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<document id="62F6A51DE4BFAD1B339F98BC0DA4E6D6" ID-DOI="10.11646/zootaxa.5519.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915949" ID-ZooBank="1EE4DD76-706A-45B7-87C9-6533710C3A72" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728586640252" checkinUser="plazi" docAuthor="Quevedo, Lucas, Capellari, Renato S. &amp; Lamas, Carlos José E." docDate="2024" docId="74455B11FFB592076083FD0D5A13FDFC" docLanguage="en" docName="zootaxa.5519.3.1.pdf" docOrigin="Zootaxa 5519 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achradocera femoralis Becker 1922" docType="treatment" docVersion="2" lastPageNumber="326" masterDocId="887C2369FFA3921E6014FF835F4EFFC1" masterDocTitle="Taxonomic revision of Achradocera Becker (Diptera: Dolichopodidae), with description of two new species" masterLastPageNumber="344" masterPageNumber="301" pageNumber="323" updateTime="1728587218636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DE3CF0F86FEFA3F86C18B3231FEB5AB2">Taxonomic revision of Achradocera Becker (Diptera: Dolichopodidae), with description of two new species</mods:title>
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<mods:namePart id="DD03FD435734162E6950E8B642F24404">Quevedo, Lucas</mods:namePart>
<mods:affiliation id="983FD734A3AB2595CE99576DDC4A4030">Universidade de São Paulo, Instituto de Biociências, Programa de Pós-Graduação em Zoologia. Rua do Matão, travessa 14, nº. 101, Cep: 05508 - 090, Cidade Universitária, São Paulo, SP, Brazil</mods:affiliation>
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<mods:namePart id="1183528EBE8107DD4A0DA7119C9CB22C">Capellari, Renato S.</mods:namePart>
<mods:affiliation id="451EC94B99C05AF53DEB31738F02409C">Instituto Federal de São Paulo. Campus Avaré. Av. Prof. Celso Ferreira da Silva, 1333 Jardim Europa II, 18707 - 150, Avaré, SP, Brazil</mods:affiliation>
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<mods:namePart id="3C7089548511E23EEE6A216A772C138D">Lamas, Carlos José E.</mods:namePart>
<mods:affiliation id="2CCA3B6C163B046663E1F34D2459F36D">Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481, 04263 - 000, Ipiranga, São Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="2890090131788478F8CA8699F6BB0C56">Zootaxa</mods:title>
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<mods:date id="C0B4C51BAB5D32DC780B438F00658851">2024</mods:date>
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<treatment id="74455B11FFB592076083FD0D5A13FDFC" LSID="urn:lsid:plazi:treatment:74455B11FFB592076083FD0D5A13FDFC" httpUri="http://treatment.plazi.org/id/74455B11FFB592076083FD0D5A13FDFC" lastPageId="25" lastPageNumber="326" pageId="22" pageNumber="323">
<subSubSection id="B4F6B98CFFB592086083FD0D5D1DFD69" box="[151,595,654,681]" pageId="22" pageNumber="323" type="nomenclature">
<paragraph id="FC53EA07FFB592086083FD0D5D1DFD69" blockId="22.[151,595,654,717]" box="[151,595,654,681]" pageId="22" pageNumber="323">
<heading id="A71B5D6BFFB592086083FD0D5D1DFD69" bold="true" box="[151,595,654,681]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
<taxonomicName id="3BEC9184FFB592086083FD0D5D1DFD69" ID-CoL="89TRS" authority="Becker, 1922" authorityName="Becker" authorityYear="1922" box="[151,595,654,681]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB592086083FD0D5D1DFD69" bold="true" box="[151,595,654,681]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB592086083FD0D5EE3FD68" bold="true" box="[151,429,654,681]" italics="true" pageId="22" pageNumber="323">Achradocera femoralis</emphasis>
<bibRefCitation id="987D97F6FFB5920861A0FD0C5D1DFD69" author="Becker, T." box="[436,595,654,681]" pageId="22" pageNumber="323" pagination="1 - 394" refId="ref26227" refString="Becker, T. (1922) Dipterologische Studien. Dolichopodidae. B. Nearktische und neotropische Region. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13 (1), 1 - 394. https: // doi. org / 10.5962 / bhl. title. 132893" type="journal article" year="1922">Becker, 1922</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB592086083FD305E1FFD0C" box="[151,337,691,717]" pageId="22" pageNumber="323" type="description">
<paragraph id="FC53EA07FFB592086083FD305E1FFD0C" blockId="22.[151,595,654,717]" box="[151,337,691,717]" pageId="22" pageNumber="323">
(
<figureCitation id="64D7F682FFB59208608BFD305FB8FD0C" box="[159,246,691,717]" captionStart="FIGURE 13" captionStartId="23.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@23.[151,1435,181,1798]" captionTargetPageId="23" captionText="FIGURE 13. Achradocera femoralis. A. Habitus, left lateral. B. Head, frontal. C. Thorax, dorsal. D. Legs, left lateral. E. Legs, focused on TII excavation. Scale bar: A, 1 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13915980" httpUri="https://zenodo.org/record/13915980/files/figure.png" pageId="22" pageNumber="323">Figs 13</figureCitation>
,
<figureCitation id="64D7F682FFB592086116FD305E6EFD0C" box="[258,288,691,717]" captionStart="FIGURE 14" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[205,1385,193,1908]" captionTargetId="figure-23@24.[192,1395,181,1919]" captionTargetPageId="24" captionText="FIGURE 14. Achradocera femoralis, male genital structures. A. Hypopygium, left lateral. B. Hypopygium, ventral. C. Phallus, ejaculatory apodeme and postgonite, left lateral. D. Postgonite, ventral. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13915982" httpUri="https://zenodo.org/record/13915982/files/figure.png" pageId="22" pageNumber="323">14</figureCitation>
,
<figureCitation id="64D7F682FFB592086138FD305E07FD0C" box="[300,329,691,717]" captionStart="FIGURE 26" captionStartId="40.[151,250,1764,1788]" captionTargetBox="[151,1432,597,1741]" captionTargetId="figure-242@40.[151,1436,597,1741]" captionTargetPageId="40" captionText="FIGURE 26. Distribution map of the genus Achradocera." figureDoi="http://doi.org/10.5281/zenodo.13916008" httpUri="https://zenodo.org/record/13916008/files/figure.png" pageId="22" pageNumber="323">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB592086083FD745E51FCEF" pageId="22" pageNumber="323" type="reference_group">
<paragraph id="FC53EA07FFB592086083FD745E51FCEF" blockId="22.[151,1436,759,814]" pageId="22" pageNumber="323">
<treatmentCitationGroup id="DCFCCD29FFB592086083FD745E51FCEF" pageId="22" pageNumber="323">
<treatmentCitation id="7D4DCC16FFB592086083FD745D2EFCCE" author="Becker, T." box="[151,608,759,783]" page="208" pageId="22" pageNumber="323" year="1922">
<taxonomicName id="3BEC9184FFB592086083FD745D2EFCCE" ID-CoL="89TRS" authority="Becker, 1922: 208" authorityName="Becker" authorityPageNumber="208" authorityYear="1922" box="[151,608,759,783]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB592086083FD745EC7FCCE" box="[151,393,759,783]" italics="true" pageId="22" pageNumber="323">Achradocera femoralis</emphasis>
<materialsCitation id="4C84E05AFFB592086182FD745D2EFCCE" box="[406,608,759,783]" collectionCode="MfN" country="Colombia" location="Colombia" pageId="22" pageNumber="323" specimenCount="1" typeStatus="lectotype">
<bibRefCitation id="987D97F6FFB592086182FD745D2EFCCE" author="Becker, T." box="[406,608,759,783]" pageId="22" pageNumber="323" pagination="1 - 394" refId="ref26227" refString="Becker, T. (1922) Dipterologische Studien. Dolichopodidae. B. Nearktische und neotropische Region. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13 (1), 1 - 394. https: // doi. org / 10.5962 / bhl. title. 132893" type="journal article" year="1922">Becker, 1922: 208</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type-locality:
<collectingCountry id="84FBAA97FFB592086318FD745C3DFCCE" box="[780,883,759,783]" name="Colombia" pageId="22" pageNumber="323">Colombia</collectingCountry>
, Barranquilla.
<typeStatus id="235754A5FFB592086402FD7B5B35FCD1" box="[1046,1147,760,784]" pageId="22" pageNumber="323" type="lectotype">Lectotype</typeStatus>
male, MfN [see Remarks section].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB592086083FCDF5DA7FB8E" pageId="22" pageNumber="323" type="diagnosis">
<paragraph id="FC53EA07FFB592086083FCDF5DA7FB8E" blockId="22.[151,1437,860,2044]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB592086083FCDF5E28FCB7" bold="true" box="[151,358,860,886]" pageId="22" pageNumber="323">Diagnosis (male).</emphasis>
Postpedicel longer than arista-like stylus; lower postoccipital setae dense. Legs mostly yellow, except for FIII, dorsal surface of FI, and distal tarsi, which are all brown infuscated; leg I simple; TII bulged ventrally in the middle, with two ventral rows of setae along the swelling, then constricted from apical 1/3 to the apex, forming excavation; IIt
<subScript id="6068E842FFB5920861F6FC595EA4FC29" attach="left" box="[482,490,986,1000]" fontSize="6" pageId="22" pageNumber="323">1</subScript>
arched, with ring of setae at basal 1/3 and anteroventral tubercule within ring; FIII bulged on basal half and concave below, bearing cluster of strong, long setae near middle of anterior surface; IIIt
<subScript id="6068E842FFB592086590FC7D5AD2FBCD" attach="left" box="[1412,1436,1022,1036]" fontSize="6" pageId="22" pageNumber="323">34</subScript>
concave ventrally. Hypopygium small; surstylus subtriangular, tapered and bearing 1 apical and 1 strong subapical seta; postgonite enlarged and covered by small setae.
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB5920860D3FBDB5EECF9E3" pageId="22" pageNumber="323" type="description">
<paragraph id="FC53EA07FFB5920860D3FBDB5A43FA3F" blockId="22.[151,1437,860,2044]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3FBDB5E3BFBB3" bold="true" box="[199,373,1112,1138]" pageId="22" pageNumber="323">Redescription.</emphasis>
Similar to
<taxonomicName id="3BEC9184FFB5920861FBFBDB5DEFFBB3" authorityName="Quevedo &amp; Capellari &amp; Lamas" authorityYear="2024" box="[495,673,1112,1138]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="angustifacies" status="stat. rev.">
<emphasis id="CE983615FFB5920861FBFBDB5DEFFBB3" box="[495,673,1112,1138]" italics="true" pageId="22" pageNumber="323">A. angustifacies</emphasis>
</taxonomicName>
<emphasis id="CE983615FFB5920862B3FBDB5C42FBB3" bold="true" box="[679,780,1112,1138]" pageId="22" pageNumber="323">
<taxonomicNameLabel id="D5AB8B6EFFB5920862B3FBDB5C42FBB3" box="[679,780,1112,1138]" pageId="22" pageNumber="323">stat. rev.</taxonomicNameLabel>
</emphasis>
, except as noted.
<emphasis id="CE983615FFB5920863C5FBDB5B5AFBB3" bold="true" box="[977,1044,1112,1138]" pageId="22" pageNumber="323">Male.</emphasis>
Body length:
<quantity id="3B1447E2FFB5920864A5FBDB5A59FBB2" box="[1201,1303,1112,1139]" metricMagnitude="-3" metricUnit="m" metricValue="2.68" pageId="22" pageNumber="323" unit="mm" value="2.68">2.68 mm</quantity>
. Wing:
<quantity id="3B1447E2FFB59208657FFBDB5F8DFB56" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="22" pageNumber="323" unit="mm" value="2.24">2.24 mm</quantity>
long,
<quantity id="3B1447E2FFB59208611EFBFF5E21FB56" box="[266,367,1148,1175]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="22" pageNumber="323" unit="mm" value="0.78">0.78 mm</quantity>
wide.
<emphasis id="CE983615FFB5920861AEFBFF5D66FB57" bold="true" box="[442,552,1148,1174]" pageId="22" pageNumber="323">Antenna.</emphasis>
Postpedicel longer than arista-like stylus; FAr ratio, 1.35.
<emphasis id="CE983615FFB5920864ADFBFF5A57FB57" bold="true" box="[1209,1305,1148,1174]" pageId="22" pageNumber="323">Thorax.</emphasis>
Bearing 10 ac and 5 dc pairs of setae.
<emphasis id="CE983615FFB5920861D3FB235D41FB7B" bold="true" box="[455,527,1184,1210]" pageId="22" pageNumber="323">Wing.</emphasis>
Lower calypter yellow with fan of about 67 brownish setae; CuAx ratio, 0.45; RMx ratio, 1.43.
<emphasis id="CE983615FFB59208614FFB475EECFB1F" bold="true" box="[347,418,1220,1246]" pageId="22" pageNumber="323">Leg I.</emphasis>
40, 35, 15, 7, 5, 4, 4. F bearing posterior row of about 4 setae at apical 1/4 apical; T with distinct apicoventral seta.
<emphasis id="CE983615FFB5920861A9FB6B5D40FAC3" bold="true" box="[445,526,1256,1282]" pageId="22" pageNumber="323">Leg II.</emphasis>
45, 50, 23, 14, 6, 5, 4. F yellow, with anterior and posterior row of about 5 setae at apical 1/4; T bearing anterior seta on basal 1/4, and 4 setae apically (1 av, 1 pv, 1 ad and 1 pd); T (
<figureCitation id="64D7F682FFB59208653BFA8F5ADAFAE7" box="[1327,1428,1292,1318]" captionStart="FIGURE 13" captionStartId="23.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@23.[151,1435,181,1798]" captionTargetPageId="23" captionText="FIGURE 13. Achradocera femoralis. A. Habitus, left lateral. B. Head, frontal. C. Thorax, dorsal. D. Legs, left lateral. E. Legs, focused on TII excavation. Scale bar: A, 1 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13915980" httpUri="https://zenodo.org/record/13915980/files/figure.png" pageId="22" pageNumber="323">Fig. 13E</figureCitation>
) bulged ventrally in middle, with two ventral rows of setae along swelling, constricted at apical 1/3 to apex, forming excavation; t
<subScript id="6068E842FFB592086132FAE55E60FAB5" attach="left" box="[294,302,1382,1396]" fontSize="6" pageId="22" pageNumber="323">1</subScript>
(
<figureCitation id="64D7F682FFB592086129FAD75ED0FAAF" box="[317,414,1364,1390]" captionStart="FIGURE 13" captionStartId="23.[151,250,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-17@23.[151,1435,181,1798]" captionTargetPageId="23" captionText="FIGURE 13. Achradocera femoralis. A. Habitus, left lateral. B. Head, frontal. C. Thorax, dorsal. D. Legs, left lateral. E. Legs, focused on TII excavation. Scale bar: A, 1 mm; BE, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13915980" httpUri="https://zenodo.org/record/13915980/files/figure.png" pageId="22" pageNumber="323">Fig. 13E</figureCitation>
) arched with small av tubercule on basal 1/5 with some spines apically, carrying ventral and posterior row of strong decreasing setae, with ad row making circlet of conspicuous setae.
<emphasis id="CE983615FFB592086498FAFB5BA9FA53" bold="true" box="[1164,1255,1400,1426]" pageId="22" pageNumber="323">Leg III.</emphasis>
32, 35, 10, 6, 4, 3, 3. F with anterior and posterior row of about 3 setae at apical 1/5; bulged on basal half and concave below, bearing cluster of strong anterior setae near middle; T bearing 3 pd, 2 ad, and 3 apical setae (1 av, 1 ad and 1 pd); T yellow; IIIt
<subScript id="6068E842FFB5920860AFFA755F9DF9C5" attach="left" box="[187,211,1526,1540]" fontSize="6" pageId="22" pageNumber="323">34</subScript>
concave ventrally.
<emphasis id="CE983615FFB5920861B9FA675D1EFA3F" bold="true" box="[429,592,1508,1534]" pageId="22" pageNumber="323">Hypopygium.</emphasis>
As in
<figureCitation id="64D7F682FFB59208628DFA675C6CFA3F" box="[665,802,1508,1534]" captionStart="FIGURE 14" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[205,1385,193,1908]" captionTargetId="figure-23@24.[192,1395,181,1919]" captionTargetPageId="24" captionText="FIGURE 14. Achradocera femoralis, male genital structures. A. Hypopygium, left lateral. B. Hypopygium, ventral. C. Phallus, ejaculatory apodeme and postgonite, left lateral. D. Postgonite, ventral. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13915982" httpUri="https://zenodo.org/record/13915982/files/figure.png" pageId="22" pageNumber="323">Fig. 14AD</figureCitation>
, similar to that of
<taxonomicName id="3BEC9184FFB5920863FAFA675BEFFA3F" authorityName="Quevedo &amp; Capellari &amp; Lamas" authorityYear="2024" box="[1006,1185,1508,1534]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="angustifacies" status="stat. rev.">
<emphasis id="CE983615FFB5920863FAFA675BEFFA3F" box="[1006,1185,1508,1534]" italics="true" pageId="22" pageNumber="323">A. angustifacies</emphasis>
</taxonomicName>
<emphasis id="CE983615FFB5920864BCFA675A43FA3F" bold="true" box="[1192,1293,1508,1534]" pageId="22" pageNumber="323">
<taxonomicNameLabel id="D5AB8B6EFFB5920864BCFA675A43FA3F" box="[1192,1293,1508,1534]" pageId="22" pageNumber="323">stat. rev.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="FC53EA07FFB5920860D3F98B5EECF9E3" blockId="22.[151,1437,860,2044]" box="[199,418,1544,1570]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3F98B5E6BF9E3" bold="true" box="[199,293,1544,1570]" pageId="22" pageNumber="323">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB5920860D3F9AF5D94F987" box="[199,730,1580,1606]" pageId="22" pageNumber="323" type="distribution">
<paragraph id="FC53EA07FFB5920860D3F9AF5D94F987" blockId="22.[151,1437,860,2044]" box="[199,730,1580,1606]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3F9AF5E2FF987" bold="true" box="[199,353,1580,1606]" pageId="22" pageNumber="323">Distribution.</emphasis>
<collectingCountry id="84FBAA97FFB592086173F9AF5E9BF987" box="[359,469,1580,1606]" name="Colombia" pageId="22" pageNumber="323">Colombia</collectingCountry>
and
<collectingCountry id="84FBAA97FFB592086218F9AF5D27F987" box="[524,617,1580,1606]" name="Curaçao" pageId="22" pageNumber="323">Curaçao</collectingCountry>
(
<figureCitation id="64D7F682FFB59208626DF9AF5D83F987" box="[633,717,1580,1606]" captionStart="FIGURE 26" captionStartId="40.[151,250,1764,1788]" captionTargetBox="[151,1432,597,1741]" captionTargetId="figure-242@40.[151,1436,597,1741]" captionTargetPageId="40" captionText="FIGURE 26. Distribution map of the genus Achradocera." figureDoi="http://doi.org/10.5281/zenodo.13916008" httpUri="https://zenodo.org/record/13916008/files/figure.png" pageId="22" pageNumber="323">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB5920860D3F9D35E9BF93B" pageId="22" pageNumber="323" type="materials_examined">
<paragraph id="FC53EA07FFB5920860D3F9D35C41F973" blockId="22.[151,1437,860,2044]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3F9D35EBDF9AB" bold="true" box="[199,499,1616,1642]" pageId="22" pageNumber="323">Type material examined.</emphasis>
<materialsCitation id="4C84E05AFFB5920861EFF9D35C41F973" collectionCode="III, MfN" pageId="22" pageNumber="323" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="CE983615FFB5920861EFF9D35DE9F9AB" bold="true" box="[507,679,1616,1642]" pageId="22" pageNumber="323">
<typeStatus id="235754A5FFB5920861EFF9D35DE9F9AB" box="[507,679,1616,1642]" pageId="22" pageNumber="323" type="lectotype">LECTOTYPE</typeStatus>
</emphasis>
[here designated]
<specimenCount id="EAEA218EFFB592086369F9D25CDAF9AA" box="[893,916,1617,1643]" count="1" pageId="22" pageNumber="323" type="male"></specimenCount>
, labelled: “Columbia | Ujhelyi”, “Baranquilla | 1912.
<collectionCode id="9AFD72C2FFB5920860F3F9F75E49F94F" box="[231,263,1652,1678]" pageId="22" pageNumber="323">III</collectionCode>
.”, “
<taxonomicName id="3BEC9184FFB59208613AF9F75E8EF94F" authorityName="Becker" authorityYear="1922" box="[302,448,1652,1678]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="CE983615FFB59208613AF9F75E8EF94F" box="[302,448,1652,1678]" italics="true" pageId="22" pageNumber="323">Achradocera</emphasis>
</taxonomicName>
| Beck.” [handwritten],
<taxonomicName id="3BEC9184FFB5920862C5F9F75C09F94F" box="[721,839,1652,1678]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB5920862C9F9F75C09F94F" box="[733,839,1652,1678]" italics="true" pageId="22" pageNumber="323">femoralis</emphasis>
</taxonomicName>
| Beck.” [handwritten], “Typus”, “MFN URI | http:// coll.mfn- | berlin.de/u/ | 09c83f” [
<collectionCode id="9AFD72C2FFB59208621AF91B5D04F973" box="[526,586,1688,1714]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="22" pageNumber="323" type="Museum">MfN</collectionCode>
, based on photo].
</materialsCitation>
</paragraph>
<paragraph id="FC53EA07FFB5920860D3F93F5E9BF93B" blockId="22.[151,1437,860,2044]" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3F93F5DE5F916" bold="true" box="[199,683,1724,1751]" pageId="22" pageNumber="323">Additional material examined. Curaçao.</emphasis>
<materialsCitation id="4C84E05AFFB5920862A7F93F5E9FF93B" collectingDate="1987-02-12" collectionCode="SMNS" collectorName="W. E. Steiner &amp; J. M. Swearingen" country="Curaçao" location="Savonet" pageId="22" pageNumber="323" specimenCount="1" specimenCount-male="1">
<specimenCount id="EAEA218EFFB5920862A7F93F5D99F917" box="[691,727,1724,1750]" count="1" pageId="22" pageNumber="323" type="male">1♂</specimenCount>
, “
<collectingCountry id="84FBAA97FFB5920862FAF93F5C34F917" box="[750,890,1724,1750]" name="Curaçao" pageId="22" pageNumber="323">CURAÇAO</collectingCountry>
|
<location id="F933BCDCFFB59208639BF93F5CA7F917" LSID="urn:lsid:plazi:treatment:74455B11FFB592076083FD0D5A13FDFC:F933BCDCFFB59208639BF93F5CA7F917" box="[911,1001,1724,1750]" country="Curaçao" name="Savonet" pageId="22" pageNumber="323">Savonet</location>
|
<date id="8852CCC7FFB5920863EAF93F5B89F917" box="[1022,1223,1724,1750]" pageId="22" pageNumber="323" value="1987-02-12">
<collectingDate id="9816352FFFB5920863EAF93F5B89F917" box="[1022,1223,1724,1750]" pageId="22" pageNumber="323" value="1987-02-12">12 February 1987</collectingDate>
</date>
|
<collectorName id="51198FD1FFB5920864CFF93E5A3FF917" box="[1243,1393,1724,1751]" pageId="22" pageNumber="323">W. E. Steiner</collectorName>
&amp; |
<collectorName id="51198FD1FFB592086083F9625E10F93B" box="[151,350,1760,1787]" pageId="22" pageNumber="323">J. M. Swearingen</collectorName>
” [
<collectionCode id="9AFD72C2FFB592086161F9635E84F93B" box="[373,458,1760,1786]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="22" pageNumber="323">SMNS</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4F6B98CFFB5920760D3F8875A13FDFC" lastPageId="25" lastPageNumber="326" pageId="22" pageNumber="323" type="discussion">
<paragraph id="FC53EA07FFB5920760D3F8875E9AFE48" blockId="22.[151,1437,860,2044]" lastBlockId="25.[151,1437,151,573]" lastPageId="25" lastPageNumber="326" pageId="22" pageNumber="323">
<emphasis id="CE983615FFB5920860D3F8875E75F8DF" bold="true" box="[199,315,1796,1822]" pageId="22" pageNumber="323">Remarks.</emphasis>
According to
<bibRefCitation id="987D97F6FFB5920861CAF8875DCCF8DF" author="Becker, T." box="[478,642,1796,1822]" pageId="22" pageNumber="323" pagination="1 - 394" refId="ref26227" refString="Becker, T. (1922) Dipterologische Studien. Dolichopodidae. B. Nearktische und neotropische Region. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13 (1), 1 - 394. https: // doi. org / 10.5962 / bhl. title. 132893" type="journal article" year="1922">Becker (1922)</bibRefCitation>
, the types of
<taxonomicName id="3BEC9184FFB59208630FF8875CE8F8DF" box="[795,934,1796,1822]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB59208630FF8875CE8F8DF" box="[795,934,1796,1822]" italics="true" pageId="22" pageNumber="323">A. femoralis</emphasis>
</taxonomicName>
were collected from
<collectingCountry id="84FBAA97FFB592086483F8865A4EF8DE" box="[1175,1280,1797,1823]" name="Paraguay" pageId="22" pageNumber="323">Paraguay</collectingCountry>
,
<collectingCountry id="84FBAA97FFB592086518F8875A25F8DF" box="[1292,1387,1796,1822]" name="Trinidad and Tobago" pageId="22" pageNumber="323">Trinidad</collectingCountry>
and
<collectingCountry id="84FBAA97FFB592086083F8AB5E4BF883" box="[151,261,1832,1858]" name="Colombia" pageId="22" pageNumber="323">Colombia</collectingCountry>
and were housed at the Hungarian Natural History Museum (HNHM, Budapest), whose collection was set on fire during a bombing in the 1956 Hungarian Revolution (
<bibRefCitation id="987D97F6FFB592086364F8CF5B52F8A7" author="Evenhuis, N. L." box="[880,1052,1868,1894]" pageId="22" pageNumber="323" pagination="1 - 178" refId="ref26782" refString="Evenhuis, N. L. (2006) Catalog of the Keroplatidae of the World. Bishop Museum Bulletins in Entomology 13, 1 - 178." type="journal article" year="2006">Evenhuis 2006</bibRefCitation>
) and, as such, the types should be considered as lost. Nevertheless, a male specimen of
<taxonomicName id="3BEC9184FFB592086315F8F35CDDF84B" authorityName="Becker" authorityYear="1922" box="[769,915,1904,1930]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="CE983615FFB592086315F8F35CDDF84B" box="[769,915,1904,1930]" italics="true" pageId="22" pageNumber="323">Achradocera</emphasis>
</taxonomicName>
from
<collectingCountry id="84FBAA97FFB5920863CAF8F35B01F84B" box="[990,1103,1904,1930]" name="Colombia" pageId="22" pageNumber="323">Colombia</collectingCountry>
, labeled as “typus” and with Beckers handwritten label
<taxonomicName id="3BEC9184FFB5920861C5F8175D06F86F" box="[465,584,1940,1966]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB5920861CAF8175D06F86F" box="[478,584,1940,1966]" italics="true" pageId="22" pageNumber="323">femoralis</emphasis>
</taxonomicName>
”, was later found in the MfN holdings in Berlin. This male type in Berlin clearly does not fit
<bibRefCitation id="987D97F6FFB59208616CF83B5D61F813" author="Becker, T." box="[376,559,1976,2002]" pageId="22" pageNumber="323" pagination="1 - 394" refId="ref26227" refString="Becker, T. (1922) Dipterologische Studien. Dolichopodidae. B. Nearktische und neotropische Region. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13 (1), 1 - 394. https: // doi. org / 10.5962 / bhl. title. 132893" type="journal article" year="1922">Beckers (1922)</bibRefCitation>
original description (stated as having unmodified legs), as the specimen has modified TI, FII, IIt
<subScript id="6068E842FFB592086162F86D5E30F83D" attach="left" box="[374,382,2030,2044]" fontSize="6" pageId="22" pageNumber="323">1</subScript>
, FIII and IIIt
<subScript id="6068E842FFB592086200F86D5D62F83D" attach="left" box="[532,556,2030,2044]" fontSize="6" pageId="22" pageNumber="323">34</subScript>
. It is not unexpected that specimens from distant localities such as
<collectingCountry id="84FBAA97FFB59208653EF85E5ADAF836" box="[1322,1428,2013,2039]" name="Paraguay" pageId="22" pageNumber="323">Paraguay</collectingCountry>
,
<collectingCountry id="84FBAA97FFBA92076083FF145FB8FF70" box="[151,246,151,177]" name="Trinidad and Tobago" pageId="25" pageNumber="326">Trinidad</collectingCountry>
and
<collectingCountry id="84FBAA97FFBA9207613EFF145ED6FF70" box="[298,408,151,177]" name="Colombia" pageId="25" pageNumber="326">Colombia</collectingCountry>
were not conspecific. We assume that Becker probably overlooked salient differences among the specimens of his type series, as he did before with
<taxonomicName id="3BEC9184FFBA920762ECFF385B53FF14" authority="Becker" authorityName="Becker" authorityYear="1922" box="[760,1053,187,213]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="pictipes">
<emphasis id="CE983615FFBA920762ECFF385C89FF15" box="[760,967,187,213]" italics="true" pageId="25" pageNumber="326">Chrysotus pictipes</emphasis>
Becker
</taxonomicName>
(see
<bibRefCitation id="987D97F6FFBA9207644CFF385AD2FF14" author="Capellari, R. S. &amp; Almeida, K. F." box="[1112,1436,187,213]" pageId="25" pageNumber="326" pagination="18 - 27" refId="ref26487" refString="Capellari, R. S. &amp; Almeida, K. F. (2024) On the identity of three species of Chrysotus Meigen (Diptera: Dolichopodidae) described from Brazil by Millard C. Van Duzee. Journal of Insect Biodiversity, 45 (1), 18 - 27. https: // doi. org / 10.12976 / jib / 2024.45.1.3" type="journal article" year="2024">Capellari &amp; Almeida (2024))</bibRefCitation>
and, in similar way, regarded as conspecific specimens from distant localities, such as
<collectingCountry id="84FBAA97FFBA9207649DFF5C5BBFFF38" box="[1161,1265,223,249]" name="Paraguay" pageId="25" pageNumber="326">Paraguay</collectingCountry>
and
<collectingCountry id="84FBAA97FFBA92076539FF5C5AD2FF38" box="[1325,1436,223,249]" name="Argentina" pageId="25" pageNumber="326">Argentina</collectingCountry>
with
<collectingRegion id="3E2824E5FFBA920760C7FE805E05FEDC" box="[211,331,259,285]" country="United States of America" name="New York" pageId="25" pageNumber="326">New York</collectingRegion>
,
<collectingCountry id="84FBAA97FFBA9207614DFE805EE8FEDC" box="[345,422,259,285]" name="United States of America" pageId="25" pageNumber="326">U.S.A.</collectingCountry>
(for
<taxonomicName id="3BEC9184FFBA920761F1FE805D90FEDD" authorityName="Becker" authorityYear="1922" box="[485,734,259,285]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="erectus">
<emphasis id="CE983615FFBA920761F1FE805D90FEDD" box="[485,734,259,285]" italics="true" pageId="25" pageNumber="326">Condylostylus erectus</emphasis>
</taxonomicName>
, see
<bibRefCitation id="987D97F6FFBA92076309FE805C8AFEDC" author="Bickel, D. J." box="[797,964,259,285]" pageId="25" pageNumber="326" pagination="545 - 560" refId="ref26319" refString="Bickel, D. J. (2003) New synonyms in the Nearctic and Neotropical Sciapodinae (Diptera: Dolichopodidae). Studia dipterologica, 9, 545 - 560." type="journal article" year="2003">Bickel (2003))</bibRefCitation>
and
<collectingCountry id="84FBAA97FFBA92076415FE805B27FEDC" box="[1025,1129,259,285]" name="Paraguay" pageId="25" pageNumber="326">Paraguay</collectingCountry>
with
<collectingRegion id="3E2824E5FFBA920764A4FE805A43FEDC" box="[1200,1293,259,285]" country="United States of America" name="Georgia" pageId="25" pageNumber="326">Georgia</collectingRegion>
,
<collectingCountry id="84FBAA97FFBA9207650FFE805A26FEDC" box="[1307,1384,259,285]" name="United States of America" pageId="25" pageNumber="326">U.S.A.</collectingCountry>
(for
<taxonomicName id="3BEC9184FFBA92076083FEA45E39FE80" authorityName="Becker" authorityYear="1922" box="[151,375,295,321]" class="Insecta" family="Dolichopodidae" genus="Mesorhaga" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="jucunda">
<emphasis id="CE983615FFBA92076083FEA45E39FE80" box="[151,375,295,321]" italics="true" pageId="25" pageNumber="326">Mesorhaga jucunda</emphasis>
</taxonomicName>
, see
<bibRefCitation id="987D97F6FFBA920761BBFEA45D19FE80" author="Bickel, D. J." box="[431,599,295,321]" pageId="25" pageNumber="326" pagination="47 - 67" refId="ref26351" refString="Bickel, D. J. (2007) The Mesoamerican Mesorhaga (Diptera: Dolichopodidae), with a taxonomic conspectus of the New World fauna. Zootaxa, 1411 (1), 47 - 67. https: // doi. org / 10.11646 / zootaxa. 1411.1.5" type="journal article" year="2007">Bickel (2007))</bibRefCitation>
. Even though the remaining type does not fit Beckers original description, it still is the name-bearer specimen of
<taxonomicName id="3BEC9184FFBA9207622BFEC85D89FEA4" box="[575,711,331,357]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFBA9207622BFEC85D89FEA4" box="[575,711,331,357]" italics="true" pageId="25" pageNumber="326">A. femoralis</emphasis>
</taxonomicName>
and we are herein designating it as the
<typeStatus id="235754A5FFBA9207646FFEC85BACFEA4" box="[1147,1250,331,357]" pageId="25" pageNumber="326" type="lectotype">lectotype</typeStatus>
to ensure proper identification of the species.
</paragraph>
<caption id="A893BA8FFFB492096083F89D5C45F898" ID-DOI="http://doi.org/10.5281/zenodo.13915980" ID-Zenodo-Dep="13915980" httpUri="https://zenodo.org/record/13915980/files/figure.png" pageId="23" pageNumber="324" startId="23.[151,250,1822,1846]" targetBox="[151,1435,181,1798]" targetPageId="23" targetType="figure">
<paragraph id="FC53EA07FFB492096083F89D5C45F898" blockId="23.[151,1436,1822,1882]" pageId="23" pageNumber="324">
<emphasis id="CE983615FFB492096083F89D5E6EF8F7" bold="true" box="[151,288,1822,1846]" pageId="23" pageNumber="324">FIGURE 13.</emphasis>
<taxonomicName id="3BEC9184FFB492096132F89D5D5EF8F7" box="[294,528,1822,1846]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFB492096132F89D5D5EF8F7" box="[294,528,1822,1846]" italics="true" pageId="23" pageNumber="324">Achradocera femoralis</emphasis>
</taxonomicName>
.
<emphasis id="CE983615FFB492096208F89D5D7BF8F7" bold="true" box="[540,565,1822,1846]" pageId="23" pageNumber="324">A.</emphasis>
Habitus, left lateral.
<emphasis id="CE983615FFB49209631EF89D5C6FF8F7" bold="true" box="[778,801,1822,1846]" pageId="23" pageNumber="324">B.</emphasis>
Head, frontal.
<emphasis id="CE983615FFB4920963ADF89D5C9FF8F7" bold="true" box="[953,977,1822,1846]" pageId="23" pageNumber="324">C.</emphasis>
Thorax, dorsal.
<emphasis id="CE983615FFB492096463F89D5BDEF8F7" bold="true" box="[1143,1168,1822,1846]" pageId="23" pageNumber="324">D.</emphasis>
Legs, left lateral.
<emphasis id="CE983615FFB492096553F89D5A11F8F7" bold="true" box="[1351,1375,1822,1846]" pageId="23" pageNumber="324">E.</emphasis>
Legs, focused on TII excavation. Scale bar: A, 1 mm; BE, 0.2 mm.
</paragraph>
</caption>
<caption id="A893BA8FFFBB92066083F8145B60F812" ID-DOI="http://doi.org/10.5281/zenodo.13915982" ID-Zenodo-Dep="13915982" httpUri="https://zenodo.org/record/13915982/files/figure.png" pageId="24" pageNumber="325" startId="24.[151,250,1943,1967]" targetBox="[205,1385,193,1908]" targetPageId="24" targetType="figure">
<paragraph id="FC53EA07FFBB92066083F8145B60F812" blockId="24.[151,1436,1943,2004]" pageId="24" pageNumber="325">
<emphasis id="CE983615FFBB92066083F8145E51F86E" bold="true" box="[151,287,1943,1967]" pageId="24" pageNumber="325">FIGURE 14.</emphasis>
<taxonomicName id="3BEC9184FFBB92066131F8145D40F86E" box="[293,526,1943,1967]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFBB92066131F8145D40F86E" box="[293,526,1943,1967]" italics="true" pageId="24" pageNumber="325">Achradocera femoralis</emphasis>
</taxonomicName>
, male genital structures.
<emphasis id="CE983615FFBB9206631EF8145C6DF86E" bold="true" box="[778,803,1943,1967]" pageId="24" pageNumber="325">A.</emphasis>
Hypopygium, left lateral.
<emphasis id="CE983615FFBB92066438F81B5B0DF871" bold="true" box="[1068,1091,1944,1968]" pageId="24" pageNumber="325">B.</emphasis>
Hypopygium, ventral.
<emphasis id="CE983615FFBB92066539F8145A08F86E" bold="true" box="[1325,1350,1943,1967]" pageId="24" pageNumber="325">C.</emphasis>
Phallus, ejaculatory apodeme and postgonite, left lateral.
<emphasis id="CE983615FFBB92066291F83F5DD0F815" bold="true" box="[645,670,1980,2004]" pageId="24" pageNumber="325">D.</emphasis>
Postgonite, ventral. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="FC53EA07FFBA920760D3FE105A13FDFC" blockId="25.[151,1437,151,573]" pageId="25" pageNumber="326">
It is worth mentioning that Beckers original concept for this species (based on the 1922 description) was the same used by
<bibRefCitation id="987D97F6FFBA92076120FE345EA1FE10" author="Robinson, H." box="[308,495,439,465]" pageId="25" pageNumber="326" refId="ref27766" refString="Robinson, H. (1970) 40. Family Dolichopodidae. In: A catalogue of the Diptera of the Americas South of United States. Departamento de Zoologia, Sap Paulo, pp. 40.1 - 40.92." type="book" year="1970">Robinson (1970)</bibRefCitation>
for a subsequent designation of a type-species for the genus. We see no need to shift the
<typeStatus id="235754A5FFBA920760ABFE585FA1FE34" box="[191,239,475,501]" pageId="25" pageNumber="326">type</typeStatus>
species of the genus, despite the change in concept of
<taxonomicName id="3BEC9184FFBA9207632BFE585C8BFE34" box="[831,965,475,501]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFBA9207632BFE585C8BFE34" box="[831,965,475,501]" italics="true" pageId="25" pageNumber="326">A. femoralis</emphasis>
</taxonomicName>
(which was maintained as the type-species of
<taxonomicName id="3BEC9184FFBA920760A0FE7C5E0BFDD8" authorityName="Becker" authorityYear="1922" box="[180,325,511,537]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="CE983615FFBA920760A0FE7C5E0BFDD8" box="[180,325,511,537]" italics="true" pageId="25" pageNumber="326">Achradocera</emphasis>
</taxonomicName>
). A concept similar to Beckers
<taxonomicName id="3BEC9184FFBA920762B2FE7C5C63FDD8" box="[678,813,511,537]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CE983615FFBA920762B2FE7C5C63FDD8" box="[678,813,511,537]" italics="true" pageId="25" pageNumber="326">A. femoralis</emphasis>
</taxonomicName>
is applied below to describe the new species
<taxonomicName id="3BEC9184FFBA9207650AFE7C5A3BFDD8" authorityName="Quevedo &amp; Capellari &amp; Lamas" authorityYear="2024" box="[1310,1397,511,537]" class="Insecta" family="Dolichopodidae" genus="Achradocera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="gimli" status="sp. nov.">
<emphasis id="CE983615FFBA9207650AFE7C5A3BFDD8" box="[1310,1397,511,537]" italics="true" pageId="25" pageNumber="326">A. gimli</emphasis>
</taxonomicName>
<emphasis id="CE983615FFBA9207656FFE7C5F86FDFC" bold="true" pageId="25" pageNumber="326">
<taxonomicNameLabel id="D5AB8B6EFFBA9207656FFE7C5F86FDFC" pageId="25" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the species may correspond to Beckers specimens from
<collectingCountry id="84FBAA97FFBA92076367FDA05C94FDFC" box="[883,986,547,573]" name="Paraguay" pageId="25" pageNumber="326">Paraguay</collectingCountry>
and
<collectingCountry id="84FBAA97FFBA9207641BFDA05B23FDFC" box="[1039,1133,547,573]" name="Trinidad and Tobago" pageId="25" pageNumber="326">Trinidad</collectingCountry>
(HNHM, destroyed).
</paragraph>
</subSubSection>
</treatment>
</document>