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<mods:title id="6AEFFD1DB71A3790FF4AE3099B167BFA">The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)</mods:title>
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<taxonomicName id="4C348BC65433950236FE78C7FCBBFEA9" ID-CoL="4N3P3" authority="Bonato, Foddai &amp; Minelli, 2002" authorityName="Bonato, Foddai &amp; Minelli" authorityYear="2002" box="[151,864,231,257]" class="Chilopoda" family="Mecistocephalidae" genus="Proterotaiwanella" kingdom="Animalia" order="Geophilomorpha" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tanabei">
<emphasis id="B9402C575433950236FE78C7FCBBFEA9" bold="true" box="[151,864,231,257]" pageId="64" pageNumber="65">
<emphasis id="B9402C575433950236FE78C7FE14FEA9" bold="true" box="[151,463,231,257]" italics="true" pageId="64" pageNumber="65">Proterotaiwanella tanabei</emphasis>
<bibRefCitation id="EFA58DB45433950237BE78C7FCBBFEA9" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[471,864,231,257]" pageId="64" pageNumber="65" pagination="1 - 12" refId="ref52646" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2002) A new mecistocephalid centipede from Ryukyu Islands and a revisitation of ' Taiwanella ' (Chilopoda: Geophilomorpha: Mecistocephalidae). Zootaxa, 86, 1 - 12." type="journal article" year="2002">Bonato, Foddai &amp; Minelli, 2002</bibRefCitation>
</emphasis>
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<paragraph id="8B8BF0455433950236FE792AFEFAFE8C" blockId="64.[151,864,231,292]" box="[151,289,266,292]" pageId="64" pageNumber="65">
<figureCitation id="130FECC05433950236FE792AFEFAFE8C" box="[151,289,266,292]" captionStart="FIGURES 7376" captionStartId="64.[151,269,1816,1840]" captionTargetBox="[294,1315,451,1764]" captionTargetId="figure-114@64.[248,1339,415,1793]" captionTargetPageId="64" captionText="FIGURES 7376. Proterotaiwanella tanabei, holotype: 73, head, dorsal; 74, anterior part of head, ventral (maxillary complex removed); 75, maxillary complex (right side), ventral; 76, forcipular segment and left forcipule, dorsal (poison calyx indicated by solid lines; setae not drawn except for the cerrus). Areolation is not drawn." pageId="64" pageNumber="65">Figs. 7376</figureCitation>
</paragraph>
<paragraph id="8B8BF0455433950236FE797AFC34FE39" blockId="64.[151,1200,346,401]" pageId="64" pageNumber="65">
[1]
<taxonomicName id="4C348BC65433950236AE797AFD71FED9" authority="Bonato et al. 2002" authorityName="Bonato" authorityYear="2002" box="[199,682,346,369]" class="Chilopoda" family="Mecistocephalidae" genus="Proterotaiwanella" kingdom="Animalia" order="Geophilomorpha" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tanabei">
<emphasis id="B9402C575433950236AE797AFE04FED9" box="[199,479,346,369]" italics="true" pageId="64" pageNumber="65">Proterotaiwanella tanabei</emphasis>
<bibRefCitation id="EFA58DB454339502378C797AFD71FED9" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[485,682,346,369]" pageId="64" pageNumber="65" pagination="1 - 12" refId="ref52646" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2002) A new mecistocephalid centipede from Ryukyu Islands and a revisitation of ' Taiwanella ' (Chilopoda: Geophilomorpha: Mecistocephalidae). Zootaxa, 86, 1 - 12." type="journal article" year="2002">
Bonato
<emphasis id="B9402C57543395023450797AFDB4FED9" box="[569,623,346,369]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2002
</bibRefCitation>
</taxonomicName>
— Zootaxa 86: 3 (original description); fig 17.
<taxonomicName id="4C348BC65433950236AE795AFD6AFE39" authority=": Bonato et al. 2003" authorityName="Bonato" authorityYear="2003" box="[199,689,378,401]" class="Chilopoda" family="Mecistocephalidae" genus="Proterotaiwanella" kingdom="Animalia" order="Geophilomorpha" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tanabei">
<emphasis id="B9402C575433950236AE795AFE04FE39" box="[199,479,378,401]" italics="true" pageId="64" pageNumber="65">Proterotaiwanella tanabei</emphasis>
:
<bibRefCitation id="EFA58DB4543395023784795AFD6AFE39" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[493,689,378,401]" pageId="64" pageNumber="65" pagination="539 - 579" refId="ref52693" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2003) Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha). Systematic Entomology, 28, 539 - 579." type="journal article" year="2003">
Bonato
<emphasis id="B9402C57543395023428795AFDACFE39" box="[577,631,378,401]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2003
</bibRefCitation>
</taxonomicName>
— Syst. Entom. 28: 550, 551
</paragraph>
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<emphasis id="B9402C575433950236FE7F38FE84F898" bold="true" box="[151,351,1816,1840]" pageId="64" pageNumber="65">FIGURES 7376.</emphasis>
<taxonomicName id="4C348BC65433950237007F39FD58F898" box="[361,643,1817,1840]" class="Chilopoda" family="Mecistocephalidae" genus="Proterotaiwanella" kingdom="Animalia" order="Geophilomorpha" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tanabei">
<emphasis id="B9402C575433950237007F39FD58F898" box="[361,643,1817,1840]" italics="true" pageId="64" pageNumber="65">Proterotaiwanella tanabei</emphasis>
</taxonomicName>
, holotype: 73, head, dorsal; 74, anterior part of head, ventral (maxillary complex removed); 75, maxillary complex (right side), ventral; 76, forcipular segment and left forcipule, dorsal (poison calyx indicated by solid lines; setae not drawn except for the cerrus). Areolation is not drawn.
</paragraph>
</caption>
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<emphasis id="B9402C575432950336FE78B8FED5FF19" box="[151,270,152,177]" italics="true" pageId="65" pageNumber="66">Diagnosis</emphasis>
. A
<taxonomicName id="4C348BC654329503375778B8FDABFF1A" box="[318,624,152,178]" class="Chilopoda" family="Mecistocephalidae" genus="Proterotaiwanella" kingdom="Animalia" order="Geophilomorpha" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9402C5754329503375778B8FDC9FF19" box="[318,530,152,177]" italics="true" pageId="65" pageNumber="66">Proterotaiwanella</emphasis>
species
</taxonomicName>
with 45 leg­bearing segments. Body length reaching at least
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. Clypeal ratio about 3.2. Forcipular article I with a very small tooth, article II without teeth, article III with a tooth; tarsungulum with a shallow basal tooth.
</paragraph>
<paragraph id="8B8BF0455432950336AF7930FB88FE82" blockId="65.[151,1437,152,1140]" box="[198,1107,272,298]" pageId="65" pageNumber="66">
<emphasis id="B9402C575432950336AF7930FEBCFE81" box="[198,359,272,297]" italics="true" pageId="65" pageNumber="66">Type material</emphasis>
.
<typeStatus id="548F4EE754329503371F7930FE3CFE82" box="[374,487,272,298]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
: female,
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long, subadult (
<bibRefCitation id="EFA58DB45432950335097930FB9CFE82" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[864,1095,272,298]" pageId="65" pageNumber="66" pagination="1 - 12" refId="ref52646" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2002) A new mecistocephalid centipede from Ryukyu Islands and a revisitation of ' Taiwanella ' (Chilopoda: Geophilomorpha: Mecistocephalidae). Zootaxa, 86, 1 - 12." type="journal article" year="2002">
Bonato
<emphasis id="B9402C575432950335D77930FC23FE81" box="[958,1016,272,297]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
, 2002
</bibRefCitation>
).
</paragraph>
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<emphasis id="B9402C575432950336AF7918FE80FEF9" box="[198,347,312,337]" italics="true" pageId="65" pageNumber="66">
<typeStatus id="548F4EE75432950336AF7918FF26FEF9" box="[198,253,312,337]" pageId="65" pageNumber="66">Type</typeStatus>
locality
</emphasis>
. Gushikawa­son, Kume Island, Ryukyu Ids (
<bibRefCitation id="EFA58DB45432950335147918FBB8FEFA" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[893,1123,312,338]" pageId="65" pageNumber="66" pagination="1 - 12" refId="ref52646" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2002) A new mecistocephalid centipede from Ryukyu Islands and a revisitation of ' Taiwanella ' (Chilopoda: Geophilomorpha: Mecistocephalidae). Zootaxa, 86, 1 - 12." type="journal article" year="2002">
Bonato
<emphasis id="B9402C575432950335B37918FBCEFEF9" box="[986,1045,312,337]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
, 2002
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BF0455432950336AF7940FB72FED2" blockId="65.[151,1437,152,1140]" box="[198,1193,352,378]" pageId="65" pageNumber="66">
<emphasis id="B9402C575432950336AF7940FEBCFED1" box="[198,359,352,377]" italics="true" pageId="65" pageNumber="66">Depository of</emphasis>
<materialsCitation id="3B5CFA185432950337077940FB7FFED2" box="[366,1188,352,378]" location="National Science Museum" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Tokyo" typeStatus="holotype">
<emphasis id="B9402C575432950337077940FDD2FED1" box="[366,521,352,377]" italics="true" pageId="65" pageNumber="66">
<typeStatus id="548F4EE75432950337077940FE44FED1" box="[366,415,352,377]" pageId="65" pageNumber="66">type</typeStatus>
material
</emphasis>
.
<location id="8EEBA69E5432950334717940FC8AFED2" LSID="urn:lsid:plazi:treatment:039D41535433950336FE78C7FCB9FBDC:8EEBA69E5432950334717940FC8AFED2" box="[536,849,352,378]" name="National Science Museum" pageId="65" pageNumber="66" stateProvince="Tokyo">National Science Museum</location>
,
<collectingRegion id="49F03EA75432950335347940FC7CFED2" box="[861,935,352,378]" country="Japan" name="Tokyo" pageId="65" pageNumber="66">Tokyo</collectingRegion>
(
<bibRefCitation id="EFA58DB45432950335DF7940FB46FED2" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[950,1181,352,378]" pageId="65" pageNumber="66" pagination="1 - 12" refId="ref52646" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2002) A new mecistocephalid centipede from Ryukyu Islands and a revisitation of ' Taiwanella ' (Chilopoda: Geophilomorpha: Mecistocephalidae). Zootaxa, 86, 1 - 12." type="journal article" year="2002">
Bonato
<emphasis id="B9402C5754329503327A7940FB95FED1" box="[1043,1102,352,377]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
, 2002
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF0455432950336AF79A8FC1EFE62" blockId="65.[151,1437,152,1140]" pageId="65" pageNumber="66">
<emphasis id="B9402C575432950336AF79A8FE73FE09" box="[198,424,392,417]" italics="true" pageId="65" pageNumber="66">Material examined</emphasis>
.
<materialsCitation id="3B5CFA185432950337D379A8FD78FE62" collectingDate="1992-02-23" collectionCode="T" collectorName="Kume Id. &amp; Tanabe" location="Ryukyu Ids" municipality="Gushikawa-son" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Okinawa">
<specimenCount id="9D323BCC5432950337D379A8FD9CFE0A" box="[442,583,392,418]" pageId="65" pageNumber="66" type="generic">1 specimen</specimenCount>
: female, subadult (
<quantity id="4CCC5DA054329503354779A8FC52FE0A" box="[814,905,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="65" pageNumber="66" unit="mm" value="21.0">21 mm</quantity>
), from
<collectingMunicipality id="6BEF6A3F54329503358C79A8FB45FE0A" box="[997,1182,392,418]" pageId="65" pageNumber="66">Gushikawa­son</collectingMunicipality>
,
<collectorName id="26C195935432950332C779A8FAF9FE0A" box="[1198,1314,392,418]" pageId="65" pageNumber="66">Kume Id.</collectorName>
,
<collectingRegion id="49F03EA754329503335B79A8FF09FE62" country="Japan" name="Okinawa" pageId="65" pageNumber="66">Okinawa Pref.</collectingRegion>
,
<location id="8EEBA69E5432950336B77990FEBCFE62" LSID="urn:lsid:plazi:treatment:039D41535433950336FE78C7FCB9FBDC:8EEBA69E5432950336B77990FEBCFE62" box="[222,359,432,458]" municipality="Gushikawa-son" name="Ryukyu Ids" pageId="65" pageNumber="66" stateProvince="Okinawa">Ryukyu Ids</location>
,
<date id="FF8AD68554329503371A7990FE34FE62" box="[371,495,432,458]" pageId="65" pageNumber="66" value="1992-02-23">
<collectingDate id="EFCE2F6D54329503371A7990FE34FE62" box="[371,495,432,458]" pageId="65" pageNumber="66" value="1992-02-23">23.II.1992</collectingDate>
</date>
,
<collectionCode id="ED2568805432950337957990FDD4FE62" box="[508,527,432,458]" name="Tavera, Department of Geology and Geophysics" pageId="65" pageNumber="66">T</collectionCode>
.
<collectorName id="26C195935432950334737990FDB5FE62" box="[538,622,432,458]" pageId="65" pageNumber="66">Tanabe</collectorName>
leg.
</materialsCitation>
,
<materialsCitation id="3B5CFA185432950334D97990FC1AFE62" box="[688,961,432,458]" collectionCode="NSMT" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
coll.
<collectionCode id="ED2568805432950334837990FCE0FE62" box="[746,827,432,458]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="65" pageNumber="66">NSMT</collectionCode>
(
<typeStatus id="548F4EE75432950335227990FC63FE62" box="[843,952,432,458]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF0455432950336AF79F8FB37FC52" blockId="65.[151,1437,152,1140]" pageId="65" pageNumber="66">
<emphasis id="B9402C575432950336AF79F8FE8BFE59" box="[198,336,472,497]" italics="true" pageId="65" pageNumber="66">Description</emphasis>
. Body size reaching at least about
<quantity id="4CCC5DA054329503348479F8FCF0FE5A" box="[749,811,472,498]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="65" pageNumber="66" unit="cm" value="2.0">2 cm</quantity>
. Body colour yellow, without dark patches; head and forcipular segment orange. Head 1.6 times as long as wide, frontal line indistinct. Antennae 3.6 times as long as the head width; apical sensilla about
<quantity id="4CCC5DA054329503340F7A08FD6AFDEA" box="[614,689,552,578]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="65" pageNumber="66" unit="mm" value="10.0">10 µm</quantity>
long. Clypeus: areolate part with 4 pairs of setae symmetrically arranged on each side, and about 30 scattered peculiar button­like sensilla; plagulae subtriangular, without setae; clypeal ratio about 3.2. Labrum: anterior ala triangular, medial margin reduced to a vertex; posterior margin of each posterior ala with about 20 well developed notches. Buccae without spiculum and without setae. Mandible: about 5 lamellae, first lamella with about 6 teeth, intermediate lamella with about 11 teeth; ventral side with small scales, producing a hairy appearance. First maxillae: distal lobe of the medial projections clavate, distal part of the telopodite attenuate. Second maxillae: telopodite with a small apical claw. Forcipular segment: coxosternum 1.1 times as wide as long; cerrus composed of two lateral groups of setae. Forcipules: article I 1.7 times as long as wide, with a small distal tooth; article II without teeth; article III with a tooth; tarsungulum with a shallow, obtuse, basal tooth; poison calyx reaching article II. A total of 45 leg­bearing segments. Last leg­bearing segment: sternum 1.2 times as wide as long; about 20 pores on each coxopleuron; telopodites slightly swollen, with a small apical tubercle bearing two tiny spines.
</paragraph>
<paragraph id="8B8BF0455432950336AF7C28FDB3FB8A" blockId="65.[151,1437,152,1140]" box="[198,616,1032,1058]" pageId="65" pageNumber="66">
<emphasis id="B9402C575432950336AF7C28FDB9FB89" box="[198,610,1032,1057]" italics="true" pageId="65" pageNumber="66">Distribution in the considered area</emphasis>
.
</paragraph>
<paragraph id="8B8BF0455432950336AF7C12FC19FBE4" blockId="65.[151,1437,152,1140]" box="[198,962,1072,1100]" pageId="65" pageNumber="66">
<materialsCitation id="3B5CFA185432950336AF7C12FC66FBE4" box="[198,957,1072,1100]" county="Kume Island'" location="Ryukyu Islands" municipality="Gushikawa-son" pageId="65" pageNumber="66" specimenCount="1">
<location id="8EEBA69E5432950336AF7C12FE5BFBE4" LSID="urn:lsid:plazi:treatment:039D41535433950336FE78C7FCB9FBDC:8EEBA69E5432950336AF7C12FE5BFBE4" box="[198,384,1074,1100]" county="Kume Island'" municipality="Gushikawa-son" name="Ryukyu Islands" pageId="65" pageNumber="66">Ryukyu Islands</location>
:
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,
<collectingCounty id="62EA88C95432950334337C12FD21FBE4" box="[602,762,1074,1100]" pageId="65" pageNumber="66">Kume Island</collectingCounty>
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(
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locality)
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.
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<emphasis id="B9402C575432950336AF7C7BFE6DFBDC" box="[198,438,1115,1140]" italics="true" pageId="65" pageNumber="66">General distribution</emphasis>
. Only known from the
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locality.
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