treatments-xml/data/21/7A/02/217A020B6E503F0AFF6C4BAEFF0CCAEA.xml
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<document id="BFCB1E08B57C04577ED94E5593D173A2" ID-CLB-Dataset="115950" ID-DOI="10.11646/zootaxa.5174.1.1" ID-GBIF-Dataset="c4bf1061-5af8-4f4a-b4ca-234b6d0ca615" ID-ISSN="1175-5326" ID-Zenodo-Dep="6972960" ID-ZooBank="F80F144B-D1E8-4587-A146-0BACFFE18FB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659936256232" checkinUser="plazi" docAuthor="Beaver, Roger A. &amp; Smith, Sarah M." docDate="2022" docId="217A020B6E503F0AFF6C4BAEFF0CCAEA" docLanguage="en" docName="zootaxa.5174.1.1.pdf" docOrigin="Zootaxa 5174 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xylosandrus geduensis Smith &amp; Beaver 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="8" masterDocId="DD437A736E543F0DFFFB4D0DFFFCCF2B" masterDocTitle="The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae" masterLastPageNumber="24" masterPageNumber="1" pageNumber="5" updateTime="1699365110226" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="373B7DF804B2D3385CA29055C2A6C95D">The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae</mods:title>
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<mods:namePart id="85CD3932F2B013EF301E78993ABD6236">Beaver, Roger A.</mods:namePart>
<mods:affiliation id="392C1E85B3B220C3A28C2F68F3E70639">161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
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<mods:namePart id="9B4083619CF2D600766C55E1A153F8FE">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="DAD81367FE5ACB9CA579606D28A192A7" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="B3D893D8C8CCFDEF8D363C13DB06EDB6">Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
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<treatment id="217A020B6E503F0AFF6C4BAEFF0CCAEA" ID-DOI="http://doi.org/10.5281/zenodo.6972972" ID-GBIF-Taxon="198633434" ID-Zenodo-Dep="6972972" LSID="urn:lsid:plazi:treatment:217A020B6E503F0AFF6C4BAEFF0CCAEA" httpUri="http://treatment.plazi.org/id/217A020B6E503F0AFF6C4BAEFF0CCAEA" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
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<paragraph id="A96CB31D6E503F09FF6C4BAEFD1FC996" blockId="4.[151,1175,1699,1798]" box="[151,739,1699,1726]" pageId="4" pageNumber="5">
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<emphasis id="9BA76F0F6E503F09FF6C4BAEFD1FC996" bold="true" box="[151,739,1699,1726]" pageId="4" pageNumber="5">
<taxonomicName id="6ED3C89E6E503F09FF6C4BAEFD82C996" authority="Smith &amp; Beaver" authorityName="Smith &amp; Beaver" authorityYear="2022" box="[151,638,1699,1726]" class="Insecta" family="Curculionidae" genus="Xylosandrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="geduensis" status="sp. nov.">
<emphasis id="9BA76F0F6E503F09FF6C4BAEFE51C995" bold="true" box="[151,429,1699,1726]" italics="true" pageId="4" pageNumber="5">Xylosandrus geduensis</emphasis>
Smith &amp; Beaver
</taxonomicName>
<taxonomicNameLabel id="8094D2746E503F09FD7E4BAEFD1FC996" box="[645,739,1699,1725]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="E1C9E0966E503F09FF6C4BCAFB6BC82D" pageId="4" pageNumber="5" type="description">
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(
<figureCitation id="31E8AF986E503F09FF644BCAFEEDC9CA" box="[159,273,1735,1761]" captionStart="FIGURES 912" captionStartId="6.[151,264,1981,2005]" captionTargetBox="[165,1423,181,1957]" captionTargetId="figure-23@6.[165,1423,181,1958]" captionTargetPageId="6" captionText="FIGURES 912. Xylosandrus geduensis holotype, 2.1 mm. Dorsal, lateral, posterior oblique and anterior views." figureDoi="http://doi.org/10.5281/zenodo.6972970" httpUri="https://zenodo.org/record/6972970/files/figure.png" pageId="4" pageNumber="5">Figs 912</figureCitation>
)
</paragraph>
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="E1C9E0966E503F09FF6C4A3EFE0FC85A" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="A96CB31D6E503F09FF6C4A3EFE0FC85A" blockId="4.[151,1437,1843,2014]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF6C4A3EFEB8C866" bold="true" box="[151,324,1843,1869]" pageId="4" pageNumber="5">Type material.</emphasis>
<materialsCitation id="19BBB9406E503F09FEB14A3EFE0FC85A" ID-GBIF-Occurrence="3880946321" accessionNumber="A17491" collectionCode="NHML" collectorName="X. Padgham" country="Bhutan" location="Bhutan" pageId="4" pageNumber="5" specimenCode="NHMUK014591825" specimenCount="1" typeStatus="holotype">
<typeStatus id="76680DBF6E503F09FEB14A3EFE49C866" box="[330,437,1843,1869]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="D1C4F38D6E503F09FDE24A3EFD91C866" box="[537,621,1843,1869]" name="Bhutan" pageId="4" pageNumber="5">Bhutan</collectingCountry>
, Gedu, 6.viii.[19]85,
<collectorName id="0426D6CB6E503F09FCA14A39FC1FC866" box="[858,995,1843,1870]" pageId="4" pageNumber="5">X. Padgham</collectorName>
coll., ex
<taxonomicName id="6ED3C89E6E503F09FBB34A39FAADC866" authority="CIE A" authorityName="CIE A" box="[1096,1361,1843,1869]" class="Magnoliopsida" family="Euphorbiaceae" genus="Trewia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="nudiflora">
<emphasis id="9BA76F0F6E503F09FBB34A39FAFFC866" box="[1096,1283,1843,1869]" italics="true" pageId="4" pageNumber="5">Trewia nudiflora</emphasis>
CIE A
</taxonomicName>
17491,
<specimenCode id="F9751B666E503F09FF6C4A5AFE83C859" box="[151,383,1879,1906]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="5" type="Museum">NHMUK014591825</specimenCode>
(
<collectionCode id="CFC22BD86E503F09FE754A5AFE1BC85A" box="[398,487,1879,1905]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="5">NHML</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F0AFF3C4A76FEFECFD2" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="A96CB31D6E503F0AFF3C4A76FEFECFD2" blockId="4.[151,1437,1843,2014]" lastBlockId="7.[151,1437,151,1473]" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4A76FEBEC8BE" bold="true" box="[199,322,1915,1941]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
<quantity id="6E2B1EF86E503F09FEBC4A76FE60C8BD" box="[327,412,1915,1942]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="5" unit="mm" value="2.1">2.1 mm</quantity>
long (n = 1); 2.33 × as long as wide. This species is distinguished by the small size; declivity obliquely truncate, abruptly separated from disc; posterolateral margins of elytra carinate to interstriae 7; declivital face with 5 punctate striae; declivital interstriae with densely uniseriate denticles; pronotum from dorsal view rounded (
<typeStatus id="76680DBF6E533F0AFF644D9AFF2DCF9A" box="[159,209,151,177]" pageId="7" pageNumber="8">type</typeStatus>
1) and lateral view basic (
<typeStatus id="76680DBF6E533F0AFE014D9AFDD0CF9A" box="[506,556,151,177]" pageId="7" pageNumber="8">type</typeStatus>
0), pronotal summit at midpoint, basal 1/2 smooth, shining, sparsely minutely punctate; sparse mycangial tuft on the pronotal base; and outer margin of protibia armed with 5 large socketed denticles.
</paragraph>
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<caption id="FDACE3956E513F08FF6C4ACAFB10C8F4" ID-DOI="http://doi.org/10.5281/zenodo.6972966" ID-Zenodo-Dep="6972966" box="[151,1260,1991,2015]" httpUri="https://zenodo.org/record/6972966/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,264,1991,2015]" targetBox="[163,1428,181,1966]" targetPageId="5">
<paragraph id="A96CB31D6E513F08FF6C4ACAFB10C8F4" blockId="5.[151,1260,1991,2015]" box="[151,1260,1991,2015]" pageId="5" pageNumber="6">
<emphasis id="9BA76F0F6E513F08FF6C4ACAFEC7C8F4" bold="true" box="[151,315,1991,2015]" pageId="5" pageNumber="6">FIGURES 58.</emphasis>
<taxonomicName id="6ED3C89E6E513F08FEBA4ACAFDE9C8F4" box="[321,533,1991,2015]" class="Insecta" family="Curculionidae" genus="Euwallacea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="9BA76F0F6E513F08FEBA4ACAFDE9C8F4" box="[321,533,1991,2015]" italics="true" pageId="5" pageNumber="6">Euwallacea insolitus</emphasis>
</taxonomicName>
holotype, 2.9 mm. Dorsal, lateral, posterior oblique and anterior views.
</paragraph>
</caption>
<caption id="FDACE3956E523F0BFF6C4AB0FAF1C8FD" ID-DOI="http://doi.org/10.5281/zenodo.6972970" ID-Zenodo-Dep="6972970" box="[151,1293,1981,2006]" httpUri="https://zenodo.org/record/6972970/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,264,1981,2005]" targetBox="[165,1423,181,1957]" targetPageId="6">
<paragraph id="A96CB31D6E523F0BFF6C4AB0FAF1C8FD" blockId="6.[151,1293,1981,2006]" box="[151,1293,1981,2006]" pageId="6" pageNumber="7">
<emphasis id="9BA76F0F6E523F0BFF6C4AB0FEBBC8FE" bold="true" box="[151,327,1981,2005]" pageId="6" pageNumber="7">FIGURES 912.</emphasis>
<taxonomicName id="6ED3C89E6E523F0BFEB64AB3FDCAC8FD" box="[333,566,1982,2006]" class="Insecta" family="Curculionidae" genus="Xylosandrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="geduensis">
<emphasis id="9BA76F0F6E523F0BFEB64AB3FDCAC8FD" box="[333,566,1982,2006]" italics="true" pageId="6" pageNumber="7">Xylosandrus geduensis</emphasis>
</taxonomicName>
holotype, 2.1 mm. Dorsal, lateral, posterior oblique and anterior views.
</paragraph>
</caption>
<subSubSection id="E1C9E0966E533F0AFF3C4C0FFE8ECB56" pageId="7" pageNumber="8" type="description">
<paragraph id="A96CB31D6E533F0AFF3C4C0FFE8ECB56" blockId="7.[151,1437,151,1473]" pageId="7" pageNumber="8">
<emphasis id="9BA76F0F6E533F0AFF3C4C0FFE3ECE37" bold="true" box="[199,450,258,284]" pageId="7" pageNumber="8">Description (female).</emphasis>
<quantity id="6E2B1EF86E533F0AFE324C0EFDE3CE36" box="[457,543,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="8" unit="mm" value="2.1">2.1 mm</quantity>
long (n = 1); 2.33 × as long as wide. Head, pronotum and elytral disc light redbrown, declivity dark red-brown, antennae and legs light brown.
<emphasis id="9BA76F0F6E533F0AFC8C4C2AFC3CCE6A" bold="true" box="[887,960,295,321]" pageId="7" pageNumber="8">Head:</emphasis>
epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; median carina present; surface shagreened, sparsely punctate, punctures, large, shallow, alutaceous, sparsely setose, setae erect, hair-like. Eyes very shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum broad, triangular, deeply impressed. Antennal scape regularly thick, approximately as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 as long as pedicel. Club 1.1 × taller than broad, subcircular, obliquely truncate,
<typeStatus id="76680DBF6E533F0AFAB64CD6FA81CEDE" box="[1357,1405,475,501]" pageId="7" pageNumber="8">type</typeStatus>
1; segment 1 corneous, encircling anterior face, margin costate.
<emphasis id="9BA76F0F6E533F0AFCA34CF2FC26CD32" bold="true" box="[856,986,511,537]" pageId="7" pageNumber="8">Pronotum:</emphasis>
1.11 × as long as wide. In dorsal view rounded,
<typeStatus id="76680DBF6E533F0AFEFF4F2EFEC8CD16" box="[260,308,547,573]" pageId="7" pageNumber="8">type</typeStatus>
1, sides parallel in basal 1/2, rounded anteriorly; anterior margin with a row of six serrations. In lateral view basic,
<typeStatus id="76680DBF6E533F0AFE954F4AFE62CD4A" box="[366,414,583,609]" pageId="7" pageNumber="8">type</typeStatus>
0, disc flat, summit at midpoint. Anterior slope with densely spaced, moderate asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, erect hair-like setae, some longer hair-like setae at anterior and lateral margins. Disc shining, alutaceous with sparse, fine punctures, glabrous. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Mycangial tuft present along basal margin, tuft broad, sparse, laterally extending to striae 3.
<emphasis id="9BA76F0F6E533F0AFCE84FDAFC95CDDA" bold="true" box="[787,873,727,753]" pageId="7" pageNumber="8">Elytra:</emphasis>
1.22 × as long as wide, 1.1 × as long as pronotum. Scutellum moderately sized, linguiform, flush with elytra, flat, shining. Elytral base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 2/3, then broadly rounded to apex. Striae punctate, not impressed. Disc occupying basal half of elytra, shining, interstriae punctate, moderately setose, setae semi-erect, hair-like. Declivity sharply distinct from disc, 5 striae visible, strial punctures larger, deeper than those of disc; interstriae with densely uniseriate denticles, denticles small, spaced 2-4 diameters of a denticle, with erect hair-like setae, as long as the distance between suture and striae 2. Apex emarginate, margins acutely carinate. Posterolateral margin carinate to interstriae 7.
<emphasis id="9BA76F0F6E533F0AFEA84EDEFE68CCC6" bold="true" box="[339,404,979,1005]" pageId="7" pageNumber="8">Legs:</emphasis>
procoxae widely separated; prosternal coxal piece flat, inconspicuous. Protibiae slender, broadest at apical 1/4; posterior face smooth; outer margin of apical 1/2 with 5 large socketed denticles, their length much longer than basal width; apical mucro prominent, strongly incurved. Meso- and metatibiae flattened, outer margins evenly rounded with 10 and 12, small similarly sized socketed denticles, their length no longer than basal width, respectively.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E533F0AFF3C498BFE3CCB8A" box="[199,448,1158,1185]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="A96CB31D6E533F0AFF3C498BFE3CCB8A" blockId="7.[151,1437,151,1473]" box="[199,448,1158,1185]" pageId="7" pageNumber="8">
<emphasis id="9BA76F0F6E533F0AFF3C498BFE9DCB8B" bold="true" box="[199,353,1158,1184]" pageId="7" pageNumber="8">Distribution.</emphasis>
<collectingCountry id="D1C4F38D6E533F0AFE9C498AFE40CB8A" box="[359,444,1159,1185]" name="Bhutan" pageId="7" pageNumber="8">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E533F0AFF3C49A6FC8DCBEE" box="[199,881,1195,1221]" pageId="7" pageNumber="8" type="biology_ecology">
<paragraph id="A96CB31D6E533F0AFF3C49A6FC8DCBEE" blockId="7.[151,1437,151,1473]" box="[199,881,1195,1221]" pageId="7" pageNumber="8">
<emphasis id="9BA76F0F6E533F0AFF3C49A6FEAACBEE" bold="true" box="[199,342,1195,1221]" pageId="7" pageNumber="8">
<collectingRegion id="6B177DFF6E533F0AFF3C49A6FF03CBEE" box="[199,255,1195,1221]" country="Afghanistan" name="Khost" pageId="7" pageNumber="8">Host</collectingRegion>
plants.
</emphasis>
<taxonomicName id="6ED3C89E6E533F0AFEA649A6FE3DCBEE" box="[349,449,1195,1221]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="9BA76F0F6E533F0AFEA649A6FE3DCBEE" box="[349,449,1195,1221]" italics="true" pageId="7" pageNumber="8">Mallotus</emphasis>
</taxonomicName>
(=
<taxonomicName id="6ED3C89E6E533F0AFE1A49A6FDD2CBEF" box="[481,558,1195,1220]" class="Magnoliopsida" family="Euphorbiaceae" genus="Trewia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="9BA76F0F6E533F0AFE1A49A6FDD2CBEF" box="[481,558,1195,1220]" italics="true" pageId="7" pageNumber="8">Trewia</emphasis>
</taxonomicName>
)
<emphasis id="9BA76F0F6E533F0AFDC549A6FD4DCBEE" box="[574,689,1195,1221]" italics="true" pageId="7" pageNumber="8">nudiflorus</emphasis>
(
<taxonomicName id="6ED3C89E6E533F0AFD3B49A6FC96CBEE" box="[704,874,1195,1221]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E533F0AFF3C49C2FC14CBC2" box="[199,1000,1231,1257]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="A96CB31D6E533F0AFF3C49C2FC14CBC2" blockId="7.[151,1437,151,1473]" box="[199,1000,1231,1257]" pageId="7" pageNumber="8">
<emphasis id="9BA76F0F6E533F0AFF3C49C2FEB2CBC2" bold="true" box="[199,334,1231,1257]" pageId="7" pageNumber="8">Etymology.</emphasis>
In reference to the
<typeStatus id="76680DBF6E533F0AFDD349C2FDA4CBC2" box="[552,600,1231,1257]" pageId="7" pageNumber="8">type</typeStatus>
locality, Gedu. Latinized adjective.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E533F0AFF3C49FEFF0CCAEA" pageId="7" pageNumber="8" type="discussion">
<paragraph id="A96CB31D6E533F0AFF3C49FEFF0CCAEA" blockId="7.[151,1437,151,1473]" pageId="7" pageNumber="8">
<emphasis id="9BA76F0F6E533F0AFF3C49FEFEC7CA26" bold="true" box="[199,315,1267,1293]" pageId="7" pageNumber="8">Remarks.</emphasis>
This species is very similar to the Indomalayan species
<taxonomicName id="6ED3C89E6E533F0AFC3149FEFA65CA26" authority="(Schedl)" baseAuthorityName="Schedl" box="[970,1433,1267,1293]" class="Insecta" family="Curculionidae" genus="Xylosandrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="derupteterminatus">
<emphasis id="9BA76F0F6E533F0AFC3149FEFAD1CA26" box="[970,1325,1267,1293]" italics="true" pageId="7" pageNumber="8">Xylosandrus derupteterminatus</emphasis>
(Schedl)
</taxonomicName>
. It can be separated by the following combination of characters (
<taxonomicName id="6ED3C89E6E533F0AFC93481AFC0ACA1A" box="[872,1014,1303,1329]" class="Insecta" family="Curculionidae" genus="Xylosandrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="geduensis">
<emphasis id="9BA76F0F6E533F0AFC93481AFC0ACA1A" box="[872,1014,1303,1329]" italics="true" pageId="7" pageNumber="8">X. geduensis</emphasis>
</taxonomicName>
given first): elongate form, 2.33 × as long as wide vs stout 1.822.0 × as long as wide (
<bibRefCitation id="CD42CEEC6E533F0AFD2B4836FC60CA7E" author="Smith, S. M. &amp; Beaver, R. A. &amp; Cognato, A. I." box="[720,924,1339,1365]" pageId="7" pageNumber="8" pagination="1 - 442" refId="ref16478" refString="Smith, S. M., Beaver, R. A. &amp; Cognato, A. I. (2020) A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China. ZooKeys, 983, 1 - 442. https: // doi. org / 10.3897 / zookeys. 983.52630" type="journal article" year="2020">
Smith
<emphasis id="9BA76F0F6E533F0AFCE64836FCABCA7E" box="[797,855,1339,1365]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2020
</bibRefCitation>
); outer margin of protibia with 5 denticles vs 4 denticles; more gradual declivital slope, declivity occupying 1/2 of elytra vs. 2/5 of elytra; and interstrial setae shorter, as long as the distance between suture and striae 2 vs. longer, as long as the distance between the suture and striae 3.
</paragraph>
</subSubSection>
</treatment>
</document>