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<document ID-DOI="http://doi.org/10.5281/zenodo.3792654" ID-GBIF-Dataset="cdc50aed-51ca-47e9-adbf-a4f2710c5114" ID-GBIF-Taxon="163748658" ID-ISSN="13132970" ID-Zenodo-Dep="3792654" ID-ZooBank="695276C3-BA8B-4BCF-9754-A86CC7B1C77D" approvalRequired="3" approvalRequired_for_matCits="2" approvalRequired_for_treatments="1" checkinTime="1587961194007" checkinUser="plazi" docAuthor="Engel, Michael &amp; Ortega-Blanco, Jaime" docDate="2008" docId="0604785CFFDAFFBDFF56E0B1E9E46DBC" docLanguage="en" docName="ZK_article_1957.pdf" docOrigin="ZooKeys 4 (4)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@68ab6cce" docStyleName="zookeys.2008.journal_article" docTitle="Electrostephanus (Electrostephanodes) petiolatus Brues 2008, combinatio revivisco" docType="treatment" docVersion="8" lastPageId="5" lastPageNumber="60" masterDocId="FA3D0024FFD8FFB8FFDAE412E86B6C66" masterDocTitle="The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae" masterLastPageNumber="64" masterPageNumber="55" pageId="2" pageNumber="57" updateTime="1636521069113" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael</mods:namePart>
<mods:affiliation>University of Kansas, Natural History Museum, Lawrence, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortega-Blanco, Jaime</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Departament dEstratigrafia, Paleontologia i Geociències Marines, Facultat de Geologia, Universitat de Barcelona, Barcelona, Spain &amp; F 597 B 757 &amp; F- 921 F- 47085 A 576641</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-12-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>55</mods:start>
<mods:end>64</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.4.49</mods:identifier>
<mods:identifier type="GBIF-Dataset">cdc50aed-51ca-47e9-adbf-a4f2710c5114</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576424</mods:identifier>
<mods:identifier type="ZooBank">695276C3-BA8B-4BCF-9754-A86CC7B1C77D</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792654" ID-GBIF-Taxon="163748658" ID-Zenodo-Dep="3792654" LSID="urn:lsid:plazi:treatment:0604785CFFDAFFBDFF56E0B1E9E46DBC" httpUri="http://treatment.plazi.org/id/0604785CFFDAFFBDFF56E0B1E9E46DBC" lastPageId="5" lastPageNumber="60" pageId="2" pageNumber="57">
<subSubSection box="[140,1048,1187,1214]" pageId="2" pageNumber="57" type="nomenclature">
<paragraph blockId="2.[140,1048,1187,1214]" box="[140,1048,1187,1214]" pageId="2" pageNumber="57">
<heading box="[140,1048,1187,1214]" fontSize="11" level="2" pageId="2" pageNumber="57" reason="3">
<emphasis bold="true" box="[140,1048,1187,1214]" pageId="2" pageNumber="57">
<taxonomicName authority="Brues" authorityName="Brues" authorityYear="2008" box="[140,792,1187,1214]" class="Insecta" family="Stephanidae" genus="Electrostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="petiolatus" status="combinatio revivisco" subGenus="Electrostephanodes">
<emphasis bold="true" box="[140,715,1187,1214]" italics="true" pageId="2" pageNumber="57">Electrostephanus (Electrostephanodes) petiolatus</emphasis>
Brues
</taxonomicName>
,
<emphasis bold="true" box="[804,1048,1187,1214]" italics="true" pageId="2" pageNumber="57">
<taxonomicNameLabel box="[804,1048,1187,1214]" pageId="2" pageNumber="57">combinatio revivisco</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="57" type="reference_group">
<paragraph blockId="2.[140,1108,1258,1460]" pageId="2" pageNumber="57">
<treatmentCitationGroup pageId="2" pageNumber="57">
<treatmentCitation author="Brues CT" box="[140,609,1258,1284]" page="14" pageId="2" pageNumber="57" year="1933">
<taxonomicName authority="Brues, 1933: 14" authorityName="Brues" authorityPageNumber="14" authorityYear="1933" box="[140,609,1258,1284]" class="Insecta" family="Stephanidae" genus="Electrostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="petiolatus">
<emphasis box="[140,422,1258,1284]" italics="true" pageId="2" pageNumber="57">Electrostephanus petiolatus</emphasis>
<bibRefCitation author="Brues CT" box="[430,609,1258,1284]" pageId="2" pageNumber="57" pagination="4 - 178" refId="ref4343" refString="Brues CT (1933) The parasitic Hymenoptera of the Baltic amber. Part 1. Bernstein Forschungen 3: 4 - 178." type="journal article" year="1933">Brues, 1933: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus box="[623,724,1258,1284]" pageId="2" pageNumber="57">holotype</typeStatus>
male, in ill-fated Königsberg Collection, presumed destroyed];
<treatmentCitation author="Aguiar AP &amp; Janzen J-W" box="[536,890,1293,1319]" page="444" pageId="2" pageNumber="57" year="1999">
<bibRefCitation author="Aguiar AP &amp; Janzen J-W" box="[536,890,1293,1319]" pageId="2" pageNumber="57" pagination="443 - 452" refId="ref4259" refString="Aguiar AP, Janzen J-W (1999) An overview of fossil Stephanidae (Hymenoptera), with description of two new taxa from Baltic amber, and key to species of Electrostephanus Brues. Entomologica Scandinavica 30 (4): 443 - 452. [Publication date: January 2000]" type="journal article" year="1999">Aguiar and Janzen, 1999: 444</bibRefCitation>
</treatmentCitation>
[key];
<treatmentCitation author="Achterberg C van" page="12" pageId="2" pageNumber="57" year="2002">
<bibRefCitation author="Achterberg C van" pageId="2" pageNumber="57" pagination="1 - 206" refId="ref4186" refString="Achterberg C van (2002) A revision of the Old World species of Megischus Brulle, Stephanus Jurine and Pseudomegischus gen. nov., with a key to the genera of the family Stephanidae (Hymenoptera: Stephanoidea). Zoologische Verhandelingen 339: 1 - 206." type="journal article" year="2002">Achterberg, 2002: 12</bibRefCitation>
</treatmentCitation>
[mentioned in
<taxonomicName authorityName="Orfila" authorityYear="1949" box="[462,640,1327,1354]" class="Insecta" family="Stephanidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Schlettereriinae">Schlettereriinae</taxonomicName>
];
<treatmentCitation author="Aguiar AP" box="[657,846,1328,1354]" page="14" pageId="2" pageNumber="57" year="2004">
<bibRefCitation author="Aguiar AP" box="[657,846,1328,1354]" pageId="2" pageNumber="57" pagination="1 - 120" refId="ref4236" refString="Aguiar AP (2004) World catalog of the Stephanidae (Hymenoptera: Stephanoidea). Zootaxa 753: 1 - 120." type="journal article" year="2004">Aguiar, 2004: 14</bibRefCitation>
</treatmentCitation>
[catalogue].
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[140,1108,1258,1460]" pageId="2" pageNumber="57">
<treatmentCitationGroup pageId="2" pageNumber="57">
<taxonomicName authority="(Brues)" baseAuthorityName="Brues" baseAuthorityYear="1933" box="[140,521,1363,1389]" class="Insecta" family="Stephanidae" genus="Denaeostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="petiolatus">
<emphasis box="[140,429,1363,1389]" italics="true" pageId="2" pageNumber="57">Denaeostephanus petiolatus</emphasis>
(Brues)
</taxonomicName>
,
<treatmentCitation author="Engel MS &amp; Grimaldi DA" box="[534,903,1363,1389]" page="1194" pageId="2" pageNumber="57" year="2004">
<bibRefCitation author="Engel MS &amp; Grimaldi DA" box="[534,903,1363,1389]" pageId="2" pageNumber="57" pagination="1192 - 1197" refId="ref4432" refString="Engel MS, Grimaldi DA (2004) The first Mesozoic stephanid wasp (Hymenoptera: Stephanidae). Journal of Paleontology 78 (6): 1192 - 1197." type="journal article" year="2004">Engel and Grimaldi, 2004: 1194</bibRefCitation>
</treatmentCitation>
[tentative transfer to genus based on description of petiole by
<bibRefCitation author="Aguiar AP &amp; Janzen J-W" box="[660,948,1398,1425]" pageId="2" pageNumber="57" pagination="443 - 452" refId="ref4259" refString="Aguiar AP, Janzen J-W (1999) An overview of fossil Stephanidae (Hymenoptera), with description of two new taxa from Baltic amber, and key to species of Electrostephanus Brues. Entomologica Scandinavica 30 (4): 443 - 452. [Publication date: January 2000]" type="journal article" year="1999">Aguiar and Janzen (1999)</bibRefCitation>
];
<treatmentCitation author="Engel MS" page="318" pageId="2" pageNumber="57" year="2005">
<bibRefCitation author="Engel MS" pageId="2" pageNumber="57" pagination="317 - 332" refId="ref4395" refString="Engel MS (2005) The crown wasp genus Electrostephanus (Hymenoptera: Stephanidae): Discovery of the female and a new species. Polskie Pismo Entomologiczne 74 (3): 317 - 332." type="journal article" year="2005">Engel, 2005: 318</bibRefCitation>
</treatmentCitation>
[mentioned].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="57" type="materials_examined">
<paragraph blockId="2.[140,1108,1504,1671]" pageId="2" pageNumber="57">
<emphasis bold="true" box="[140,320,1504,1530]" pageId="2" pageNumber="57">
<emphasis box="[140,200,1504,1530]" inLineHeading="true" pageId="2" pageNumber="57" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608819270" collectionCode="AMNH" collectorName="Baltic &amp; Eocene &amp; Lutetian &amp; Formerly &amp; Jens-Wilhelm Janzen" pageId="2" pageNumber="57" specimenCode="B-JWJ-260" specimenCount="2" typeStatus="neotype">
<typeStatus box="[328,428,1505,1531]" pageId="2" pageNumber="57">Neotype</typeStatus>
(
<emphasis bold="true" box="[447,642,1504,1530]" pageId="2" pageNumber="57">here designated</emphasis>
) male;
<collectionCode box="[732,826,1504,1530]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="57">AMNH</collectionCode>
<specimenCode box="[835,972,1504,1530]" collectionCode="B-JWJ-" pageId="2" pageNumber="57">B-JWJ-260</specimenCode>
,
<collectorName box="[985,1053,1504,1530]" pageId="2" pageNumber="57">Baltic</collectorName>
amber,
<collectorName box="[196,281,1540,1566]" pageId="2" pageNumber="57">Eocene</collectorName>
(
<collectorName box="[301,408,1539,1566]" pageId="2" pageNumber="57">Lutetian</collectorName>
), and labeled “
<typeStatus box="[593,699,1539,1565]" pageId="2" pageNumber="57">Neotype</typeStatus>
,
<taxonomicName authority="Brues" authorityName="Brues" box="[716,1103,1539,1566]" class="Insecta" family="Stephanidae" genus="Electrostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="petiolatus">
<emphasis box="[716,1021,1539,1565]" italics="true" pageId="2" pageNumber="57">Electrostephanus petiolatus</emphasis>
Brues
</taxonomicName>
, desig. Engel &amp; Ortega-Blanco [red label]”.
<collectorName box="[681,789,1574,1600]" pageId="2" pageNumber="57">Formerly</collectorName>
part of the
<collectorName pageId="2" pageNumber="57">Jens-Wilhelm Janzen</collectorName>
collection.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[185,684,1645,1671]" pageId="2" pageNumber="57" type="diagnosis">
<paragraph blockId="2.[140,1108,1504,1671]" box="[185,684,1645,1671]" pageId="2" pageNumber="57">
<emphasis bold="true" box="[185,312,1645,1671]" pageId="2" pageNumber="57">
<emphasis box="[185,307,1645,1671]" inLineHeading="true" pageId="2" pageNumber="57" reason="1">Diagnosis</emphasis>
.
</emphasis>
As for the subgenus (
<emphasis box="[558,668,1645,1671]" italics="true" pageId="2" pageNumber="57">vide supra</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58" type="description">
<paragraph blockId="3.[140,1108,166,903]" pageId="3" pageNumber="58">
<emphasis bold="true" box="[185,335,166,192]" pageId="3" pageNumber="58">
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="3" pageNumber="58" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[345,401,166,192]" italics="true" pageId="3" pageNumber="58">Male</emphasis>
: Total body length (from head anterior margin to metasoma posterior margin)
<quantity box="[355,467,202,228]" metricMagnitude="-3" metricUnit="m" metricValue="9.84" pageId="3" pageNumber="58" unit="mm" value="9.84">9.84 mm</quantity>
; forewing length (from tegula to apex)
<quantity box="[944,1057,201,228]" metricMagnitude="-3" metricUnit="m" metricValue="5.04" pageId="3" pageNumber="58" unit="mm" value="5.04">5.04 mm</quantity>
. Integument dark brown to black (where evident) with scattered, thin, simple, erect or suberect setae as noted. Head globular, with compound eyes occupying around onehalf of lateral surface, eyes well separated from preoccipital area; “crown” composed of 5 tubercles arising anterior to series of four transverse carinae between lateral ocelli around compound eyes, median ocellus set just anterior to series of carinae; tubercles arranged with anteriormost tubercle medial on frons, lateral tubercles paired in longitudinal series parallel to compound eye; integument of face rugulose, integument posterior to carinae and on gena impunctate and smooth. Antennae with 23 articles, arising from clypeus boundary at about compound eye midlength; scape about twice as long as wide, somewhat ovoid, surface bordering malar space slightly flattened; pedicel about one-half scape length, almost as wide as long; first flagellomere as long as pedicel but one-half width; second flagellomere around four times longer than wide, slightly less than twice length of first flagellomere; third through fifth subequal in length and shape; remaining flagellomeres progressively shorter, except apicalmost flagellomere with tapered apex. Maxillary palpus 5-segmented, elongate, elbowed between MP2 (maxillary palpomere II) and MP3, MP1 and MP2 thicker than MP35, MP1 shortest, MP
<subScript attach="left" box="[183,192,819,833]" fontSize="6" pageId="3" pageNumber="58">2</subScript>
about twice length of MP
<subScript attach="left" box="[489,498,819,833]" fontSize="6" pageId="3" pageNumber="58">1</subScript>
, MP
<subScript attach="left" box="[556,565,819,833]" fontSize="6" pageId="3" pageNumber="58">3</subScript>
slightly less than twice length of MP
<subScript attach="left" box="[983,992,819,833]" fontSize="6" pageId="3" pageNumber="58">2</subScript>
, MP
<subScript attach="left" box="[1049,1058,819,833]" fontSize="6" pageId="3" pageNumber="58">4</subScript>
and MP
<subScript attach="left" box="[183,192,854,868]" fontSize="6" pageId="3" pageNumber="58">5</subScript>
equal in length to MP
<subScript attach="left" box="[459,468,854,868]" fontSize="6" pageId="3" pageNumber="58">3</subScript>
. Labial palpus short, apparently 3-segmented (base obscured), slightly widening apically except LP
<subScript attach="left" box="[641,650,889,903]" fontSize="6" pageId="3" pageNumber="58">3</subScript>
with acutely pointed apex.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3784816" ID-Zenodo-Dep="3784816" box="[140,1025,1654,1676]" httpUri="https://zenodo.org/record/3784816/files/figure.png" pageId="3" pageNumber="58" startId="3.[140,174,1654,1676]" targetBox="[140,1108,950,1634]" targetPageId="3">
<paragraph blockId="3.[140,1025,1654,1676]" box="[140,1025,1654,1676]" pageId="3" pageNumber="58">
<emphasis bold="true" box="[140,204,1654,1676]" pageId="3" pageNumber="58">Fig. 1.</emphasis>
<typeStatus box="[210,289,1655,1676]" pageId="3" pageNumber="58">Neotype</typeStatus>
male of
<taxonomicName authority="Brues in Baltic amber" authorityName="Brues in Baltic Amber" box="[370,810,1654,1676]" class="Insecta" family="Stephanidae" genus="Electrostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="petiolatus">
<emphasis box="[370,604,1654,1675]" italics="true" pageId="3" pageNumber="58">Electrostephanus petiolatus</emphasis>
Brues in Baltic amber
</taxonomicName>
(
<collectionCode box="[822,898,1654,1675]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="58">AMNH</collectionCode>
<specimenCode box="[905,1014,1654,1676]" collectionCode="B-JWJ-" pageId="3" pageNumber="58">B-JWJ-260</specimenCode>
).
</paragraph>
</caption>
<paragraph blockId="4.[140,1108,166,1671]" pageId="4" pageNumber="59">Pronotum long, transversely striate along neck, near articulation with head, remainder of surface imbricate and irregular, laterally and dorsoventrally micro-rugulose, posterior portion with sparse, erect, long, simple setae; propleuron finely imbricate, with scattered shallow punctures. Mesoscutum, mesoscutellum, and metanotum apparently largely imbricate, with scattered shallow punctures (direct view of this surface slightly obscured), with sparse, erect, long, simple setae; pleura largely coarsely and irregularly punctured, punctures deep, large, and nearly contiguous, forming irregular network, except posterior half of mesopleuron with large, impunctate and imbricate area bordered posteriorly by dorsoventral column of coarse, large, punctures along boundary with metapleuron; metapleuron with longitudinal series of irregular rugae, posteriorly with coarse, irregular, contiguous punctures like those on anterior portion of mesopleuron except more deeply impressed.</paragraph>
<paragraph blockId="4.[140,1108,166,1671]" pageId="4" pageNumber="59">Pro- and mesocoxae short and cylindrical, imbricate; metacoxa massive, wider at base, oriented posteriorly, imbricate except basally somewhat rugose; trochanters narrow at base, gently and slightly widening apically, with distinct trochantellus; femora finely imbricate; pro- and mesofemora with proximal thirds narrowly petiolate; metafemur fusiform, widest at mid-point; ventral surface of metafemur with a blunt, triangular, principal tooth at first third of length, a more acute and longer tooth near midlength, and a tooth near apex in apical third of length; three minor teeth or protuberances between medial tooth and others, such protuberances slightly closer to medial tooth, another protuberance just prior to distalmost principal tooth; a thin, long seta arising from each protuberance; metafemur with scattered, erect, long, slightly fuscous setae; tibiae finely imbricate, thin and elongate, with nearly basal halves narrowly petiolate, with a row of short spines on anterior and posterior surfaces of pro- and mesotibia; metatibia with several long setae at apex, inner surface with thin patch of microtrichia; tibial spurs 1-2-2, stout and short; metabasitarsus with a row of stouter setae anteriorly and posteriorly; remaining tarsomeres with two distinct long, thick distal setae; tarsal relative proportions (all in comparison to associated basitarsus = 1) foreleg: 1: 0.5: 0.3: 0.3: 0.4; midlegs: 1: 0.4: 0.4: 0.3: 0.4; hind legs: 1: 0.4: 0.25: 0.8: 0.5; fourth tarsomere with distal ventral surface elongate, extending under most of fifth tarsomere; fifth tarsomere arising from near midlength of fourth tarsomere; pretarsal ungues (= claws) long, curved, and simple.</paragraph>
<paragraph blockId="4.[140,1108,166,1671]" pageId="4" pageNumber="59">
Wing membranes hyaline. Forewing with parallel-sided, dark brown pterostigma, slightly more than three times longer than high; Rs apex not reaching completely wing margin but extending well beyond pterostigma, demarcating a long, open marginal cell; basal vein (first free abscissa of M) arched proximally along basal half, about three times as long as first free abscissa of Rs, about twice 1m-cu length and approximately parallel to 1m-cu, demarcating a trapezoidal medial cell (= discal cell); Rs+M
<subScript attach="left" box="[997,1006,1417,1431]" fontSize="6" pageId="4" pageNumber="59">b</subScript>
and bullae absent; submarginal cell pentagonal, wider apically, r-rs slightly less than one-half length of second free abscissa Rs; cubital cell rectangular, three times longer than high; M and Cu almost reaching wing margin as nebulous veins; 3A, 2cu-a, and apical third of 2A nebulous; posterior margin of wing with very short, thin setae. Hind wing with four distal hamuli; only Sc+R present, thin and tubular.
</paragraph>
<paragraph blockId="4.[140,1108,166,1671]" lastBlockId="5.[140,1108,166,475]" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
Metasoma slender, elongate, terga and sterna not fused laterally, integument finely imbricate except pseudo-petiole rugulose, sternum with irregular transverse rugae ba-sally; first metasomal tergum and sternum forming a narrow, tubular pseudo-petiole (a “true” petiole in
<taxonomicName authorityName="Leach" authorityYear="1815" box="[362,501,202,228]" class="Insecta" family="Stephanidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="family">Stephanidae</taxonomicName>
have the tergum and sternum fused laterally), about five times longer than wide; metasomal segments IIV subequal in length, remaining terga progressively shorter, second and third metasomal segments about three times longer than wide; gastral terga with exceptionally sparse setae, setae suberect, simple and long; gastral sterna with sparse, erect or suberect, elongate setae; parameres exposed, broad, tapering gradually to bluntly rounded apices, with fringe of dense, erect, moderate-length, slightly fuscous setae at apex.
</paragraph>
<paragraph blockId="5.[140,1108,166,475]" box="[185,399,448,475]" pageId="5" pageNumber="60">
<emphasis box="[185,261,448,474]" italics="true" pageId="5" pageNumber="60">Female</emphasis>
: Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>