treatments-xml/data/21/7A/02/217A020B6E503F09FF6C4D9AFF31C97A.xml
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<document id="F21B92D01A0D2A427AD519F495F3E87D" ID-CLB-Dataset="115950" ID-DOI="10.11646/zootaxa.5174.1.1" ID-GBIF-Dataset="c4bf1061-5af8-4f4a-b4ca-234b6d0ca615" ID-ISSN="1175-5326" ID-Zenodo-Dep="6972960" ID-ZooBank="F80F144B-D1E8-4587-A146-0BACFFE18FB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659936256232" checkinUser="plazi" docAuthor="Beaver, Roger A. &amp; Smith, Sarah M." docDate="2022" docId="217A020B6E503F09FF6C4D9AFF31C97A" docLanguage="en" docName="zootaxa.5174.1.1.pdf" docOrigin="Zootaxa 5174 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euwallacea insolitus Smith &amp; Beaver 2022, sp. nov." docType="treatment" docUuid="C268A443-EE7A-4892-BC62-BA81618D361A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="5" masterDocId="DD437A736E543F0DFFFB4D0DFFFCCF2B" masterDocTitle="The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae" masterLastPageNumber="24" masterPageNumber="1" pageNumber="5" updateTime="1699365110226" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DAC3417DC5C228FABEF180FEAB5E4710">The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae</mods:title>
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<mods:name id="72D26C76E15C8A3880552DA886551142" type="personal">
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<mods:namePart id="F4B6B3F7D31440CF42C11F80ED5D22E3">Beaver, Roger A.</mods:namePart>
<mods:affiliation id="73EEF8E66FA7AFC07133D4E9FF96AE50">161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
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<mods:namePart id="5FD5BDE89A6E16A3CBEBB8821BB31650">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="C4DC7045A553364068F1D660DC276605" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="2BC8ED6ACBB2AF52245B4AAB2071F135">Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
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<mods:title id="D2B945D88946EB413F9F9F1C502818D7">Zootaxa</mods:title>
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<mods:date id="F71B228ACA4D74039C9231D19C405B79">2022</mods:date>
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<mods:number id="B7C3C2B80780FF1D6DEC3842E8463DD3">2022-08-08</mods:number>
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<treatment id="217A020B6E503F09FF6C4D9AFF31C97A" ID-DOI="http://doi.org/10.5281/zenodo.6972968" ID-GBIF-Taxon="198633506" ID-Zenodo-Dep="6972968" LSID="urn:lsid:zoobank.org:act:C268A443-EE7A-4892-BC62-BA81618D361A" httpUri="http://treatment.plazi.org/id/217A020B6E503F09FF6C4D9AFF31C97A" lastPageNumber="5" pageId="4" pageNumber="5">
<subSubSection id="E1C9E0966E503F09FF6C4D9AFD35CF9A" box="[151,713,151,178]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="A96CB31D6E503F09FF6C4D9AFD35CF9A" blockId="4.[151,1185,151,249]" box="[151,713,151,178]" pageId="4" pageNumber="5">
<heading id="F22404716E503F09FF6C4D9AFD35CF9A" bold="true" box="[151,713,151,178]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="9BA76F0F6E503F09FF6C4D9AFD35CF9A" bold="true" box="[151,713,151,178]" pageId="4" pageNumber="5">
<taxonomicName id="6ED3C89E6E503F09FF6C4D9AFD9FCF9A" authority="Smith &amp; Beaver" authorityName="Smith &amp; Beaver" authorityYear="2022" box="[151,611,151,178]" class="Insecta" family="Curculionidae" genus="Euwallacea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="insolitus" status="sp. nov.">
<emphasis id="9BA76F0F6E503F09FF6C4D9AFE6FCF99" bold="true" box="[151,403,151,178]" italics="true" pageId="4" pageNumber="5">Euwallacea insolitus</emphasis>
Smith &amp; Beaver
</taxonomicName>
<taxonomicNameLabel id="8094D2746E503F09FD904D9AFD35CF9A" box="[619,713,151,177]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="E1C9E0966E503F09FF6C4DB6FB5DCFD2" pageId="4" pageNumber="5" type="description">
<paragraph id="A96CB31D6E503F09FF6C4DB6FEF7CFFD" blockId="4.[151,1185,151,249]" box="[151,267,187,214]" pageId="4" pageNumber="5">
(
<figureCitation id="31E8AF986E503F09FF644DB6FEFFCFFD" box="[159,259,187,214]" captionStart="FIGURES 58" captionStartId="5.[151,264,1991,2015]" captionTargetBox="[163,1428,181,1966]" captionTargetId="figure-17@5.[160,1428,181,1967]" captionTargetPageId="5" captionText="FIGURES 58. Euwallacea insolitus holotype, 2.9 mm. Dorsal, lateral, posterior oblique and anterior views." figureDoi="http://doi.org/10.5281/zenodo.6972966" httpUri="https://zenodo.org/record/6972966/files/figure.png" pageId="4" pageNumber="5">Figs 58</figureCitation>
)
</paragraph>
<paragraph id="A96CB31D6E503F09FF6C4DD2FB5DCFD2" blockId="4.[151,1185,151,249]" box="[151,1185,223,249]" pageId="4" pageNumber="5">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="DD7589C86E503F09FD7B4DD2FB5DCFD2" box="[640,1185,223,249]" pageId="4" pageNumber="5">C268A443-EE7A-4892-BC62-BA81618D361A</uuid>
</uri>
</paragraph>
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<subSubSection id="E1C9E0966E503F09FF6C4C2AFDA7CE4E" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="A96CB31D6E503F09FF6C4C2AFDA7CE4E" blockId="4.[151,1437,295,1617]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF6C4C2AFEBBCE6A" bold="true" box="[151,327,295,321]" pageId="4" pageNumber="5">Type material.</emphasis>
<materialsCitation id="19BBB9406E503F09FEAB4C2AFDA7CE4E" ID-GBIF-Occurrence="3880946310" accessionNumber="A17491" collectionCode="TDRI, CIE, NHML" collectorName="A. J, D. E. Padgham" country="Bhutan" location="Gedu" pageId="4" pageNumber="5" specimenCode="NHMUK014591818" specimenCount="1" typeStatus="holotype">
<typeStatus id="76680DBF6E503F09FEAB4C2AFE47CE6A" box="[336,443,295,321]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="D1C4F38D6E503F09FDDD4C2AFD86CE6A" box="[550,634,295,321]" name="Bhutan" pageId="4" pageNumber="5">Bhutan</collectingCountry>
,
<location id="AC0CE5C66E503F09FD7C4C2AFD3ACE6A" LSID="urn:lsid:plazi:treatment:217A020B6E503F09FF6C4D9AFF31C97A:AC0CE5C66E503F09FD7C4C2AFD3ACE6A" box="[647,710,295,321]" country="Bhutan" name="Gedu" pageId="4" pageNumber="5">Gedu</location>
,
<collectionCode id="CFC22BD86E503F09FD284C25FCEBCE69" box="[723,791,296,322]" pageId="4" pageNumber="5">TDRI</collectionCode>
, 6.viii.[19]85,
<collectorName id="0426D6CB6E503F09FC444C2AFB66CE6A" box="[959,1178,295,322]" pageId="4" pageNumber="5">A.J, D.E. Padgham</collectorName>
coll., ex
<taxonomicName id="6ED3C89E6E503F09FAF34C2AFA9ACE69" box="[1288,1382,295,322]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="9BA76F0F6E503F09FAF34C2AFAC1CE6A" box="[1288,1341,295,321]" italics="true" pageId="4" pageNumber="5">Acer</emphasis>
sp.
</taxonomicName>
<collectionCode id="CFC22BD86E503F09FA944C2AFA60CE6A" box="[1391,1436,295,321]" pageId="4" pageNumber="5">CIE</collectionCode>
A17491,
<specimenCode id="F9751B666E503F09FF044C46FE1BCE4D" box="[255,487,331,358]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="5" type="Museum">NHMUK014591818</specimenCode>
(
<collectionCode id="CFC22BD86E503F09FE0D4C46FDB3CE4E" box="[502,591,331,357]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="5">NHML</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C4C62FD72CEDD" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="A96CB31D6E503F09FF3C4C62FCB7CEFA" blockId="4.[151,1437,295,1617]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4C62FEBECEA2" bold="true" box="[199,322,367,393]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
<quantity id="6E2B1EF86E503F09FEBC4C62FE61CEA1" box="[327,413,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="mm" value="2.9">2.9 mm</quantity>
long (n = 1); 2.42 × as long as wide. This species is distinguished by its pronotum conical in dorsal view (
<typeStatus id="76680DBF6E503F09FEDE4C9EFEABCE86" box="[293,343,403,429]" pageId="4" pageNumber="5">type</typeStatus>
0); obliquely truncate antennal club with two sutures on the posterior face (
<typeStatus id="76680DBF6E503F09FB634C9EFB36CE86" box="[1176,1226,403,429]" pageId="4" pageNumber="5">type</typeStatus>
2); elytra unarmed by tubercles and granules, and posterolateral margin rounded.
</paragraph>
<paragraph id="A96CB31D6E503F09FF3C4CD6FD72CEDD" blockId="4.[151,1437,295,1617]" box="[199,654,475,502]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4CD6FE7CCEDE" bold="true" box="[199,384,475,501]" pageId="4" pageNumber="5">Similar species.</emphasis>
<taxonomicName id="6ED3C89E6E503F09FE7D4CD6FD72CEDD" box="[390,654,475,502]" class="Insecta" family="Curculionidae" genus="Anisandrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9BA76F0F6E503F09FE7D4CD6FDFACEDE" box="[390,518,475,501]" italics="true" pageId="4" pageNumber="5">Anisandrus</emphasis>
Ferrari spp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C4CF2FBFBCA26" pageId="4" pageNumber="5" type="description">
<paragraph id="A96CB31D6E503F09FF3C4CF2FBFBCA26" blockId="4.[151,1437,295,1617]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4CF2FE3CCD32" bold="true" box="[199,448,511,537]" pageId="4" pageNumber="5">Description (female).</emphasis>
<quantity id="6E2B1EF86E503F09FE3D4CF2FDE6CD31" box="[454,538,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="mm" value="2.9">2.9 mm</quantity>
long (n = 1); 2.42 × as long as wide. Body dark brown. Legs and antennae brown.
<emphasis id="9BA76F0F6E503F09FF6C4F2EFF1CCD16" bold="true" box="[151,224,547,573]" pageId="4" pageNumber="5">Head:</emphasis>
epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface alutaceous, sparsely punctate, setose; punctures bearing a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum broadly triangular, slightly impressed. Antennal scape regularly thick. Funicle 4-segmented. Club longer than wide, obliquely truncate,
<typeStatus id="76680DBF6E503F09FB2D4F9DFAFACD81" box="[1238,1286,656,682]" pageId="4" pageNumber="5">type</typeStatus>
2; segment 1 corneous, transverse on anterior face, occupying basal half; segment 2 narrow, corneous; segments 1 and 2 present on posterior face.
<emphasis id="9BA76F0F6E503F09FEA64FDAFE23CDDA" bold="true" box="[349,479,727,753]" pageId="4" pageNumber="5">Pronotum:</emphasis>
0.92 × as long as wide. In dorsal view conical,
<typeStatus id="76680DBF6E503F09FC1D4FD5FBEACDD9" box="[998,1046,728,754]" pageId="4" pageNumber="5">type</typeStatus>
0, sides convex, conical anteriorly; anterior margin without serrations. In lateral view tall,
<typeStatus id="76680DBF6E503F09FCF24FF1FCC5CC3D" box="[777,825,764,790]" pageId="4" pageNumber="5">type</typeStatus>
2, disc flat, summit at apical 2/5. Anterior slope with dense low, broad asperities, becoming lower and more strongly transverse towards summit. Disc shagreened with moderately dense, minute punctures, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded.
<emphasis id="9BA76F0F6E503F09FD754E6AFD18CCAA" bold="true" box="[654,740,871,897]" pageId="4" pageNumber="5">Elytra:</emphasis>
1.50 × as long as wide, 1.64 × as long as pronotum. Scutellum moderately sized, linguiform, shining, flush with elytra, flat. Elytral base transverse, edge oblique and unarmed by granules, humeral angles rounded, parallel-sided in basal 3/5, then broadly rounded to apex. Striae and interstriae flush, unarmed by tubercles or granules; interstriae opalescent. Disc flat, striae not impressed, with large, deep punctures separated by 1.52 diameters of a puncture, setose; each puncture bearing a semi-erect hair-like seta, increasing in length towards declivity; interstriae flat, punctures subequal to strial punctures, each bearing an erect seta longer than the width of an interstria. Declivital slope very gradual, occupying approximately 1/3 of elytra, declivital face weakly convex; strial punctures very large, shallow, much larger and deeper than those of disc, setose, setae short, as long as strial punctures; interstriae sparsely, minutely punctate, setose, setae long, erect. Posterolateral margin rounded.
<emphasis id="9BA76F0F6E503F09FEA049A6FE60CBEE" bold="true" box="[347,412,1195,1221]" pageId="4" pageNumber="5">Legs:</emphasis>
protibiae distinctly triangular, broadest at apical 1/4; posterior face smooth; apical 1/2 of outer margin with 7 moderate socketed denticles, their length as long as basal width. Meso- and metatibiae flattened; outer margin evenly rounded with 8 moderate socketed denticles, respectively.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C481AFE3CCA1A" box="[199,448,1303,1329]" pageId="4" pageNumber="5" type="distribution">
<paragraph id="A96CB31D6E503F09FF3C481AFE3CCA1A" blockId="4.[151,1437,295,1617]" box="[199,448,1303,1329]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C481AFE9DCA1A" bold="true" box="[199,353,1303,1329]" pageId="4" pageNumber="5">Distribution.</emphasis>
<collectingCountry id="D1C4F38D6E503F09FE9C481AFE40CA1A" box="[359,444,1303,1329]" name="Bhutan" pageId="4" pageNumber="5">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C4836FDC3CA7E" box="[199,575,1339,1365]" pageId="4" pageNumber="5" type="biology_ecology">
<paragraph id="A96CB31D6E503F09FF3C4836FDC3CA7E" blockId="4.[151,1437,295,1617]" box="[199,575,1339,1365]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4836FEAACA7E" bold="true" box="[199,342,1339,1365]" pageId="4" pageNumber="5">
<collectingRegion id="6B177DFF6E503F09FF3C4836FF03CA7E" box="[199,255,1339,1365]" country="Afghanistan" name="Khost" pageId="4" pageNumber="5">Host</collectingRegion>
plants.
</emphasis>
<taxonomicName id="6ED3C89E6E503F09FEA64836FDC0CA7E" box="[349,572,1339,1365]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis id="9BA76F0F6E503F09FEA64836FE6ECA7E" box="[349,402,1339,1365]" italics="true" pageId="4" pageNumber="5">Acer</emphasis>
(Sapindaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C4852FDACCAB6" pageId="4" pageNumber="5" type="etymology">
<paragraph id="A96CB31D6E503F09FF3C4852FDACCAB6" blockId="4.[151,1437,295,1617]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C4852FEB2CA52" bold="true" box="[199,334,1375,1401]" pageId="4" pageNumber="5">Etymology.</emphasis>
L.
<taxonomicName id="6ED3C89E6E503F09FE894852FE2DCA52" box="[370,465,1375,1401]" class="Insecta" family="Curculionidae" genus="Euwallacea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="9BA76F0F6E503F09FE894852FE2DCA52" box="[370,465,1375,1401]" italics="true" pageId="4" pageNumber="5">insolitus</emphasis>
</taxonomicName>
= unusual, strange. An adjective. In reference to the unusual combination of characters exhibited by this species (see remarks.)
</paragraph>
</subSubSection>
<subSubSection id="E1C9E0966E503F09FF3C48AAFF31C97A" pageId="4" pageNumber="5" type="discussion">
<paragraph id="A96CB31D6E503F09FF3C48AAFF31C97A" blockId="4.[151,1437,295,1617]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FF3C48AAFEC7CAEA" bold="true" box="[199,315,1447,1473]" pageId="4" pageNumber="5">Remarks.</emphasis>
This is a rather unique
<taxonomicName id="6ED3C89E6E503F09FDBB48AAFCE0CAEA" box="[576,796,1447,1473]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FDBB48AAFD38CAEA" box="[576,708,1447,1473]" italics="true" pageId="4" pageNumber="5">Euwallacea</emphasis>
species
</taxonomicName>
with an idiosyncratic combination of characters. Overall, the species is rather similar to
<taxonomicName id="6ED3C89E6E503F09FE1748C6FD3FCACE" box="[492,707,1483,1509]" pageId="4" pageNumber="5">
<emphasis id="9BA76F0F6E503F09FE1748C6FD90CACE" box="[492,620,1483,1509]" italics="true" pageId="4" pageNumber="5">Anisandrus</emphasis>
species
</taxonomicName>
which often have a similar
<taxonomicName id="6ED3C89E6E503F09FC0048C6FB96CACE" box="[1019,1130,1483,1509]" form="with" pageId="4" pageNumber="5" rank="form">form with</taxonomicName>
a conical
<typeStatus id="76680DBF6E503F09FB2748C1FAF0CACD" box="[1244,1292,1484,1510]" pageId="4" pageNumber="5">type</typeStatus>
0 pronotum, gradual elytral declivity, rounded posterolateral elytra margin, and unarmed interstriae. It can be readily distinguished from
<taxonomicName id="6ED3C89E6E503F09FF2F4B1EFEA8C906" box="[212,340,1555,1581]" class="Insecta" family="Curculionidae" genus="Anisandrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="9BA76F0F6E503F09FF2F4B1EFEA8C906" box="[212,340,1555,1581]" italics="true" pageId="4" pageNumber="5">Anisandrus</emphasis>
</taxonomicName>
by the lack of a pronotal mycangial tuft and
<typeStatus id="76680DBF6E503F09FCBC4B19FC8BC905" box="[839,887,1556,1582]" pageId="4" pageNumber="5">type</typeStatus>
2 antennal club with two sutures on the posterior face.
</paragraph>
</subSubSection>
</treatment>
</document>