="14" pageNumber="499">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1334,1418,1605,1626]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, a midventral copulatory gland is present in
|
||
<taxonomicName authorityName="De Wit, Rota & Erseus" authorityYear="2009" box="[1268,1421,1635,1656]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="breviductus">
|
||
<emphasis box="[1268,1421,1635,1656]" italics="true" pageId="14" pageNumber="499">G. breviductus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="De Wit & Erseus" authorityYear="2007" box="[861,1061,1665,1687]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="novacaledonia">
|
||
<emphasis box="[861,1061,1665,1687]" italics="true" pageId="14" pageNumber="499">G. novacaledonia</emphasis>
|
||
</taxonomicName>
|
||
too.
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[1126,1321,1665,1687]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[1126,1321,1665,1687]" italics="true" pageId="14" pageNumber="499">Grania chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1328,1417,1666,1687]" pageId="14" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, however, differs from all the species mentioned above by the short length of the sperm funnel; in the other three species these funnels are about four or five times longer than wide. Furthermore,
|
||
<taxonomicName authorityName="De Wit & Erseus" authorityYear="2007" box="[1163,1280,1788,1809]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis box="[1163,1280,1788,1809]" italics="true" pageId="14" pageNumber="499">G. galbina</emphasis>
|
||
</taxonomicName>
|
||
has its male pore surrounded by a large granular mass,
|
||
<taxonomicName authorityName="De Wit, Rota & Erseus" authorityYear="2009" box="[1266,1421,1818,1840]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="14" pageNumber="499" phylum="Annelida" rank="species" species="breviductus">
|
||
<emphasis box="[1266,1421,1818,1840]" italics="true" pageId="14" pageNumber="499">G. breviductus</emphasis>
|
||
</taxonomicName>
|
||
has the communication between the spermatheca and the oesophagus in the middle of segment V, and
|
||
<taxonomicName authorityName="De Wit & Erseus" authorityYear="2007" box="[164,356,197,218]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="novacaledonia">
|
||
<emphasis box="[164,356,197,218]" italics="true" pageId="15" pageNumber="500">G. novacaledonia</emphasis>
|
||
</taxonomicName>
|
||
possesses glands at the spermathecal pores, none of which is the case for
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[560,689,227,249]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[560,689,227,249]" italics="true" pageId="15" pageNumber="500">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[696,779,228,249]" pageId="15" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="15.[164,779,197,555]" pageId="15" pageNumber="500">
|
||
The ovoid spermathecal ampulla, the length of the sperm funnel, the distribution of the lateral chaetae, and the absence of a penial stylet make
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[164,295,350,371]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[164,295,350,371]" italics="true" pageId="15" pageNumber="500">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[302,387,350,371]" pageId="15" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see discussion above) similar to the Irish
|
||
<taxonomicName authority="Locke & Coates, 1998" authorityName="Locke & Coates" authorityYear="1998" box="[276,692,380,402]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="mira">
|
||
<emphasis box="[276,423,381,402]" italics="true" pageId="15" pageNumber="500">Grania mira</emphasis>
|
||
<bibRefCitation author="Locke JM & Coates KA" box="[434,692,380,402]" pageId="15" pageNumber="500" pagination="1101 - 1114" refId="ref17882" refString="Locke JM, Coates KA. 1998. A new species of Grania (Enchytraeidae, Clitellata, Annelida) and redescription of Grania pusilla from a rocky shore in SE Ireland. Journal of Natural History 32: 1101 - 1114." type="journal article" year="1998">Locke & Coates, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, but in addition to being geographically distant from this species,
|
||
<taxonomicName authority="Prantoni & Wit & Erséus, 2016" authorityName="Prantoni & Wit & Erséus" authorityYear="2016" box="[256,382,442,463]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="chilensis" status="sp. nov.">
|
||
<emphasis box="[256,382,442,463]" italics="true" pageId="15" pageNumber="500">G. chilensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[389,471,442,463]" pageId="15" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName authorityName="Locke & Coates" authorityYear="1998" box="[615,701,442,463]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="15" pageNumber="500" phylum="Annelida" rank="species" species="mira">
|
||
<emphasis box="[615,701,442,463]" italics="true" pageId="15" pageNumber="500">G. mira</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a midventral copulatory gland in segment XIV, and the lack of thickened muscles along the male ducts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |