171 lines
17 KiB
XML
171 lines
17 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.5.5.3" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333681" ID-ZooBank="DDE5EFC6-DD26-4B92-836B-7C91D4CBBE32" approvalRequired="4" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="2" checkinTime="1668766693217" checkinUser="plazi" docAuthor="SHCHERBAKOV, DMITRY E." docDate="2022" docId="038E7975217FA259DB72FBCBFF44991C" docLanguage="en" docName="palaeoentomology.5.5.3.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.3" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Strivicia SHCHERBAKOV 2022, gen. nov." docType="treatment" docVersion="2" lastPageNumber="434" masterDocId="FFB7010D217FA258D858FF9FFF9A9C11" masterDocTitle="A new genus of Ipsviciidae (Hemiptera, Cicadomorpha) with a tegminal strigil from the Triassic of Madygen" masterLastPageNumber="438" masterPageNumber="434" pageNumber="434" updateTime="1668767062596" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus of Ipsviciidae (Hemiptera, Cicadomorpha) with a tegminal strigil from the Triassic of Madygen</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>SHCHERBAKOV, DMITRY E.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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<mods:extent unit="page">
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<mods:start>434</mods:start>
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<mods:end>438</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.5.3</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.5.5.3</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7333681</mods:identifier>
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<mods:identifier type="ZooBank">DDE5EFC6-DD26-4B92-836B-7C91D4CBBE32</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:038E7975217FA259DB72FBCBFF44991C" httpUri="http://treatment.plazi.org/id/038E7975217FA259DB72FBCBFF44991C" lastPageId="1" lastPageNumber="434" pageId="0" pageNumber="434">
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<subSubSection pageId="0" pageNumber="434" type="nomenclature">
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<paragraph blockId="0.[810,1452,1108,1206]" box="[810,1009,1108,1132]" pageId="0" pageNumber="434">
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<heading bold="true" box="[810,1009,1108,1132]" fontSize="10" level="2" pageId="0" pageNumber="434" reason="3">
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<emphasis bold="true" box="[810,1009,1108,1132]" pageId="0" pageNumber="434">
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<taxonomicName authority="SHCHERBAKOV, 2022" authorityName="SHCHERBAKOV" authorityYear="2022" box="[810,902,1108,1132]" class="Insecta" family="Ipsviciidae" genus="Strivicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="434" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[810,902,1108,1132]" italics="true" pageId="0" pageNumber="434">Strivicia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[908,1009,1108,1132]" pageId="0" pageNumber="434" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="0.[810,1452,1108,1206]" pageId="0" pageNumber="434">
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urn:lsid:zoobank.org:act:
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B747C684-E04C-422A-ACB3- EE
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<date box="[842,948,1182,1206]" pageId="0" pageNumber="434" value="2537-07-10">07102537</date>
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FA
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</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="434" type="type_taxon">
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<paragraph blockId="0.[810,1452,1255,2092]" pageId="0" pageNumber="434">
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<emphasis bold="true" box="[810,973,1255,1280]" pageId="0" pageNumber="434">
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<typeStatus box="[810,866,1256,1280]" pageId="0" pageNumber="434">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName authority="SHCHERBAKOV, 2022" authorityName="SHCHERBAKOV" authorityYear="2022" box="[995,1174,1256,1280]" class="Insecta" family="Ipsviciidae" genus="Strivicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="434" phylum="Arthropoda" rank="species" species="davidi" status="sp. nov.">
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<emphasis box="[995,1174,1256,1280]" italics="true" pageId="0" pageNumber="434">Strivicia davidi</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1196,1297,1256,1280]" pageId="0" pageNumber="434">
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<taxonomicNameLabel box="[1196,1297,1256,1280]" pageId="0" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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; by present designation.
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</paragraph>
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<paragraph blockId="0.[810,1452,1255,2092]" box="[863,1206,1329,1354]" pageId="0" pageNumber="434">
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<emphasis bold="true" box="[863,1056,1329,1354]" pageId="0" pageNumber="434">Included species.</emphasis>
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<typeStatus box="[1062,1115,1330,1354]" pageId="0" pageNumber="434">Type</typeStatus>
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species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="434" type="etymology">
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<paragraph blockId="0.[810,1452,1255,2092]" pageId="0" pageNumber="434">
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<emphasis bold="true" box="[863,992,1367,1391]" pageId="0" pageNumber="434">Etymology.</emphasis>
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From Latin
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<emphasis box="[1142,1235,1366,1390]" italics="true" pageId="0" pageNumber="434">stridulus</emphasis>
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(creaking, grating) and the genus
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<taxonomicName authorityName="Tillyard" authorityYear="1919" box="[963,1047,1404,1427]" class="Insecta" family="Ipsviciidae" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="434" phylum="Arthropoda" rank="genus">
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<emphasis box="[963,1047,1404,1427]" italics="true" pageId="0" pageNumber="434">Ipsvicia</emphasis>
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</taxonomicName>
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; gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="434" type="diagnosis">
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<paragraph blockId="0.[810,1452,1255,2092]" pageId="0" pageNumber="434">
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<emphasis bold="true" box="[863,980,1440,1464]" pageId="0" pageNumber="434">Diagnosis.</emphasis>
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Tegmen small, length less than
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<quantity box="[1374,1447,1440,1464]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="434" unit="mm" value="5.0">5 mm</quantity>
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, weakly elongate, widest before midlength, tapered beyond; postclaval margin convex; costal area distal to costal fracture sloped ventrally, with finely striate strigil; bSc deeply arched; R stem short; dSc sigmoidal, moderately long, reaching at most 1/2 tegmen length; both R stem and dSc convex; R distal to dSc with 3 anterior branches; M and CuA beyond short triangular basal cell fused for some distance, then divergent up to ~1/2 tegmen length, medial area lanceolate; single crossvein
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<emphasis box="[993,1031,1773,1796]" italics="true" pageId="0" pageNumber="434">r-m</emphasis>
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; posterior M branch fused to CuA
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<subScript attach="right" box="[1414,1422,1789,1803]" fontSize="6" pageId="0" pageNumber="434">1</subScript>
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; 7 apical cells; ocellate spots in radial and medial areas.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="1" lastPageNumber="435" pageId="0" pageNumber="434" type="discussion">
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<paragraph blockId="0.[810,1452,1255,2092]" lastBlockId="1.[136,778,615,1293]" lastPageId="1" lastPageNumber="435" pageId="0" pageNumber="434">
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<emphasis bold="true" box="[863,974,1847,1871]" pageId="0" pageNumber="434">Remarks.</emphasis>
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Distinct from all known ipsviciids in the presence of a strigil. The new genus is more primitive than the Late Triassic genera
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<taxonomicName authority="Tillyard, 1919" authorityName="Tillyard" authorityYear="1919" box="[1148,1401,1920,1945]" class="Insecta" family="Ipsviciidae" genus="Ipsvicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="434" phylum="Arthropoda" rank="genus">
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<emphasis box="[1148,1232,1921,1944]" italics="true" pageId="0" pageNumber="434">Ipsvicia</emphasis>
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<bibRefCitation author="Tillyard, R. J." box="[1244,1401,1920,1945]" pageId="0" pageNumber="434" pagination="857 - 896" refId="ref3533" refString="Tillyard, R. J. (1919) Mesozoic insects of Queensland. No. 7. Hemiptera Homoptera; with a note on the phylogeny of the suborder. Proceedings of the Linnean Society of New South Wales, 44, 857 - 896." type="journal article" year="1919">Tillyard, 1919</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName authority="Tillyard, 1922" authorityName="Tillyard" authorityYear="1922" box="[810,1103,1957,1981]" class="Insecta" family="Ipsviciidae" genus="Ipsviciopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="434" phylum="Arthropoda" rank="genus">
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<emphasis box="[810,936,1958,1981]" italics="true" pageId="0" pageNumber="434">Ipsviciopsis</emphasis>
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<bibRefCitation author="Tillyard, R. J." box="[945,1103,1957,1981]" pageId="0" pageNumber="434" pagination="447 - 470" refId="ref3580" refString="Tillyard, R. J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Planipennia. Proceedings of the Linnean Society of New South Wales, 47, 447 - 470." type="journal article" year="1922">Tillyard, 1922</bibRefCitation>
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</taxonomicName>
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, but is similar to these latter in the tegmen shape, deeply arched bSc, long sigmoidal dSc, lanceolate medial area, and distal R branches in an anterior pecten opposed to the posterior M+CuA pecten. The two latter genera have more elongate tegmina with a straight postclaval margin aligned with the commissural margin, horizontal costal area, concave R stem, concave longitudinal dSc reaching beyond 1/2 tegmen length, 2 or 3 crossveins
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<emphasis box="[733,771,725,748]" italics="true" pageId="1" pageNumber="435">r-m</emphasis>
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, and more numerous apical cells. The new genus is similar to the Middle Triassic (Anisian) genus
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<taxonomicName authority="Papier, Nel, Grauvogel-Stamm & Gall, 1997" authorityName="Papier, Nel, Grauvogel-Stamm & Gall" authorityYear="1997" class="Insecta" family="Tettigoniidae" genus="Triassophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus">
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<emphasis box="[532,696,797,821]" italics="true" pageId="1" pageNumber="435">Triassophyllum</emphasis>
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<bibRefCitation author="Papier, F. & Nel, A. & Grauvogel-Stamm, L. & Gall, J. - C." pageId="1" pageNumber="435" pagination="71 - 77" refId="ref3032" refString="Papier, F., Nel, A., Grauvogel-Stamm, L. & Gall, J. - C. (1997) La plus ancienne sauterelle Tettigoniidae, Orthoptera (Trias, NE France): mimetisme ou exaptation? Palaontologische Zeitschrift, 71, 71 - 77. https: // doi. org / 10.1007 / BF 03022547" type="journal article" year="1997">Papier, Nel, Grauvogel-Stamm & Gall, 1997</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation box="[562,629,833,857]" captionStart="FIGURE 1" captionStartId="1.[136,235,493,517]" captionTargetBox="[151,1436,189,470]" captionTargetId="figure-684@1.[151,1436,189,470]" captionTargetPageId="1" captionText="FIGURE 1. Triassophyllum leopardii Papier, Nel, Grauvogel-Stamm & Gall, 1997, tegmina (images flipped). A, Specimen no. 7882 (Papier et al., 1997, fig. 2B–D; arrows, costal fracture and dSc). B, Specimen no. 9116. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7333683" httpUri="https://zenodo.org/record/7333683/files/figure.png" pageId="1" pageNumber="435">Fig. 1</figureCitation>
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; described in
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<taxonomicName box="[136,254,869,893]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
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, later transferred to
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<taxonomicName authorityName="Tillyard" authorityYear="1919" box="[476,597,869,893]" class="Insecta" family="Ipsviciidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="family">Ipsviciidae</taxonomicName>
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; Shcherbakov in
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<bibRefCitation author="Gorochov, A. V." box="[136,310,906,930]" pageId="1" pageNumber="435" pagination="272 - 279" refId="ref2596" refString="Gorochov, A. V. (2005) [Review of Triassic Orthoptera with descriptions of new and little known] axa: Part 2]. Paleontologicheskii Zhurnal, 3, 46 - 54. Published in English in Paleontological Journal, 39 (3), 272 - 279." type="journal article" year="2005">Gorochov, 2005</bibRefCitation>
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;
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<bibRefCitation author="Lambkin, K. J." box="[319,483,906,930]" pageId="1" pageNumber="435" pagination="503 - 520" refId="ref2698" refString="Lambkin, K. J. (2020) Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera: Cicadomorpha: Scytinopteroidea). Zootaxa, 4860 (4), 503 - 520. https: // doi. org / 10.11646 / zootaxa. 4860.4.2" type="journal article" year="2020">Lambkin, 2020</bibRefCitation>
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) in the moderately long dSc and presence of ocellate spots in the radial and medial areas, but in the latter genus tegmina are much larger, with rich venation, numerous crossveins and dSc originating more distally and reaching beyond 1/2 tegmen length.
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<taxonomicName authority="SHCHERBAKOV, 2022" authorityName="SHCHERBAKOV" authorityYear="2022" box="[687,777,1051,1074]" class="Insecta" family="Ipsviciidae" genus="Strivicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[687,777,1051,1074]" italics="true" pageId="1" pageNumber="435">Strivicia</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[136,245,1087,1111]" pageId="1" pageNumber="435">
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<taxonomicNameLabel box="[136,245,1087,1111]" pageId="1" pageNumber="435" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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is distinct from incompletely known Triassic genera
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<taxonomicName authority="Fujiyama, 1973" authorityName="Fujiyama" authorityYear="1973" box="[216,523,1124,1148]" class="Insecta" family="Ipsviciidae" genus="Ipsvicioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus">
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<emphasis box="[216,343,1124,1148]" italics="true" pageId="1" pageNumber="435">Ipsvicioides</emphasis>
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<bibRefCitation author="Fujiyama, I." box="[351,523,1124,1148]" pageId="1" pageNumber="435" pagination="331 - 386" refId="ref2558" refString="Fujiyama, I. (1973) Mesozoic insect fauna of East Asia. Part I. Introduction and Upper Triassic faunas. Bulletin of the National Science Museum, Tokyo, 16, 331 - 386." type="journal article" year="1973">Fujiyama, 1973</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName authority="Lin, 1982" authorityName="Lin" authorityYear="1982" box="[536,774,1124,1148]" class="Insecta" family="Ipsviciidae" genus="Doryscarta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus">
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<emphasis box="[536,657,1124,1147]" italics="true" pageId="1" pageNumber="435">Doryscarta</emphasis>
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Lin, 1982
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</taxonomicName>
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, and
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<taxonomicName authority="Lin, 1982" authorityName="Lin" authorityYear="1982" box="[192,492,1160,1184]" class="Insecta" family="Ipsviciidae" genus="Shaanxioscarta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus">
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<emphasis box="[192,358,1160,1184]" italics="true" pageId="1" pageNumber="435">Shaanxioscarta</emphasis>
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Lin, 1982
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</taxonomicName>
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, and the Early Jurassic genus
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<taxonomicName authority="Becker-Migdisova, 1962" authorityName="Becker-Migdisova" authorityYear="1962" box="[209,599,1196,1220]" class="Insecta" family="Ipsviciidae" genus="Ipsviciella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus">
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<emphasis box="[209,320,1196,1220]" italics="true" pageId="1" pageNumber="435">Ipsviciella</emphasis>
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<bibRefCitation author="Becker-Migdisova, E. E." box="[331,599,1196,1220]" pageId="1" pageNumber="435" pagination="89 - 104" refId="ref2339" refString="Becker-Migdisova, E. E. (1962) Some new Hemiptera and Psocoptera. Paleontologicheskii Zhurnal, 89 - 104. [In Russian]" type="book chapter" year="1962">Becker-Migdisova, 1962</bibRefCitation>
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</taxonomicName>
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at least in only three anterior branches on R distal to dSc and the shape of tegmen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |