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<document id="BF531C5924A733095DCA0C9BEB616DC0" ID-DOI="10.5281/zenodo.281575" ID-GBIF-Dataset="20855354-eb05-401e-96a9-8761ea097506" ID-ISSN="1175-5326" ID-Zenodo-Dep="281575" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460493090668" checkinUser="plazi" docAuthor="Müller, Andreas" docDate="2012" docId="03C0BF77FFBAFFAD1DB1FADF1EEDFC85" docLanguage="en" docName="zt03355p050.pdf" docOrigin="Zootaxa 3355" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Chelostoma (Gyrodromella) aegaeicum Müller, 2012, spec. nov." docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="FFF9C70FFFBDFFA41D26FFE31D75FF91" masterDocTitle="New European bee species of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)" masterLastPageNumber="50" masterPageNumber="29" pageNumber="36" updateTime="1698310480346" updateUser="plazi">
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<mods:title id="8AD31A0B7D62D2A815C6B362150F0A12">New European bee species of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)</mods:title>
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<mods:namePart id="CF98549D86C3D5B76958902E5D96286C">Müller, Andreas</mods:namePart>
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<mods:date id="6AD4F828E0B63D1A56BE81CA18251998">2012</mods:date>
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<treatment id="03C0BF77FFBAFFAD1DB1FADF1EEDFC85" ID-DOI="http://doi.org/10.5281/zenodo.3507593" ID-GBIF-Taxon="119520195" ID-Zenodo-Dep="3507593" LSID="urn:lsid:plazi:treatment:03C0BF77FFBAFFAD1DB1FADF1EEDFC85" httpUri="http://treatment.plazi.org/id/03C0BF77FFBAFFAD1DB1FADF1EEDFC85" lastPageId="9" lastPageNumber="38" pageId="7" pageNumber="36">
<subSubSection id="C3735DEAFFBAFFA31DB1FADF1F84FAC4" box="[151,753,1339,1366]" pageId="7" pageNumber="36" type="nomenclature">
<paragraph id="8BD60E61FFBAFFA31DB1FADF1F84FAC4" blockId="7.[151,753,1339,1366]" box="[151,753,1339,1366]" pageId="7" pageNumber="36">
<heading id="D09EB90DFFBAFFA31DB1FADF1F84FAC4" bold="true" box="[151,753,1339,1366]" fontSize="11" level="1" pageId="7" pageNumber="36" reason="1">
<emphasis id="B91DD273FFBAFFA31DB1FADF1F84FAC4" bold="true" box="[151,753,1339,1366]" pageId="7" pageNumber="36">
<taxonomicName id="4C6975E2FFBAFFA31DB1FADF1F07FAC7" ID-CoL="8SFYH" box="[151,626,1340,1366]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aegaeicum" status="sp. nov." subGenus="Gyrodromella">
<emphasis id="B91DD273FFBAFFA31DB1FADF1F07FAC7" bold="true" box="[151,626,1340,1366]" italics="true" pageId="7" pageNumber="36">Chelostoma (Gyrodromella) aegaeicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFBAFFA31F5FFAD81F84FAC4" box="[633,753,1339,1365]" pageId="7" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3735DEAFFBAFFA31DB1FA611F3DF9C1" pageId="7" pageNumber="36" type="materials_examined">
<paragraph id="8BD60E61FFBAFFA31DB1FA6119B6FA09" blockId="7.[151,1437,1410,1616]" box="[151,1219,1410,1432]" pageId="7" pageNumber="36">
<typeStatus id="54D2B0C3FFBAFFA31DB1FA611D8CFA09" box="[151,249,1410,1432]" pageId="7" pageNumber="36" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F37E4EF1FFBAFFA31C24FA601C39FA09" box="[258,332,1411,1432]" name="Greece" pageId="7" pageNumber="36">Greece</collectingCountry>
,
<emphasis id="B91DD273FFBAFFA31C71FA601CD1FA09" box="[343,420,1411,1432]" italics="true" pageId="7" pageNumber="36">Lesvos:</emphasis>
<quantity id="4C91A384FFBAFFA31C8CFA611C9EFA09" box="[426,491,1410,1432]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="36" unit="km" value="3.8">3.8km</quantity>
SSE of Agiasos,
<quantity id="4C91A384FFBAFFA31FBAFA611FACFA09" box="[668,729,1410,1432]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="36" unit="m" value="760.0">760m</quantity>
,
<date id="FFD728A1FFBAFFA31FC5FA611E3EFA09" box="[739,843,1410,1432]" pageId="7" pageNumber="36" value="2004-05-24">24.5.2004</date>
, ɗ (leg. A. Kyriakopoulos), [
<collectionCode id="ED7896A4FFBAFFA31956FA6119CCFA09" box="[1136,1209,1410,1432]" httpUri="http://grbio.org/cool/3kp9-a263" name="Eidgenoessische Technische Hochschule-Zentrum" pageId="7" pageNumber="36">ETHZ</collectionCode>
].
</paragraph>
<paragraph id="8BD60E61FFBAFFA31DB1FA411F3DF9C1" blockId="7.[151,1437,1410,1616]" pageId="7" pageNumber="36">
<typeStatus id="54D2B0C3FFBAFFA31DB1FA411D8BFA26" box="[151,254,1442,1463]" pageId="7" pageNumber="36" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F37E4EF1FFBAFFA31C2DFA411C20FA26" box="[267,341,1442,1463]" name="Greece" pageId="7" pageNumber="36">Greece</collectingCountry>
,
<emphasis id="B91DD273FFBAFFA31C43FA411CD0FA26" box="[357,421,1442,1463]" italics="true" pageId="7" pageNumber="36">Crete:</emphasis>
Kato Simi,
<quantity id="4C91A384FFBAFFA31F08FA411F02FA26" box="[558,631,1442,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="36" unit="m" value="1000.0">1000m</quantity>
, 6.
<date id="FFD728A1FFBAFFA31F85FA421E78FA26" box="[675,781,1441,1463]" pageId="7" pageNumber="36" value="2003-12-05">12.5.2003</date>
, 1 Ψ, 2 ɗ (leg. Sausa); Lasithi,
<quantity id="4C91A384FFBAFFA31957FA4219DEFA26" box="[1137,1195,1441,1463]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="36" unit="km" value="15.0">15km</quantity>
NW of Males,
<quantity id="4C91A384FFBAFFA3187AFA4218EDFA26" box="[1372,1432,1441,1463]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="36" unit="m" value="850.0">850m</quantity>
,
<date id="FFD728A1FFBAFFA31DE1FA231C5AFA47" box="[199,303,1472,1494]" pageId="7" pageNumber="36" value="2003-05-14">14.5.2003</date>
, 4 Ψ (leg. Sausa).
<emphasis id="B91DD273FFBAFFA31CD2FA221F37FA47" box="[500,578,1473,1494]" italics="true" pageId="7" pageNumber="36">Lesvos:</emphasis>
Aghiossos, 11.5.
<date id="FFD728A1FFBAFFA31FD9FA231E1CFA47" box="[767,873,1472,1494]" pageId="7" pageNumber="36" value="1980-05-13">13.5.1980</date>
, 1 Ψ, 1 ɗ (leg. H. Teunissen);
<quantity id="4C91A384FFBAFFA31997FA231988FA47" box="[1201,1277,1472,1494]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="7" pageNumber="36" unit="km" value="2.45">2.45km</quantity>
W of Vatousa,
<quantity id="4C91A384FFBAFFA31DE1FA3C1C76FA65" box="[199,259,1503,1524]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="36" unit="m" value="200.0">200m</quantity>
,
<date id="FFD728A1FFBAFFA31C2BFA3C1C00FA65" box="[269,373,1503,1524]" pageId="7" pageNumber="36" value="2004-04-29">29.4.2004</date>
, 1 ɗ (leg. O. Messinger);
<quantity id="4C91A384FFBAFFA31F5FFA3D1FCCFA65" box="[633,697,1502,1524]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="36" unit="km" value="3.8">3.8km</quantity>
SSE of Agiasos,
<quantity id="4C91A384FFBAFFA31E4FFA3D1ED3FA65" box="[873,934,1502,1524]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="36" unit="m" value="760.0">760m</quantity>
,
<date id="FFD728A1FFBAFFA31E89FA3D1962FA65" box="[943,1047,1502,1524]" pageId="7" pageNumber="36" value="2004-05-24">24.5.2004</date>
, 2 ɗ (leg. A. Kyriakopoulos);
<quantity id="4C91A384FFBAFFA3186EFA3D18FDFA65" box="[1352,1416,1502,1524]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="36" unit="km" value="3.5">3.5km</quantity>
S of Agiasos,
<quantity id="4C91A384FFBAFFA31C62FA1E1CF5F982" box="[324,384,1533,1555]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="36" unit="m" value="860.0">860m</quantity>
,
<date id="FFD728A1FFBAFFA31CAAFA1E1C92F982" box="[396,487,1533,1555]" pageId="7" pageNumber="36" value="2004-06-09">9.6.2004</date>
, 1 ɗ (leg. A. Kyriakopoulos);
<quantity id="4C91A384FFBAFFA31E02FA1E1E11F982" box="[804,868,1533,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="36" unit="km" value="1.2">1.2km</quantity>
W of Stipsi,
<quantity id="4C91A384FFBAFFA31ED7FA1D1958F982" box="[1009,1069,1534,1555]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="36" unit="m" value="330.0">330m</quantity>
,
<date id="FFD728A1FFBAFFA3191FFA1E19D4F982" box="[1081,1185,1533,1555]" pageId="7" pageNumber="36" value="2006-05-19">19.5.2006</date>
, 1 Ψ (leg. C. Zografou).
<collectingCountry id="F37E4EF1FFBAFFA31DE1F9F81C64F9A0" box="[199,273,1563,1585]" name="Turkey" pageId="7" pageNumber="36">Turkey</collectingCountry>
,
<emphasis id="B91DD273FFBAFFA31C3DF9FF1C18F9A3" box="[283,365,1564,1586]" italics="true" pageId="7" pageNumber="36">Ankara:</emphasis>
Soguksu env.,
<quantity id="4C91A384FFBAFFA31F2FF9F81F37F9A0" box="[521,578,1563,1585]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="36" unit="km" value="80.0">80km</quantity>
N of Ankara,
<quantity id="4C91A384FFBAFFA31FF0F9FF1E6BF9A0" box="[726,798,1564,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="36" unit="m" value="1300.0">1300m</quantity>
,
<date id="FFD728A1FFBAFFA31E0FF9F81EE4F9A0" box="[809,913,1563,1585]" pageId="7" pageNumber="36" value="2001-06-14">14.6.2001</date>
, 1 ɗ (leg. K. Denes).
<emphasis id="B91DD273FFBAFFA31949F9FE19CAF9A3" box="[1135,1215,1565,1586]" italics="true" pageId="7" pageNumber="36">Mersin:</emphasis>
Cornelek,
<quantity id="4C91A384FFBAFFA31816F9F8181FF9A0" box="[1328,1386,1563,1585]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="36" unit="km" value="40.0">40km</quantity>
E of Mut,
<date id="FFD728A1FFBAFFA31DD8F9D91C13F9C1" box="[254,358,1594,1616]" pageId="7" pageNumber="36" value="1996-05-29">29.5.1996</date>
, 1 ɗ (leg. M. Halada).
</paragraph>
</subSubSection>
<subSubSection id="C3735DEAFFBAFFA31DB1F99E1D99F848" pageId="7" pageNumber="36" type="diagnosis">
<paragraph id="8BD60E61FFBAFFA31DB1F99E1D99F848" blockId="7.[151,1436,1661,2009]" pageId="7" pageNumber="36">
<emphasis id="B91DD273FFBAFFA31DB1F99E1C64F907" bold="true" box="[151,273,1661,1686]" pageId="7" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="4C6975E2FFBAFFA31C31F99D1CC4F904" box="[279,433,1662,1685]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B91DD273FFBAFFA31C31F99D1CC4F904" box="[279,433,1662,1685]" italics="true" pageId="7" pageNumber="36">C. aegaeicum</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C6975E2FFBAFFA31F7DF99D1FCFF904" box="[603,698,1662,1685]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B91DD273FFBAFFA31F7DF99D1FCFF904" box="[603,698,1662,1685]" italics="true" pageId="7" pageNumber="36">nasutum</emphasis>
</taxonomicName>
group of the subgenus
<emphasis id="B91DD273FFBAFFA31E9BF99E192CF904" box="[957,1113,1661,1685]" italics="true" pageId="7" pageNumber="36">Gyrodromella</emphasis>
(
<bibRefCitation id="EFF87390FFBAFFA3194EF99E184AF904" author="Sedivy" box="[1128,1343,1661,1685]" pageId="7" pageNumber="36" refString="Sedivy, C., Praz, C. J., Muller, A., Widmer, A. &amp; Dorn, S. (2008) Patterns of host-plant choice in bees of the genus Chelostoma: the constraint hypothesis of host-range evolution in bees. Evolution, 62, 2487 - 2507." type="journal article" year="2008">
Sedivy
<emphasis id="B91DD273FFBAFFA31998F99D199AF904" box="[1214,1263,1661,1685]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
., 2008
</bibRefCitation>
;
<bibRefCitation id="EFF87390FFBAFFA3186CF99E1DA6F928" author="Muller" pageId="7" pageNumber="36" refString="Muller, A. (2012) Palaearctic osmiine bees. Systematics and biology of a fascinating group of solitary bees. ETH Zurich. Available from http: // blogs. ethz. ch / osmiini / (29.2.2012)." type="book" year="2012">Müller, 2012</bibRefCitation>
), which consists of four species. The females of the
<taxonomicName id="4C6975E2FFBAFFA31E00F9411EF0F928" box="[806,901,1698,1721]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B91DD273FFBAFFA31E00F9411EF0F928" box="[806,901,1698,1721]" italics="true" pageId="7" pageNumber="36">nasutum</emphasis>
</taxonomicName>
group can be separated from the females of all other
<taxonomicName id="4C6975E2FFBAFFA31DF1F9261C28F94C" box="[215,349,1733,1757]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B91DD273FFBAFFA31DF1F9261C28F94C" box="[215,349,1733,1757]" italics="true" pageId="7" pageNumber="36">Chelostoma</emphasis>
</taxonomicName>
species by the protruding clypeus, which preapically declines sharply at right angles towards the base of the labrum, and by the distinct mediobasal impressions on terga 23. No morphological characters are known so far to reliably distinguish
<taxonomicName id="4C6975E2FFBAFFA31F08F8ED1FBDF8B4" box="[558,712,1806,1829]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B91DD273FFBAFFA31F08F8ED1FBDF8B4" box="[558,712,1806,1829]" italics="true" pageId="7" pageNumber="36">C. aegaeicum</emphasis>
</taxonomicName>
from the other three species of the
<taxonomicName id="4C6975E2FFBAFFA3197FF8ED19CDF8B4" box="[1113,1208,1806,1829]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B91DD273FFBAFFA3197FF8ED19CDF8B4" box="[1113,1208,1806,1829]" italics="true" pageId="7" pageNumber="36">nasutum</emphasis>
</taxonomicName>
group in the female sex. In contrast, the male of
<taxonomicName id="4C6975E2FFBAFFA31CF3F8D11F1AF8D8" box="[469,623,1842,1865]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B91DD273FFBAFFA31CF3F8D11F1AF8D8" box="[469,623,1842,1865]" italics="true" pageId="7" pageNumber="36">C. aegaeicum</emphasis>
</taxonomicName>
is easily recognizable by the form of the median elevation on sternum 2 (
<figureCitation id="135212E4FFBAFFA31DB9F8B61D97F8FC" box="[159,226,1877,1901]" captionStart="FIGURE 9 16. 9" captionStartId="4.[151,250,1726,1749]" captionTargetBox="[152,1434,194,1701]" captionTargetId="figure@4.[151,1436,193,1705]" captionTargetPageId="4" captionText="FIGURE 9 16. 9: Chelostoma aegaeicum spec. nov., median elevation on sternum 2 of ɗ. 10: Chelostoma aegaeicum spec. nov., terga 6 7 of ɗ. 11: Hoplitis occidentalis spec. nov., terga 5 7 of ɗ. 12: Hoplitis occidentalis spec. nov., apical hair band on sternum 5 of ɗ. 13: Hoplitis monticola spec. nov., left half of scutum of Ψ. 14: Hoplitis monticola spec. nov., antennal segments 6 13 of ɗ. 15: Hoplitis caucasicola spec. nov., clypeus of Ψ. 16: Hoplitis caucasicola spec. nov., genitalia of ɗ." httpUri="https://zenodo.org/record/281577/files/figure.png" pageId="7" pageNumber="36">Fig. 9</figureCitation>
) in combination with the shape of the two lateral teeth of tergum 7, which are more or less parallel-sided and widely spaced basally (
<figureCitation id="135212E4FFBAFFA31CBAF89A1C9AF800" box="[412,495,1913,1937]" captionStart="FIGURE 9 16. 9" captionStartId="4.[151,250,1726,1749]" captionTargetBox="[152,1434,194,1701]" captionTargetId="figure@4.[151,1436,193,1705]" captionTargetPageId="4" captionText="FIGURE 9 16. 9: Chelostoma aegaeicum spec. nov., median elevation on sternum 2 of ɗ. 10: Chelostoma aegaeicum spec. nov., terga 6 7 of ɗ. 11: Hoplitis occidentalis spec. nov., terga 5 7 of ɗ. 12: Hoplitis occidentalis spec. nov., apical hair band on sternum 5 of ɗ. 13: Hoplitis monticola spec. nov., left half of scutum of Ψ. 14: Hoplitis monticola spec. nov., antennal segments 6 13 of ɗ. 15: Hoplitis caucasicola spec. nov., clypeus of Ψ. 16: Hoplitis caucasicola spec. nov., genitalia of ɗ." httpUri="https://zenodo.org/record/281577/files/figure.png" pageId="7" pageNumber="36">Fig. 10</figureCitation>
). In
<taxonomicName id="4C6975E2FFBAFFA31F07F8991FC7F800" box="[545,690,1913,1937]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="hebraeum">
<emphasis id="B91DD273FFBAFFA31F07F8991FC7F800" box="[545,690,1913,1937]" italics="true" pageId="7" pageNumber="36">C. hebraeum</emphasis>
</taxonomicName>
(BENOIST), the form of the median elevation on sternum 2 is similar, but the shape of the two lateral teeth of tergum 7 clearly differs, being broadly triangular and close together basally.
</paragraph>
</subSubSection>
<caption id="DF165EE9FFB5FFAC1DB1F95D1C2BF8A1" httpUri="https://zenodo.org/record/281579/files/figure.png" pageId="8" pageNumber="37" targetBox="[151,1432,194,1703]" targetPageId="8">
<paragraph id="8BD60E61FFB5FFAC1DB1F95D1C2BF8A1" blockId="8.[151,1437,1726,1840]" pageId="8" pageNumber="37">
<emphasis id="B91DD273FFB5FFAC1DB1F95D1C33F942" bold="true" box="[151,326,1726,1749]" pageId="8" pageNumber="37">FIGURE 2531.</emphasis>
25:
<taxonomicName id="4C6975E2FFB5FFAC1C55F95D1F43F945" box="[371,566,1726,1748]" class="Insecta" family="Megachilidae" genus="Hoplitis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lithodorae" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1C55F95D1F43F945" box="[371,566,1726,1748]" italics="true" pageId="8" pageNumber="37">Hoplitis lithodorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC1F1AF95C1FD6F945" box="[572,675,1727,1748]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, galeae of proboscis of Ψ. 26:
<taxonomicName id="4C6975E2FFB5FFAC1EFFF95D19EEF945" box="[985,1179,1726,1748]" class="Insecta" family="Megachilidae" genus="Hoplitis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lithodorae" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1EFFF95D19EEF945" box="[985,1179,1726,1748]" italics="true" pageId="8" pageNumber="37">Hoplitis lithodorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC1984F95C187CF945" box="[1186,1289,1727,1748]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, antenna of ɗ. 27:
<taxonomicName id="4C6975E2FFB5FFAC1D9BF93E1C0BF962" box="[189,382,1757,1779]" class="Insecta" family="Megachilidae" genus="Hoplitis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lithodorae" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1D9BF93E1C0BF962" box="[189,382,1757,1779]" italics="true" pageId="8" pageNumber="37">Hoplitis lithodorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC1CA2F93D1C9EF962" box="[388,491,1758,1779]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, appendage of sternum 6 and tergum 7 of ɗ. 28:
<taxonomicName id="4C6975E2FFB5FFAC1EF3F93E19FAF962" box="[981,1167,1757,1779]" class="Insecta" family="Megachilidae" genus="Hoplitis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="manuelae" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1EF3F93E19FAF962" box="[981,1167,1757,1779]" italics="true" pageId="8" pageNumber="37">Hoplitis manuelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC19B3F93D1989F962" box="[1173,1276,1758,1779]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, terga 67 of ɗ. 29:
<taxonomicName id="4C6975E2FFB5FFAC1D9BF91F1CF0F880" box="[189,389,1788,1809]" class="Insecta" family="Megachilidae" genus="Protosmia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1D9BF91F1CF0F880" box="[189,389,1788,1809]" italics="true" pageId="8" pageNumber="37">Protosmia montana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC1CADF91F1C87F880" box="[395,498,1788,1809]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, head of Ψ. 30:
<taxonomicName id="4C6975E2FFB5FFAC1FB6F91F1E22F880" box="[656,855,1788,1809]" class="Insecta" family="Megachilidae" genus="Protosmia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC1FB6F91F1E22F880" box="[656,855,1788,1809]" italics="true" pageId="8" pageNumber="37">Protosmia montana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC1E7BF91F1EB1F880" box="[861,964,1788,1809]" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, sterna 16 of ɗ. 31:
<taxonomicName id="4C6975E2FFB5FFAC19BDF91F1816F880" box="[1179,1379,1788,1809]" class="Insecta" family="Megachilidae" genus="Protosmia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="B91DD273FFB5FFAC19BDF91F1816F880" box="[1179,1379,1788,1809]" italics="true" pageId="8" pageNumber="37">Protosmia montana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E6F08FFB5FFAC184FF91F1DB1F8A1" pageId="8" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
, genitalia of ɗ.
</paragraph>
</caption>
<subSubSection id="C3735DEAFFB5FFAD1DEFF8B41EF4FD86" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37" type="description">
<paragraph id="8BD60E61FFB5FFAC1DEFF8B41F63F802" blockId="8.[151,1436,1879,2012]" pageId="8" pageNumber="37">
<emphasis id="B91DD273FFB5FFAC1DEFF8B41C2FF8E1" bold="true" box="[201,346,1879,1904]" pageId="8" pageNumber="37">Description.</emphasis>
Female: Body length
<quantity id="4C91A384FFB5FFAC1F7EF8B41FC5F8FE" box="[600,688,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="8" pageNumber="37" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79mm</quantity>
. Morphologically identical to
<taxonomicName id="4C6975E2FFB5FFAC192EF8BB19FCF8FE" box="[1032,1161,1880,1903]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B91DD273FFB5FFAC192EF8BB19FCF8FE" box="[1032,1161,1880,1903]" italics="true" pageId="8" pageNumber="37">C. nasutum</emphasis>
</taxonomicName>
(PÉREZ), C.
<emphasis id="B91DD273FFB5FFAC183AF8B418E9F8FE" box="[1308,1436,1879,1903]" italics="true" pageId="8" pageNumber="37">handlirschi</emphasis>
(SCHLETTERER) and
<taxonomicName id="4C6975E2FFB5FFAC1C58F89F1F7AF805" box="[382,527,1916,1940]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="hebraeum">
<emphasis id="B91DD273FFB5FFAC1C58F89F1F7AF805" box="[382,527,1916,1940]" italics="true" pageId="8" pageNumber="37">C. hebraeum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD60E61FFB5FFAD1DEFF87C1EF4FD86" blockId="8.[151,1436,1879,2012]" lastBlockId="9.[151,1437,151,788]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
Male: Body length
<quantity id="4C91A384FFB5FFAC1C87F87C1F72F826" box="[417,519,1951,1975]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="8" pageNumber="37" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811mm</quantity>
.
<emphasis id="B91DD273FFB5FFAC1F34F87C1F22F826" box="[530,599,1951,1975]" italics="true" pageId="8" pageNumber="37">Head:</emphasis>
Lower genal area with a dense tuft of rather long and whitish hairs. Preoccipital ridge carinate. Second segment of the labial palpus about
<date id="FFD728A1FFB5FFAC1E40F8201EF7F84A" box="[870,898,1987,2011]" pageId="8" pageNumber="37">4x</date>
as long as the first segment.
<emphasis id="B91DD273FFB5FFAC19E5F8271868F84D" box="[1219,1309,1988,2012]" italics="true" pageId="8" pageNumber="37">Thorax:</emphasis>
Punctation of scutum and scutellum rather dense with interspaces varying between the diameter of one half and two punctures. Basal zone of the propodeum declivous, shorter than the metanotum and displaying fine longitudinal carinae.
<emphasis id="B91DD273FFB4FFAD1DB1FF021C60FF69" box="[151,277,225,248]" italics="true" pageId="9" pageNumber="38">Metasoma:</emphasis>
Apical margin of terga 14 with white hair bands, which are broadly interrupted medially. Apical margin of terga 23 and to a lesser extent also of terga 1 and 4 distinctly concave. Terga less densely punctate than scutum and scutellum with interspaces up to the diameter of three punctures. Tergum 7 with an apical pit and two lateral teeth, which are parallel-sided and separated from each other by a large and roundish incision (
<figureCitation id="135212E4FFB4FFAD181EFEAF18FAFEF5" box="[1336,1423,332,356]" captionStart="FIGURE 9 16. 9" captionStartId="4.[151,250,1726,1749]" captionTargetBox="[152,1434,194,1701]" captionTargetId="figure@4.[151,1436,193,1705]" captionTargetPageId="4" captionText="FIGURE 9 16. 9: Chelostoma aegaeicum spec. nov., median elevation on sternum 2 of ɗ. 10: Chelostoma aegaeicum spec. nov., terga 6 7 of ɗ. 11: Hoplitis occidentalis spec. nov., terga 5 7 of ɗ. 12: Hoplitis occidentalis spec. nov., apical hair band on sternum 5 of ɗ. 13: Hoplitis monticola spec. nov., left half of scutum of Ψ. 14: Hoplitis monticola spec. nov., antennal segments 6 13 of ɗ. 15: Hoplitis caucasicola spec. nov., clypeus of Ψ. 16: Hoplitis caucasicola spec. nov., genitalia of ɗ." httpUri="https://zenodo.org/record/281577/files/figure.png" pageId="9" pageNumber="38">Fig. 10</figureCitation>
). Median elevation on sternum 2 with a tuft of whitish hairs on each side in its posterior half. Lowermost part of the elevation concave, rounded anteriorly and slightly protruding downwards, constricted in the middle and slightly emarginated posteriorly (
<figureCitation id="135212E4FFB4FFAD1C9EFE541F74FE5E" box="[440,513,439,463]" captionStart="FIGURE 9 16. 9" captionStartId="4.[151,250,1726,1749]" captionTargetBox="[152,1434,194,1701]" captionTargetId="figure@4.[151,1436,193,1705]" captionTargetPageId="4" captionText="FIGURE 9 16. 9: Chelostoma aegaeicum spec. nov., median elevation on sternum 2 of ɗ. 10: Chelostoma aegaeicum spec. nov., terga 6 7 of ɗ. 11: Hoplitis occidentalis spec. nov., terga 5 7 of ɗ. 12: Hoplitis occidentalis spec. nov., apical hair band on sternum 5 of ɗ. 13: Hoplitis monticola spec. nov., left half of scutum of Ψ. 14: Hoplitis monticola spec. nov., antennal segments 6 13 of ɗ. 15: Hoplitis caucasicola spec. nov., clypeus of Ψ. 16: Hoplitis caucasicola spec. nov., genitalia of ɗ." httpUri="https://zenodo.org/record/281577/files/figure.png" pageId="9" pageNumber="38">Fig. 9</figureCitation>
). Sternum 3 with two lateral spots of black bristles. Apical third of sternum 4 densely covered with short and yellowish bristles conspicuously bent at the apex. Apical margin of sternum 5 fringed with a comb of long and moderately wavy yellowish hairs.
</paragraph>
</subSubSection>
<subSubSection id="C3735DEAFFB4FFAD1DEFFDC01D9BFD7E" pageId="9" pageNumber="38" type="distribution">
<paragraph id="8BD60E61FFB4FFAD1DEFFDC01931FDAD" blockId="9.[151,1437,151,788]" box="[201,1092,547,572]" pageId="9" pageNumber="38">
<emphasis id="B91DD273FFB4FFAD1DEFFDC01C17FDAD" bold="true" box="[201,354,547,572]" pageId="9" pageNumber="38">Distribution.</emphasis>
Known so far from Crete, Lesvos and the western half of
<collectingCountry id="F37E4EF1FFB4FFAD1EC8FDC71935FDAD" box="[1006,1088,548,572]" name="Turkey" pageId="9" pageNumber="38">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8BD60E61FFB4FFAD1DEFFDAB1D9BFD7E" blockId="9.[151,1437,151,788]" pageId="9" pageNumber="38">
<emphasis id="B91DD273FFB4FFAD1DEFFDAB1C2FFDF1" bold="true" box="[201,346,584,608]" pageId="9" pageNumber="38">Host plants.</emphasis>
As the other species of the
<taxonomicName id="4C6975E2FFB4FFAD1FB8FDAA1F88FDF1" box="[670,765,585,608]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B91DD273FFB4FFAD1FB8FDAA1F88FDF1" box="[670,765,585,608]" italics="true" pageId="9" pageNumber="38">nasutum</emphasis>
</taxonomicName>
group all appear to be strict specialists of
<taxonomicName id="4C6975E2FFB4FFAD19CEFDAB181EFDF1" box="[1256,1387,584,608]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DD273FFB4FFAD19CEFDAB181EFDF1" box="[1256,1387,584,608]" italics="true" pageId="9" pageNumber="38">Campanula</emphasis>
</taxonomicName>
and possibly also related genera of the
<taxonomicName id="4C6975E2FFB4FFAD1F3BFD8F1FBBFD15" box="[541,718,620,644]" class="Magnoliopsida" family="Campanulaceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="family">Campanulaceae</taxonomicName>
(
<bibRefCitation id="EFF87390FFB4FFAD1FFBFD8F1EC0FD15" author="Sedivy" box="[733,949,620,644]" pageId="9" pageNumber="38" refString="Sedivy, C., Praz, C. J., Muller, A., Widmer, A. &amp; Dorn, S. (2008) Patterns of host-plant choice in bees of the genus Chelostoma: the constraint hypothesis of host-range evolution in bees. Evolution, 62, 2487 - 2507." type="journal article" year="2008">
Sedivy
<emphasis id="B91DD273FFB4FFAD1E15FD8E1E19FD15" box="[819,876,620,644]" italics="true" pageId="9" pageNumber="38">et al.</emphasis>
, 2008
</bibRefCitation>
;
<bibRefCitation id="EFF87390FFB4FFAD1EE7FD8F1923FD15" author="Muller" box="[961,1110,620,644]" pageId="9" pageNumber="38" refString="Muller, A. (2012) Palaearctic osmiine bees. Systematics and biology of a fascinating group of solitary bees. ETH Zurich. Available from http: // blogs. ethz. ch / osmiini / (29.2.2012)." type="book" year="2012">Müller, 2012</bibRefCitation>
),
<taxonomicName id="4C6975E2FFB4FFAD194CFD8E1876FD15" box="[1130,1283,621,644]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B91DD273FFB4FFAD194CFD8E1876FD15" box="[1130,1283,621,644]" italics="true" pageId="9" pageNumber="38">C. aegaeicum</emphasis>
</taxonomicName>
is probably a
<taxonomicName id="4C6975E2FFB4FFAD1DB1FD6C1C3DFD36" box="[151,328,655,679]" class="Magnoliopsida" family="Campanulaceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="family">Campanulaceae</taxonomicName>
oligolege as well. In fact, the only pollen sample available so far consists of pollen of
<taxonomicName id="4C6975E2FFB4FFAD1801FD6C1D95FD5D" class="Magnoliopsida" family="Campanulaceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="family">Campanulaceae</taxonomicName>
, and several individuals of
<taxonomicName id="4C6975E2FFB4FFAD1F37FD561FDFFD5D" box="[529,682,693,716]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B91DD273FFB4FFAD1F37FD561FDFFD5D" box="[529,682,693,716]" italics="true" pageId="9" pageNumber="38">C. aegaeicum</emphasis>
</taxonomicName>
were collected on
<taxonomicName id="4C6975E2FFB4FFAD1E5BFD56180CFD5D" box="[893,1401,692,716]" class="Magnoliopsida" family="Campanulaceae" genus="Legousia" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="species" species="speculum-veneris">
<emphasis id="B91DD273FFB4FFAD1E5BFD5619DAFD5D" box="[893,1199,692,716]" italics="true" pageId="9" pageNumber="38">Legousia speculum-veneris</emphasis>
(Campanulaceae)
</taxonomicName>
on Lesvos.
</paragraph>
</subSubSection>
<subSubSection id="C3735DEAFFB4FFAD1DEFFD1F1EEDFC85" box="[201,920,764,788]" pageId="9" pageNumber="38" type="etymology">
<paragraph id="8BD60E61FFB4FFAD1DEFFD1F1EEDFC85" blockId="9.[151,1437,151,788]" box="[201,920,764,788]" pageId="9" pageNumber="38">
<emphasis id="B91DD273FFB4FFAD1DEFFD1F1C25FC85" bold="true" box="[201,336,764,788]" pageId="9" pageNumber="38">Etymology.</emphasis>
aegaeicus = occurring on islands in the Aegean sea.
</paragraph>
</subSubSection>
</treatment>
</document>