236 lines
38 KiB
XML
236 lines
38 KiB
XML
<document id="B97CB3AA7192ABEBBA7C786033AB480B" ID-DOI="10.11646/zootaxa.4044.2.2" ID-GBIF-Dataset="8b09a6a1-35c3-407c-a4b5-076c4af030db" ID-ISSN="1175-5326" ID-Zenodo-Dep="241241" ID-ZooBank="7000893A-A8BC-4196-B76D-385C97B96FBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338858631" checkinUser="plazi" docAuthor="Kanao, Taisuke & Maruyama, Munetoshi" docDate="2015" docId="0395F739FFA2BF2DFF021CCBFC81B8A0" docLanguage="en" docName="zt04044p223.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discoxenus cambodiensis Kanao & Maruyama, 2015, n. sp." docType="treatment" docVersion="11" lastPageNumber="209" masterDocId="FFAC8F41FFA5BF25FF951C5CFFC1BC65" masterDocTitle="Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="223" masterPageNumber="201" pageNumber="208" updateTime="1698619795159" updateUser="plazi">
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<mods:title id="B50F6F33EF3D4807097C19AB02834404">Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="ED164EAE72EE4E50033F373401F94D67" type="personal">
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<mods:namePart id="B535F1D7914B573E7A0B422C5A5A2F11">Kanao, Taisuke</mods:namePart>
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<mods:roleTerm id="33DAE34F276AF70F5F3A3B44ACA8B3E5">Author</mods:roleTerm>
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<mods:namePart id="B0EA91BF1F815F8D8F7A887B49779364">Maruyama, Munetoshi</mods:namePart>
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<mods:typeOfResource id="5F16685D4A87AAD99423A738C58E7DC5">text</mods:typeOfResource>
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<mods:title id="10D69287D71A5A09A1D03391F8E58FE4">Zootaxa</mods:title>
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<mods:part id="29BC08229A6C99FC706A34F6407F05DF">
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<mods:date id="60622CB7FB186F48CB8523A0949E396F">2015</mods:date>
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<mods:number id="14C0EF490D96F51E232FE8AAC9DD1EA3">4044</mods:number>
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<mods:detail id="8043B715038120BF09124B9AE7A91B1E" type="issue">
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<mods:identifier id="2161E7860C5B65CEEADD41CDA82976EE" type="DOI">10.11646/zootaxa.4044.2.2</mods:identifier>
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<treatment id="0395F739FFA2BF2DFF021CCBFC81B8A0" ID-DOI="http://doi.org/10.5281/zenodo.6111777" ID-GBIF-Taxon="119644729" ID-Zenodo-Dep="6111777" LSID="urn:lsid:plazi:treatment:0395F739FFA2BF2DFF021CCBFC81B8A0" httpUri="http://treatment.plazi.org/id/0395F739FFA2BF2DFF021CCBFC81B8A0" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
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<subSubSection id="C32615A4FFA2BF22FF021CCBFEA7BCB6" pageId="7" pageNumber="208" type="nomenclature">
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<paragraph id="8B83462FFFA2BF22FF021CCBFDD6BCD7" blockId="7.[151,535,151,211]" box="[151,535,151,178]" pageId="7" pageNumber="208">
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<heading id="D0CBF143FFA2BF22FF021CCBFDD6BCD7" bold="true" box="[151,535,151,178]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
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<emphasis id="B9489A3DFFA2BF22FF021CCBFDD6BCD7" bold="true" box="[151,535,151,178]" pageId="7" pageNumber="208">
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<taxonomicName id="4C3C3DACFFA2BF22FF021CCBFE0CBCD4" ID-CoL="6CZCG" box="[151,461,151,177]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="cambodiensis" status="sp. nov.">
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<emphasis id="B9489A3DFFA2BF22FF021CCBFE0CBCD4" bold="true" box="[151,461,151,177]" italics="true" pageId="7" pageNumber="208">Discoxenus cambodiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B2746FFA2BF22FE411CC4FDD6BCD7" box="[468,535,152,178]" pageId="7" pageNumber="208" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B83462FFFA2BF22FF021CE6FEA7BCB6" blockId="7.[151,535,151,211]" box="[151,358,186,211]" pageId="7" pageNumber="208">
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(
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<figureCitation id="13075AAAFFA2BF22FF0A1CE6FECBBCB6" box="[159,266,186,211]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1288,1311]" captionTargetBox="[151,1436,625,1267]" captionTargetId="figure@2.[151,1436,625,1268]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus. 1 – 2. Discoxenus latiabdominalis n. sp.: 1. Dorsal view, 2. Ventral view. 3 – 4. D. cambodiensis n. sp.: 3. Dorsal view, 4. Ventral view." httpUri="https://zenodo.org/record/241242/files/figure.png" pageId="7" pageNumber="208">Figs. 3, 4</figureCitation>
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,
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<figureCitation id="13075AAAFFA2BF22FE831CE6FE9FBCB6" box="[278,350,186,211]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="7" pageNumber="208">38–53</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32615A4FFA2BF22FF021D5EFB59BE72" pageId="7" pageNumber="208" type="materials_examined">
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<paragraph id="8B83462FFFA2BF22FF021D5EFDEEBD5A" blockId="7.[151,1437,258,715]" pageId="7" pageNumber="208">
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<emphasis id="B9489A3DFFA2BF22FF021D5EFE11BD7E" bold="true" box="[151,464,258,283]" pageId="7" pageNumber="208">
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<typeStatus id="5487F88DFFA2BF22FF021D5EFF13BD7E" box="[151,210,258,283]" pageId="7" pageNumber="208">Type</typeStatus>
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materials.
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<typeStatus id="5487F88DFFA2BF22FECF1D5EFE0ABD7E" box="[346,459,258,283]" pageId="7" pageNumber="208" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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♂,
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<quantity id="4CC4EBCAFFA2BF22FE6B1D5FFDF8BD7E" box="[510,569,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="1.0">1 km</quantity>
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S of Angkor Wat, Siem Reap,
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<collectingCountry id="F32B06BFFFA2BF22FC351D5EFBD7BD7E" box="[928,1046,258,283]" name="Cambodia" pageId="7" pageNumber="208">Cambodia</collectingCountry>
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,
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<date id="FF8260EFFFA2BF22FBB61D5FFB7BBD7E" box="[1059,1210,259,283]" pageId="7" pageNumber="208" value="2012-08-18">18 VIII 2012</date>
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, Maruyama M. leg. (Colony no. MMCB-T-2012-2-001).
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</paragraph>
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<paragraph id="8B83462FFFA2BF22FF521D16FB59BE72" blockId="7.[151,1437,258,715]" pageId="7" pageNumber="208">
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<emphasis id="B9489A3DFFA2BF22FF521D16FE89BD06" bold="true" box="[199,328,330,355]" pageId="7" pageNumber="208">
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<typeStatus id="5487F88DFFA2BF22FF521D16FE82BD06" box="[199,323,330,355]" pageId="7" pageNumber="208" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectingCountry id="F32B06BFFFA2BF22FEC51D16FE09BD06" box="[336,456,330,355]" name="Cambodia" pageId="7" pageNumber="208">Cambodia</collectingCountry>
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: 11♂♂, 5♀♀, same data as
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<typeStatus id="5487F88DFFA2BF22FC9D1D16FCABBD06" box="[776,874,330,355]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="9D3A8DA6FFA2BF22FCE91D17FC61BD06" box="[892,928,331,355]" pageId="7" pageNumber="208" type="generic">1♂</specimenCount>
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, 1♀, completely dissected). 11♂♂, 13♀♀,
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<quantity id="4CC4EBCAFFA2BF22FA1B1D17FF7ABDE2" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="1.0">1 km</quantity>
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SW of Angkor Wat, Siem Reap,
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<date id="FF8260EFFFA2BF22FDA11D33FD0BBDE2" box="[564,714,367,391]" pageId="7" pageNumber="208" value="2012-08-20">20 VIII 2012</date>
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, Maruyama M. leg. (Colony no. MMCB-T-2012-2-0011). 1♀,
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<quantity id="4CC4EBCAFFA2BF22FA1B1D33FF7ABDCE" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="1.0">1 km</quantity>
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W of Banteay Prei, Siem Reap,
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<date id="FF8260EFFFA2BF22FDB61DCFFD76BDCE" box="[547,695,403,427]" pageId="7" pageNumber="208" value="2012-08-21">21 VIII 2012</date>
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, Maruyama M. leg. (Colony no. MMCB-T-2012-2-0019).
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<specimenCount id="9D3A8DA6FFA2BF22FAD71DCFFAA7BDCE" box="[1346,1382,403,427]" pageId="7" pageNumber="208" type="generic">1♂</specimenCount>
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, 1♀, same date and locality data as the former specimens (Colony no. MMCB-T-2012-2-0020). 3♂♂, 7♀♀,
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<quantity id="4CC4EBCAFFA2BF22FABE1DEBFAA4BDAA" box="[1323,1381,438,463]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="1.0">1 km</quantity>
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S of Ankor Wat, Charles De Gaulle, Siem Reap, Maruyama M. leg. (Colony no. MMCB-T-2012-2-0022). 2♂♂, 8♀♀,
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<quantity id="4CC4EBCAFFA2BF22FF021DA2FF24BE72" box="[151,229,510,535]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="208" unit="km" value="1.6">1.6 km</quantity>
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SW of Angkor Wat, Siem Reap,
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<date id="FF8260EFFFA2BF22FDCF1DA3FD2EBE72" box="[602,751,511,535]" pageId="7" pageNumber="208" value="2014-08-21">21 VIII 2014</date>
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, Kakizoe S. leg. (Colony no. SK010).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32615A4FFA2BF22FF521E7EFC1DBEAE" pageId="7" pageNumber="208" type="diagnosis">
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<paragraph id="8B83462FFFA2BF22FF521E7EFC1DBEAE" blockId="7.[151,1437,258,715]" pageId="7" pageNumber="208">
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<emphasis id="B9489A3DFFA2BF22FF521E7EFE80BE5E" bold="true" box="[199,321,546,571]" pageId="7" pageNumber="208">Diagnosis.</emphasis>
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This species is easily distinguished from all other species by the macrochaetotaxy of tergites III– VIII (6, 6, 6, 4, 4, 4). This species is most similar to
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<taxonomicName id="4C3C3DACFFA2BF22FD601E1BFC08BE3B" box="[757,969,582,606]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="latiabdominalis">
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<emphasis id="B9489A3DFFA2BF22FD601E1BFC08BE3B" box="[757,969,582,606]" italics="true" pageId="7" pageNumber="208">D. latiabdominalis</emphasis>
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</taxonomicName>
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, but is distinguishable from it by a more slender body (
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<figureCitation id="13075AAAFFA2BF22FEA81E36FE71BEE6" box="[317,432,618,643]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1288,1311]" captionTargetBox="[151,1436,625,1267]" captionTargetId="figure@2.[151,1436,625,1268]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus. 1 – 2. Discoxenus latiabdominalis n. sp.: 1. Dorsal view, 2. Ventral view. 3 – 4. D. cambodiensis n. sp.: 3. Dorsal view, 4. Ventral view." httpUri="https://zenodo.org/record/241242/files/figure.png" pageId="7" pageNumber="208">Figs. 3, 4</figureCitation>
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), abdominal sternites V–VII without setae except for posterior margin, paramere with narrower paramerite which is approximately 4 times wider than condylite (
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<figureCitation id="13075AAAFFA2BF22FC7E1ED2FB81BEC2" box="[1003,1088,654,679]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="7" pageNumber="208">Fig. 52</figureCitation>
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), and spermatheca with longer basal part which is approximately 2 times longer than apical part (
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<figureCitation id="13075AAAFFA2BF22FCEE1EEEFC0EBEAE" box="[891,975,690,715]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="7" pageNumber="208">Fig. 53</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF4316A7FFA2BF22FF021BDFFCD0BB90" httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="7" pageNumber="208" targetBox="[156,1429,775,1891]" targetPageId="7">
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<paragraph id="8B83462FFFA2BF22FF021BDFFCD0BB90" blockId="7.[151,1436,1923,2037]" pageId="7" pageNumber="208">
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<emphasis id="B9489A3DFFA2BF22FF021BDFFE94BBFC" bold="true" box="[151,341,1923,1946]" pageId="7" pageNumber="208">FIGURES 38–53.</emphasis>
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<taxonomicName id="4C3C3DACFFA2BF22FEC91BD9FDA2BBFF" box="[348,611,1924,1946]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="cambodiensis" status="sp. nov.">
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<emphasis id="B9489A3DFFA2BF22FEC91BD9FDA2BBFF" box="[348,611,1924,1946]" italics="true" pageId="7" pageNumber="208">Discoxenus cambodiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B2746FFA2BF22FDFF1BD8FD64BBFC" box="[618,677,1924,1945]" pageId="7" pageNumber="208" rank="species">
|
||
<emphasis id="B9489A3DFFA2BF22FDFF1BD8FD64BBFC" bold="true" box="[618,677,1924,1945]" pageId="7" pageNumber="208">n. sp.</emphasis>
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</taxonomicNameLabel>
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: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca.
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</paragraph>
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</caption>
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<subSubSection id="C32615A4FFADBF2DFF521CCBFBB7B819" pageId="8" pageNumber="209" type="description">
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<paragraph id="8B83462FFFADBF2DFF521CCBFD41BE72" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
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<emphasis id="B9489A3DFFADBF2DFF521CCBFE98BCD5" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="209">Description.</emphasis>
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Head (
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<figureCitation id="13075AAAFFADBF2DFE231CCBFDCEBCD5" box="[438,527,151,176]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 38</figureCitation>
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) approximately 1.5 times wider than long, with anterior margin of clypeus rounded. Antenna (
|
||
<figureCitation id="13075AAAFFADBF2DFEE61CE0FE22BCB1" box="[371,483,188,213]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1288,1311]" captionTargetBox="[151,1436,625,1267]" captionTargetId="figure@2.[151,1436,625,1268]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus. 1 – 2. Discoxenus latiabdominalis n. sp.: 1. Dorsal view, 2. Ventral view. 3 – 4. D. cambodiensis n. sp.: 3. Dorsal view, 4. Ventral view." httpUri="https://zenodo.org/record/241242/files/figure.png" pageId="8" pageNumber="209">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="13075AAAFFADBF2DFE641CE0FDCFBCB1" box="[497,526,188,212]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">39</figureCitation>
|
||
) with segment I longer than other segments; segment II as long as segment IV, with 3 macrosetae; segment III bowl shaped, shorter than other segments; segment IV wider than long, dilated distally; segment V wider than other segments, wider than long; segments VI–X successively narrowed distally; segments VI–VII wider than long; segment VIII subquadrate; segments IX–X longer than wide; segment XI approximately 2.3 times longer than wide, widest at middle. Labrum (
|
||
<figureCitation id="13075AAAFFADBF2DFC461D10FBEBBD01" box="[979,1066,332,357]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 40</figureCitation>
|
||
, left side) with anterior margin concave; median projection of apodeme rounded; 5–6 lateral setae present in ventral view (
|
||
<figureCitation id="13075AAAFFADBF2DFB291D33FAD0BDED" box="[1212,1297,367,392]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 40</figureCitation>
|
||
, right side). Mandibles (
|
||
<figureCitation id="13075AAAFFADBF2DFE881DC8FE64BDC9" box="[285,421,404,429]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Figs. 41, 42</figureCitation>
|
||
) covered with more than 80 pores. Left mandible (
|
||
<figureCitation id="13075AAAFFADBF2DFC7D1DC8FBFBBDC9" box="[1000,1082,404,429]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 41</figureCitation>
|
||
) moderately produced adorally at middle. Right mandible (
|
||
<figureCitation id="13075AAAFFADBF2DFE471DEBFDE6BDB5" box="[466,551,439,464]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 42</figureCitation>
|
||
) with a small and rounded tooth. Maxillary palpal segment III approximately 2.3 times longer than wide. Mentum (
|
||
<figureCitation id="13075AAAFFADBF2DFDD61D80FD54BD91" box="[579,661,476,501]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 43</figureCitation>
|
||
) 2 times wider than long, covered with more than 70 pores. Labium with prementum covered with 15–20 pores.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF521E79FEB8BEE1" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
|
||
Pronotum (
|
||
<figureCitation id="13075AAAFFADBF2DFEDD1E78FE5FBE59" box="[328,414,548,573]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 44</figureCitation>
|
||
) sparsely covered with more than 80 macrosetae. Elytron (
|
||
<figureCitation id="13075AAAFFADBF2DFBC01E78FB6BBE58" box="[1109,1194,548,573]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 45</figureCitation>
|
||
) transverse, sparsely covered with yellow setae, 21–23 macrosetae present on disc. Metaventrite 1.3 times longer than mesoventrite, without macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF521ED3FC37BF5D" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
|
||
Tergites III–VIII (
|
||
<figureCitation id="13075AAAFFADBF2DFE1A1ED3FE10BECD" box="[399,465,655,680]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1288,1311]" captionTargetBox="[151,1436,625,1267]" captionTargetId="figure@2.[151,1436,625,1268]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus. 1 – 2. Discoxenus latiabdominalis n. sp.: 1. Dorsal view, 2. Ventral view. 3 – 4. D. cambodiensis n. sp.: 3. Dorsal view, 4. Ventral view." httpUri="https://zenodo.org/record/241242/files/figure.png" pageId="8" pageNumber="209">Fig. 3</figureCitation>
|
||
) glabrous, almost without setae except for each posterior margin. Tergite VIII (
|
||
<figureCitation id="13075AAAFFADBF2DFAD61ED3FA55BECD" box="[1347,1428,655,680]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 46</figureCitation>
|
||
) with posterior margin moderately rounded, 2 pairs of macrosetae present at posterior margin, with 4 short setae at middle of posterior margin. Macrochaetotaxy of abdominal tergites III–VIII = 6, 6, 6, 4, 4, 4. Sternites III–IV (
|
||
<figureCitation id="13075AAAFFADBF2DFAFA1E8BFF67BF71" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1288,1311]" captionTargetBox="[151,1436,625,1267]" captionTargetId="figure@2.[151,1436,625,1268]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus. 1 – 2. Discoxenus latiabdominalis n. sp.: 1. Dorsal view, 2. Ventral view. 3 – 4. D. cambodiensis n. sp.: 3. Dorsal view, 4. Ventral view." httpUri="https://zenodo.org/record/241242/files/figure.png" pageId="8" pageNumber="209">Fig. 4</figureCitation>
|
||
) partially covered with setae. Sternites V–VIII almost nude. Sternites III–VIII densely covered with minute pores anteriorly. Sternites III–VI with a row of 4–8 macrosetae at posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF521F18FEECBFA2" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9489A3DFFADBF2DFF521F18FEC9BF39" box="[199,264,836,860]" italics="true" pageId="8" pageNumber="209">Male.</emphasis>
|
||
Sternite VIII (
|
||
<figureCitation id="13075AAAFFADBF2DFE211F18FDC9BF39" box="[436,520,836,861]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 47</figureCitation>
|
||
) with 4 pairs of macrosetae on posterior half, minute setae present at posterior margin. Median lobe of aedeagus (
|
||
<figureCitation id="13075AAAFFADBF2DFD8E1F3BFD60BFE5" box="[539,673,871,896]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Figs. 50, 51</figureCitation>
|
||
) with basal capsule longer than apical lobe. Paramere (
|
||
<figureCitation id="13075AAAFFADBF2DFA9F1F3BFA9ABFE5" box="[1290,1371,871,896]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 52</figureCitation>
|
||
) with paramerite approximately 4 times wider than condylite; velar sac sclerite with 3 setae; apical lobe with 3 minute setae at apex.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF521F88FBD6B86A" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9489A3DFFADBF2DFF521F88FEDABF89" box="[199,283,980,1004]" italics="true" pageId="8" pageNumber="209">Female</emphasis>
|
||
. Sternite VIII (
|
||
<figureCitation id="13075AAAFFADBF2DFE591F88FDE1BF89" box="[460,544,980,1005]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 48</figureCitation>
|
||
) with 3 pairs of macrosetae, several minute setae present at posterior margin. Spermatheca (
|
||
<figureCitation id="13075AAAFFADBF2DFEA21FABFE49B875" box="[311,392,1015,1040]" captionStart="FIGURES 38 – 53" captionStartId="7.[151,264,1923,1946]" captionTargetBox="[156,1429,775,1891]" captionTargetId="figure@7.[151,1436,767,1901]" captionTargetPageId="7" captionText="FIGURES 38 – 53. Discoxenus cambodiensis n. sp.: 38. Head, 39. Antenna, 40. Labrum (left side = dorsal view, right side = ventral view), 41. Mandible, left, 42. Mandible, right, 43. Mentum, 44. Pronotum, 45. Elytron, right, 46. Tergite VIII, 47. Sternite VIII, male, 48. Sternite VIII, female, 49. Tergites IX and X, 50. Median lobe of aedeagus, ventral view, 51. Median lobe of aedeagus, lateral view, 52. Paramere, 53. Spermatheca." httpUri="https://zenodo.org/record/241246/files/figure.png" pageId="8" pageNumber="209">Fig. 53</figureCitation>
|
||
) with basal part more than 2 times longer than apical part.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF521840FBB7B819" blockId="8.[151,1437,151,1221]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9489A3DFFADBF2DFF521840FEB3B850" bold="true" box="[199,370,1052,1077]" pageId="8" pageNumber="209">Measurement.</emphasis>
|
||
Body length = average
|
||
<quantity id="4CC4EBCAFFADBF2DFD171840FD27B851" box="[642,742,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="8" pageNumber="209" unit="mm" value="1.91">1.91 mm</quantity>
|
||
(2.11–
|
||
<quantity id="4CC4EBCAFFADBF2DFCA31840FC5AB851" box="[822,923,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="8" pageNumber="209" unit="mm" value="1.75">1.75 mm</quantity>
|
||
, N = 10), pronotal length = average
|
||
<quantity id="4CC4EBCAFFADBF2DFAAD1840FA5DB851" box="[1336,1436,1052,1077]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="209" unit="mm" value="0.66">0.66 mm</quantity>
|
||
(
|
||
<quantity id="4CC4EBCAFFADBF2DFF0B1863FE84B832" box="[158,325,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="8" pageNumber="209" unit="mm" value="0.665" valueMax="0.68" valueMin="0.65">0.65–0.68 mm</quantity>
|
||
, N = 10), pronotal width = average
|
||
<quantity id="4CC4EBCAFFADBF2DFD411863FCF6B832" box="[724,823,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="209" unit="mm" value="0.96">0.96 mm</quantity>
|
||
(0.92–1.00 mm, N = 10), elytral length = average
|
||
<quantity id="4CC4EBCAFFADBF2DFAFE1863FF02B819" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="209" unit="mm" value="0.46">0.46 mm</quantity>
|
||
(
|
||
<quantity id="4CC4EBCAFFADBF2DFF441838FEB9B819" box="[209,376,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.1" metricValueMin="4.3" pageId="8" pageNumber="209" unit="mm" value="0.47" valueMax="0.51" valueMin="0.43">0.43–0.51 mm</quantity>
|
||
, N = 10), elytral width = average
|
||
<quantity id="4CC4EBCAFFADBF2DFD661838FC96B819" box="[755,855,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="8" pageNumber="209" unit="mm" value="0.63">0.63 mm</quantity>
|
||
(
|
||
<quantity id="4CC4EBCAFFADBF2DFCF01838FBCDB819" box="[869,1036,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.6" metricValueMin="6.1" pageId="8" pageNumber="209" unit="mm" value="0.635" valueMax="0.66" valueMin="0.61">0.61–0.66 mm</quantity>
|
||
, N = 10).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32615A4FFADBF2DFF5218DBFC81B8A0" pageId="8" pageNumber="209" type="etymology">
|
||
<paragraph id="8B83462FFFADBF2DFF5218DBFA91B8C5" blockId="8.[151,1437,151,1221]" box="[199,1360,1159,1185]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9489A3DFFADBF2DFF5218DBFE8FB8C5" bold="true" box="[199,334,1159,1184]" pageId="8" pageNumber="209">Etymology.</emphasis>
|
||
The specific Latinized epithet is derived from
|
||
<collectingCountry id="F32B06BFFFADBF2DFCCE18DBFC0FB8C5" box="[859,974,1159,1184]" name="Cambodia" pageId="8" pageNumber="209">Cambodia</collectingCountry>
|
||
where its
|
||
<typeStatus id="5487F88DFFADBF2DFBD718D5FBB3B8C4" box="[1090,1138,1161,1185]" pageId="8" pageNumber="209">type</typeStatus>
|
||
locality was found.
|
||
</paragraph>
|
||
<paragraph id="8B83462FFFADBF2DFF5218F0FC81B8A0" blockId="8.[151,1437,151,1221]" box="[199,832,1196,1221]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9489A3DFFADBF2DFF5218F0FE9EB8A0" bold="true" box="[199,351,1196,1221]" pageId="8" pageNumber="209">Host species.</emphasis>
|
||
<taxonomicName id="4C3C3DACFFADBF2DFEF318F0FCFDB8A0" authority="Ahmad, 1965" authorityName="Ahmad" authorityYear="1965" box="[358,828,1196,1221]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="maesodensis">
|
||
<emphasis id="B9489A3DFFADBF2DFEF318F0FD58B8A1" box="[358,665,1196,1220]" italics="true" pageId="8" pageNumber="209">Odontotermes maesodensis</emphasis>
|
||
<bibRefCitation id="EFAD3BDEFFADBF2DFD3518F0FCFDB8A0" author="Ahmad" box="[672,828,1196,1221]" pageId="8" pageNumber="209" refString="Ahmad, M. (1965) Termites (Isoptera) of Thailand. Bulletin of the American Museum of Natural History, 131 (1), 1 - 114." type="journal article" year="1965">Ahmad, 1965</bibRefCitation>
|
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</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |