127 lines
14 KiB
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127 lines
14 KiB
XML
<document id="40FEEE9B5F71E1E5EE48F04C857B7239" ID-DOI="10.1111/j.1096-3642.2011.00748.x" ID-ISSN="0024-4082" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630453203665" checkinUser="valdenar" docAuthor="Jadin, Robert C., Smith, Eric N. & Campbell, Jonathan A." docDate="2011" docId="03957E16FFB1FFD3FEDEFC0C185EDC3D" docLanguage="en" docName="j.1096-3642.2011.00748.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00748.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Ophryacus COPE 1887" docType="treatment" docVersion="3" lastPageNumber="954" masterDocId="FFAC066EFFBAFFD8FF95FFB81C16DA00" masterDocTitle="Unravelling a tangle of Mexican serpents: a systematic revision of highland pitvipers" masterLastPageNumber="958" masterPageNumber="943" pageNumber="954" updateTime="1699276107582" updateUser="plazi">
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<mods:titleInfo id="0387DA8D15D1223070CA1AD57FF2147D">
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<mods:title id="F3201F03A98014437FD852E1EB8DDDE2">Unravelling a tangle of Mexican serpents: a systematic revision of highland pitvipers</mods:title>
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<mods:namePart id="A9DCE91D793AA880EC66FB8016126573">Jadin, Robert C.</mods:namePart>
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<mods:namePart id="63498D2A02BDCDE8D4BE5C41FFB07DA2">Smith, Eric N.</mods:namePart>
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<mods:namePart id="372204D92656990AC18B9B786B2B5A0D">Campbell, Jonathan A.</mods:namePart>
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<mods:typeOfResource id="A9E01D6ACD2BABF959698A411E481041">text</mods:typeOfResource>
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<mods:title id="03A1FEB8A34484BAB0831AB2F76B555B">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="3E2409120616014C685917D0ED41A40B">
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<mods:date id="579CB222A245A62CB25DAC33630540F5">2011</mods:date>
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<mods:number id="AAC23F4ECBCE49BEA3C48F74C2B45984">2011-10-25</mods:number>
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<mods:number id="9E9027BD0DD4147C6D6CEF0574B6C4C8">163</mods:number>
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<mods:number id="18839736AA50D3798A3C00F38ED43518">3</mods:number>
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<mods:url id="B5C0CEA6B630F4B2914B47D88FF635FB">http://dx.doi.org/10.1111/j.1096-3642.2011.00748.x</mods:url>
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<mods:classification id="2A4449ECC7A8E5614CDE513BA1C3E56C">journal article</mods:classification>
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<mods:identifier id="DE20E25DF7ED51AED8836150032ABF1F" type="DOI">10.1111/j.1096-3642.2011.00748.x</mods:identifier>
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<mods:identifier id="CE77E35D409466996DCFC4CBC2C19E89" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03957E16FFB1FFD3FEDEFC0C185EDC3D" ID-DOI="http://doi.org/10.5281/zenodo.5492159" ID-Zenodo-Dep="5492159" LSID="urn:lsid:plazi:treatment:03957E16FFB1FFD3FEDEFC0C185EDC3D" httpUri="http://treatment.plazi.org/id/03957E16FFB1FFD3FEDEFC0C185EDC3D" lastPageNumber="954" pageId="11" pageNumber="954">
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<subSubSection id="C3269C8BFFB1FFD3FEDEFC0C1E75D9C9" box="[331,611,948,969]" pageId="11" pageNumber="954" type="nomenclature">
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<paragraph id="8B83CF00FFB1FFD3FEDEFC0C1E75D9C9" blockId="11.[331,611,948,969]" box="[331,611,948,969]" pageId="11" pageNumber="954">
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<heading id="D0CB786CFFB1FFD3FEDEFC0C1E75D9C9" box="[331,611,948,969]" centered="true" fontSize="8" level="2" pageId="11" pageNumber="954" reason="8">
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<taxonomicName id="4C3CB483FFB1FFD3FEDEFC0C1E75D9C9" authority="COPE, 1887" authorityName="COPE" authorityYear="1887" box="[331,611,948,969]" class="Reptilia" family="Viperidae" genus="Ophryacus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="genus">
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<emphasis id="B9481312FFB1FFD3FEDEFC0C1DC4D9C7" box="[331,466,948,969]" italics="true" pageId="11" pageNumber="954">OPHRYACUS</emphasis>
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COPE, 1887
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3269C8BFFB1FFD3FF31FC63185EDC3D" pageId="11" pageNumber="954" type="materials_examined">
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<paragraph id="8B83CF00FFB1FFD3FF31FC631DD6DE0E" blockId="11.[164,779,985,1039]" pageId="11" pageNumber="954">
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<emphasis id="B9481312FFB1FFD3FF31FC631E1CD9EF" box="[164,522,986,1008]" italics="true" pageId="11" pageNumber="954">
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<typeStatus id="548771A2FFB1FFD3FF31FC631CCED9F0" box="[164,216,987,1008]" pageId="11" pageNumber="954">Type</typeStatus>
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species: Trigonocephalus
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</emphasis>
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[
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<taxonomicName id="4C3CB483FFB1FFD3FDB0FC621E6AD9EF" box="[549,636,986,1007]" class="Reptilia" family="Viperidae" genus="Atropos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="genus">
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<emphasis id="B9481312FFB1FFD3FDB0FC621E6AD9EF" box="[549,636,986,1007]" italics="true" pageId="11" pageNumber="954">Atropos</emphasis>
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</taxonomicName>
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]
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<taxonomicName id="4C3CB483FFB1FFD3FD03FC621D0EDE0E" authority="Jan, 1859" authorityName="Jan" authorityYear="1859" class="Reptilia" family="Viperidae" genus="Ophryacus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="species" species="undulatus">
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<emphasis id="B9481312FFB1FFD3FD03FC621F1DD9EF" box="[662,779,986,1007]" italics="true" pageId="11" pageNumber="954">undulatus</emphasis>
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Jan, 1859
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</taxonomicName>
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, by monotypy.
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</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FF31FB801E58DEA9" blockId="11.[164,779,1080,1193]" pageId="11" pageNumber="954">
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<emphasis id="B9481312FFB1FFD3FF31FB801D34DE4D" box="[164,290,1080,1101]" italics="true" pageId="11" pageNumber="954">Etymology:</emphasis>
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The generic name is derived from the Greek
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<emphasis id="B9481312FFB1FFD3FF6DFBEE1D52DE6B" box="[248,324,1110,1131]" italics="true" pageId="11" pageNumber="954">ophrys</emphasis>
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, meaning brow, and the Latin
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<emphasis id="B9481312FFB1FFD3FD44FBEF1F12DE6C" box="[721,772,1111,1132]" italics="true" pageId="11" pageNumber="954">acus</emphasis>
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, meaning pointed, obviously in reference to the distinctive supraocular spine-like scale.
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</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FF31FB6B1E82DF82" blockId="11.[164,778,1234,1410]" pageId="11" pageNumber="954">
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<emphasis id="B9481312FFB1FFD3FF31FB6B1D15DEE8" box="[164,259,1235,1256]" italics="true" pageId="11" pageNumber="954">Content:</emphasis>
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The genus
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<taxonomicName id="4C3CB483FFB1FFD3FE00FB6A1E06DEE7" authorityName="COPE" authorityYear="1887" box="[405,528,1234,1255]" class="Reptilia" family="Viperidae" genus="Ophryacus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="genus">
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<emphasis id="B9481312FFB1FFD3FE00FB6A1E06DEE7" box="[405,528,1234,1255]" italics="true" pageId="11" pageNumber="954">Ophryacus</emphasis>
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</taxonomicName>
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contains only
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<taxonomicName id="4C3CB483FFB1FFD3FD56FB6B1CEDDF06" authorityName="Jan" authorityYear="1859" class="Reptilia" family="Viperidae" genus="Ophryacus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="species" species="undulatus">
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<emphasis id="B9481312FFB1FFD3FD56FB6B1CEDDF06" italics="true" pageId="11" pageNumber="954">O. undulatus</emphasis>
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</taxonomicName>
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confined to the highlands of the Sierra Madre Oriental (
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<collectingRegion id="49F801E2FFB1FFD3FE8CFAA81D60DF26" box="[281,374,1296,1318]" country="Mexico" name="Hidalgo" pageId="11" pageNumber="954">Hidalgo</collectingRegion>
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,
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<collectingRegion id="49F801E2FFB1FFD3FE12FAA81DF9DF25" box="[391,495,1296,1317]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="11" pageNumber="954">Veracruz</collectingRegion>
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,
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<collectingRegion id="49F801E2FFB1FFD3FD94FAA81E45DF26" box="[513,595,1296,1318]" country="Mexico" name="Puebla" pageId="11" pageNumber="954">Puebla</collectingRegion>
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), the Mesa del Sur (
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<collectingRegion id="49F801E2FFB1FFD3FF48FA961D23DF44" box="[221,309,1326,1348]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="954">Oaxaca</collectingRegion>
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), and the Sierra Madre del Sur (
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<collectingRegion id="49F801E2FFB1FFD3FD3AFA961F11DF43" box="[687,775,1326,1347]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="954">Oaxaca</collectingRegion>
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,
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<collectingRegion id="49F801E2FFB1FFD3FF31FAF51D19DF63" box="[164,271,1357,1379]" country="Mexico" name="Guerrero" pageId="11" pageNumber="954">Guerrero</collectingRegion>
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), where it occurs in pine-oak and cloud forest (
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<bibRefCitation id="EFADB2F1FFB1FFD3FF63FAD31E37DF81" author="Campbell JA & Lamar WW" box="[246,545,1387,1409]" pageId="11" pageNumber="954" refId="ref10047" refString="Campbell JA, Lamar WW. 2004. Venomous reptiles of the Western Hemisphere. 2 vols. Ithaca, NY: Comstock Publishing Associates." type="book" year="2004">Campbell & Lamar, 2004</bibRefCitation>
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: map 84).
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</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FF31FA131E93DFC0" blockId="11.[164,645,1450,1472]" box="[164,645,1450,1472]" pageId="11" pageNumber="954">
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<emphasis id="B9481312FFB1FFD3FF31FA131D4EDFC0" box="[164,344,1451,1472]" italics="true" pageId="11" pageNumber="954">Common name:</emphasis>
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Mexican horned pitviper.
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</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FF31FA52183CD8C5" blockId="11.[164,779,1514,1903]" lastBlockId="11.[826,1441,198,1597]" pageId="11" pageNumber="954">
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<emphasis id="B9481312FFB1FFD3FF31FA521DD1DFFF" box="[164,455,1514,1535]" italics="true" pageId="11" pageNumber="954">Definition and diagnosis:</emphasis>
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Rostral broader than high, moderately to distinctly concave; three preoculars, upper largest and undivided, middle not fused with supralacunal, lower small, somewhat excluded from margin of orbit; three to four supraoculars along dorsal margin of eye including supraocular spine; ten to 13 supralabials; lip margin not scalloped; nine to 14 infralabials; single scale above eye forming long, relatively slender spine, slightly compressed to subcircular in cross section, not occupying most of dorsal margin of orbit, tip pointed; adjacent scales along dorsal ocular margin often also modified, projecting slightly; canthals and internasals often raised into short spines or with especially high keels; scales in the supraocular region small and keeled; ten to 20 (usually 12–18) intersupraoculars; top of head covered with small scales, most having tubercular keels; second supralabial usually separated from prelacunal by single small subfoveal; subocular and supralabial series separated by two to four rows of small, roundish scales; 21 mid-dorsal scale rows; middorsals at midbody not noticeably broad, obtusely rounded; keel generally extending to tip of scale or nearly so, apical pits not apparent; free portion of apex of dorsal scales moderate in extent, barely overlapping contiguous scale; interstitial epidermal fold at cranial end of scale well developed; 157–178 ventrals; 37–57 subcaudals, divided; tail spine straight, about as long as preceding two to three subcaudals, pointed or obtusely rounded.
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</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FCC6FD761971DFC2" blockId="11.[826,1441,198,1597]" pageId="11" pageNumber="954">Frontal bones with concave dorsal surface, strongly elevated margins, moderately longer than wide; postfrontals moderate in size, not contacting frontal, comprising about equal amount of dorsal perimeter of orbit as parietals; posterolateral edges of dorsal surface of parietals forming distinct flat shelf continuing onto parietal as a raised ridge; junction between parietal and pro-otic irregular, not particularly angular; anterior portion of ectopterygoid possessing a shallow depression on medial side accommodating attachment of ectopterygoid retractor muscle; ectopterygoid noticeably longer than expanded, flattened base of pterygoid (posterior to articulation with ectopterygoid) with flat shaft tapering posteriorly; apex of choanal process positioned at about midlength on palatine, process moderately reduced in height, apex broadly rounded; dorsal surface of parietal roughly triangular; zero to one (usually zero) palatine teeth, seven to ten pterygoid teeth, seven to nine dentary teeth; pterygoid teeth extending to level of articulation of pterygoid with ectopterygoid; maxillary fang relatively short, only slightly longer than height of maxilla; fang at rest extending to level of suture between supralabials 7 and 8; splenial and angular bones fused; haemapophyses in contact distally.</paragraph>
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<paragraph id="8B83CF00FFB1FFD3FCC6FA73185EDC3D" blockId="11.[826,1441,198,1597]" pageId="11" pageNumber="954">Dorsum with zig-zag pattern; ground colour olivebrown, green, or grey, sometimes orange or yellow pigment present; dorsal scales usually finely mottled or speckled with black.</paragraph>
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</subSubSection>
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</treatment>
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</document> |